Published May 3, 2024
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How to describe a new species in zoology and avoid mistakes
Authors/Creators
- 1. Division of Ecology and Evolution, Research School of Biology, Australian National University, Acton, ACT 2601 and Australian National Insect Collection, National Research Collections Australia, GPO Box 1700, Canberra, ACT 2601, Australia
- 2. Department of Life Science, National Taiwan Normal University, Taipei, Taiwan 116, Republic of China
- 3. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Apartado 14-0434, Lima- 14, Peru
Description
Braby, Michael F., Hsu, Yu-Feng, Lamas, Gerardo (2024): How to describe a new species in zoology and avoid mistakes. Zoological Journal of the Linnean Society 202 (4): 1-16, DOI: 10.1093/zoolinnean/zlae043, URL: https://doi.org/10.1093/zoolinnean/zlae043
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- urn:lsid:plazi.org:pub:FFCF8510FFF2FFF9FF89FFB0F9510F26
References
- Agapow P-M, Bininda-Emonds ORP, Crandall KA et al. The impact of species concept on biodiversity studies. The Quarterly Review of Biology 2004;79:161-79. https://doi.org/10.1086/383542
- Aplin KP. 'Amateur' taxonomy in Australian herpetology -help or hindrance? Monitor, Journal of the Victorian Herpetological Society 1999;10:104-9.
- Aplin KP, Donnellan SC. An extended description of the Pilbara death adder, Acanthophis wellsi Hoser (Serpentes: Elapidae), with notes on the desert death adder, A. pyrrhus Boulenger, and identification of a possible hybrid zone. Records of the Western Australian Museum 1999;19:277-98.
- Atkins AF. Skipper butterflies: their origins? A preliminary discussion on the higher classification of the higher systematics and a proposed phylogeny of the Hesperiidae. News Bulletin of the Entomological Society of Queensland 2005;33:24-34.
- Atkins AF. A list of butterflies (Papilionoidea) and skipper butterflies (Hesperioidea) found within a 10km diameter area centered on the village of Eudlo in the hinterland of the sunshine coast with a note on their migratory habits. Metamorphosis Australia. Magazine of the Butterfly and Other Invertebrates Club 2012;66:4-11.
- Atkins AF.Skipper butterflies (Hesperoidea [sic]) and bar codes:how old are they, and how do the Aussie species fit in to the scheme of things? (Part one: the sub-family Trapezitinae and their ilk). Metamorphosis Australia. Magazine of the Butterfly and Other Invertebrates Club 2017;86:4-21.
- Bacher S. Still not enough taxonomists: reply to Joppa et al. Trends in Ecology and Evolution 2012;27:65-6; author reply 66. https://doi. org/10.1016/j.tree.2011.11.003
- Balakrishnan R. Species concepts, species boundaries and species identification: a view from the tropics. Systematic Biology 2005;54:689-93. https://doi.org/10.1080/10635150590950308
- Balint Z, Benyamini D. The identity of the nomen dubium Penaincisalia patagonaevaga Johnson, 1990 (Lepidoptera: Lycaenidae, Polyommatinae). Opuscula Zoologica 2020;51:213-7. https://doi. org/10.18348/opzool.2020.2.213
- Beaver EP, Braby MF, Glatz RV et al. Systemtatic revision of the Ogyris idmo (Hewitson, 1862) species group (Lepidoptera: Lycaenidae): implications for the conservation management of Australia's most threatened butterflies. Invertebrate Systematics 2023;37:677-701. https://doi.org/10.1071/is23032
- Borkent A. Diagnosing diagnoses -can we improve our taxonomy? Zookeys 2021;1071:43-8. https://doi.org/10.3897/ zookeys.1071.72904
- Borrell B. The big name hunters. Nature 2007;446:253-5. https://doi. org/10.1038/446253a
- Braby MF. Are taxonomic publications involving nomenclatural acts on early view code compliant? Austral Entomology 2018;57:371-6. https://doi.org/10.1111/aen.12372
- Braby MF, Eastwood RG. Revised taxonomic status of Pseudalmenus barringtonensis Waterhouse, 1928stat.rev.(Lepidoptera:Lycaenidae): uncovering Australia's greatest taxonomic fraud. Invertebrate Systematics 2019;33:530-43.
- Braby MF, Williams MR. Biosystematics and conservation biology: critical scientific disciplines for the management of insect biological diversity. Austral Entomology 2016;55:1-17. https://doi.org/10.1111/ aen.12158
- Braby MF, Zwick A. Taxonomic revision of the Taractrocera ilia (Waterhouse) complex (Lepidoptera: Hesperiidae) from north-western Australia and mainland New Guinea based on morphological and molecular data. Invertebrate Systematics 2015;29:487- 509. https://doi.org/10.1071/is15028
- Braby MF,Eastwood RG, Murray N.The subspecies concept in butterflies: has its application in taxonomy and conservation biology outlived its usefulness? Biological Journal of the Linnean Society 2012;106:699- 716. https://doi.org/10.1111/j.1095-8312.2012.01909.x
- Brown RW. Composition of Scientific Words. A Manual of Methods and a Lexicon of Materials for the Practice of Logotechnics, revised edn. Washington, DC: Smithsonian Institution Press, 1956.
- Campbell S. The Loch Ness Monster: The Evidence. Wellingborough, Northamptonshire: Aquarian Press, 1986, 128.
- Ceriaco LMP, Gutierrez EE, Dubois A. Photography-based taxonomy is inadequate, unneccessary, and potentially harmful for biological sciences. Zootaxa 2016;4196:435-45.
- Ceriaco LMP, Aescht E, Ahyong ST et al. Renaming taxa on ethical grounds threatens nomenclatural stability and scientific communication. Zoological Journal of the Linnean Society 2023;197:283-6.
- Cook LG, Edwards RD, Crisp MD et al. Need morphology always be required for new species descriptions? Invertebrate Systematics 2010;24:322-6. https://doi.org/10.1071/is10011
- Costello MJ, May RM, Stork NE. Can we name Earth's species before they go extinct? Science 2013;339:413-6. https://doi.org/10.1126/ science.1230318
- Costello MJ, Lane M, Wilson S et al. Factors influencing when species are first named and estimating global species richness. Global Ecology and Conservation 2015a;4:243-54. https://doi.org/10.1016/j. gecco.2015.07.001
- Costello MJ, Vanhoorne B, Appeltans W. Conservation of biodiversity through taxonomy,data publication,and collaborative infrastructures. Conservation Biology 2015b;29:1094-9. https://doi.org/10.1111/ cobi.12496
- D'Abrera B. Butterflies of the Holarctic Region. Part III Nymphalidae (concl.), Libytheidae, Riodinidae & Lycaenidae. Victoria: Hill House, 1993, 335-524, i -vii.
- Denzer W, Kaiser H. Naming and gaming: The illicit taxonomic practice of 'nomenclatural harvesting' and how to avoid it. Journal of Zoology 2023;320:161-8. https://doi.org/10.1111/jzo.13061
- Donegan TM.New species and subspecies descriptions do not and should not always requrie a dead type specimen. Zootaxa 2008;1761:37-48.
- Donegan TM. Type specimens, samples of live individuals and the Galapagos Pink Land Iguana. Zootaxa 2009;2201:12-20.
- Dubois A. Proposed rules for the incorporation of nomina of higherranked zoological taxa in the International Code of Zoological Nomenclature. 1. Some general questions, concepts and terms of biological nomenclature. Zoosystema 2005;27:365-426.
- Dubois A, Nemesio A. Does nomenclatural availability of nomina of new species or subspecies require the deposition of vouchers in collections? Zootaxa 2007;1409:1-22.
- Eitschberger U, Racheli T. Catasticta studies (Lepidoptera: Pieridae). Neue Entomologische Nachrichten 1998;41:4-94.
- Fujioka T. Zephyrus (Theclini butterflies) in the world (2)-genus Ussuriana and Laeosopis. Butterflies (Tokyo) 1992;2:3-18.
- Fujioka T. Zephyrus (Theclini butterflies) in the world (3)-genus Antigius and allied genera. Butterflies (Tokyo) 1993;4:3-20.
- Garnett ST, Christidis L. Taxonomy anarchy hampers conservation. Nature 2017;546:25-7. https://doi.org/10.1038/546025a
- Gentile G, Snell H. Conolophus marthae sp. nov. (Squamata, Iguanidae), a new species of land iguana from the Galapagos archipegalo.Zootaxa 2009;2201:1-10.
- Gillman LN, Wright SD. Restoring indigenous names in taxonomy. Communications Biology 2020;3:609. https://doi.org/10.1038/ s42003-020-01344-y
- Godfray HCJ. Challenges for taxonomy.Nature 2002;417:17-9. https:// doi.org/10.1038/417017a
- Grund R. The identification of Gahnia Forst & Forst. F. (Cyperaceae) eating Hesperiidae (Lepidoptera) using immature stages. Victorian Entomologist 1998;28:20-32.
- Gu TT, Wu H, Yang F et al. Genomic analysis reveals a cryptic pangolin species. Proceedings of the National Academy of Sciences of the United States of America 2023;120:1-12.
- Guedes P, Alves-Martins F, Arribas JM et al. Eponyms have no place in 21st-century biological nomenclature. Nature Ecology and Evolution 2023;7:1157-60. https://doi.org/10.1038/s41559-023-02022-y
- Halstead LB, Goriup PD, Middleton JA. The Loch Ness Monster. Nature 1976;259:75-6. https://doi.org/10.1038/259075c0
- Hamilton CA, Shockley FW, Simmons R et al. The future for a prominent taxonomy. Insect Systematics and Diversity 2021;5:1-2.
- Hammer TA, Thiele KR. (119-122) Proposals to amend Articles 51 and 56 and Division III, to allow the rejection of culturally offensive and inappropriate names. Taxon 2021;70:1392-4. https://doi. org/10.1002/tax.12620
- Hay RW, Houston TF, Williams AAE et al. Bring Back the Butterflies. Butterfly Gardening for Western Australians. Perth: Western Australian Museum, [1994], viii+72.
- ICZN. International Code of Zoological Nomenclature, 4th edn. London: The International Trust for Zoological Nomenclature, 1999.
- ICZN. Amendment of Articles 8, 9, 10, 21 and 78 of the International Code of Zoological Nomenclature to expand and refine methods of publication.Bulletin of Zoological Nomenclature 2012;69:161-9.
- Inomata T. On the original description of three Luehdorfia taxa in China (Papilionidae). Tyo to Ga 1995;45:239-41.
- Isaac NJB, Mallet J, Mace GM. Taxonomic inflation: its influence on macroecolgy and conservation. Trends in Ecology and Evolution 2004;19:464-9. https://doi.org/10.1016/j.tree.2004.06.004
- Johnson K. Penaincisalia, a new genus of 'elfin'-like butterflies from the high Andes (Lepidoptera: Lycaenidae). Pan-Pacific Entomologist 1990;66:97-125.
- Johnson K. Genera and species of the Neotropical 'elfin'-like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae). Reports of the Museum of Natural History, University of Wisconsin 1992;22:1-279.
- Johnson K, Matusik D. Five new species and one new subspecies of butterflies from the Sierra de Baoruco of Hispaniola. Annals of the Carnegie Museum 1988;57:221-54. https://doi.org/10.5962/p.215185
- Johnson K, Matusik D. Addendum:a new species of Tmolus (Lycaenidae) from Hispaniola. In: Schwartz A (ed.), The butterflies of Hispaniola. Gainesville: University of Florida Press, 1989, 525-9.
- Johnson K, Eisele RC, MacPherson BN. The 'hairstreak butterflies' (Lycaenidae, Theclinae) of north-western Argentina. II. Strymon, sensu stricto. Bulletin of the Allyn Museum 1990;130:1-77.
- Johnson K, Austin GT, Le Crom J-F et al. The Strephonina, a new infratribe of the Eumaeini with description of fourteen new genera (Lycaenidae). Revista de Theclinae Colombianos 1997;1:1-61.
- de Jong R. Phylogeny and biogeography of the genus Taractrocera Butler, 1870 (Lepidoptera: Hesperiidae), an example of Southeast Asian-Australian interchange. Zoologische Mededelingen 2004;78:383-415.
- Joppa LN, Roberts DL, Pimm SL. The population ecology and social behaviour of taxonomists.Trends in Ecology and Evolution 2011;26:551- 3. https://doi.org/10.1016/j.tree.2011.07.010
- Kaiser H. The Taxon Filter, a novel mechanism designed to facilitate the relationship between taxonomy and nomenclature, vis-a-vis the utility of the Code's Article 81 (the Commission's plenary power). Bulletin of Zoological Nomenclature 2013;70:293-302. https://doi. org/10.21805/bzn.v70i4.a17
- Kaiser H. Best practices in herpetological taxonomy: errata and addenda. Herpetological Review 2014;45:257-68.
- Kaiser H, Crother BI, Kelly CMR et al. Best practices; in the 21st century, taxonomic decisioins in Herpetology are acceptable only when supported by a body of evidence via peer-review. Herpetological Review 2013;44:8-23.
- Kitching RL. Biodiversity and taxonomy: impediment or opportunity. In: Moritz C, Kikkawa J (eds), Conservation Biology in Australia and Oceania.Chipping Norton, Australia: Surrey Beatty, 1993, 253-68.
- Knapp S, Lamas G, Lughadha EN et al. Stability or stasis in the names of organisms: the evolving codes of nomenclature. Philosophical Transactions of the Royal Society of London, Series B: Biological Sciences 2004;359:611-22. https://doi.org/10.1098/rstb.2003.1445
- Krell F-T. Suppressing works of contemporary authors using the Code's publication requirements is neither easy nor advisable. Bulletin of Zoological Nomenclature 2021;78:61-7.
- Krell F-T, Marshall SA. New species described from photographs: Yes? No? Sometimes? A fierce debate and a new declaration of the ICZN. Insect Systematics and Diversity 2017;1:3-19. https://doi. org/10.1093/isd/ixx004
- Krell F-T, Pape T. Electronic publications need registration in ZooBank to be available. Bulletin of Zoological Nomenclature 2015;72:245-51. https://doi.org/10.21805/bzn.v72i3.a2
- Krell F-T, Wheeler QD. Specimen collection: plan for the future. Science 2014;344:815-6.
- Lamas G. A critical review of J.Y. Miller's checklist of the neotropical Castniidae (Lepidoptera). Revista Peruana de Entomologia 1995;37:73-87.
- Lamas G, Bollino M. Revisional notes on the 'amastris ' group of Catasticta Butler, 1870 (Lepidoptera: Pieridae), with descriptions of new species and subspecies. Zootaxa 2004;605:1-19.
- Lamb J. Vanessa kiwa. Moths and Butterflies of New Zealand Trust 2023;46:11-2.
- Lambert O, Bianucci G, Post K et al. The giant bite of a new raptorial sperm whale from the Miocene epoch of Peru. Nature 2010a;466:105-8. https://doi.org/10.1038/nature09067
- Lambert O, Bianucci G, Post K et al. Corrigendum: The giant bite of a new raptorial sperm whale from the Miocene epoch of Peru. Nature 2010b;466:1134-1134. https://doi.org/10.1038/nature09381
- Lawton JH. Nessiteras Rhombopteryx. Oikos 1996;77:378-80. https:// doi.org/10.2307/3545927
- Lee C. Distribution and habitats of three Luehdorfia species in China. Yadoriga 1982;107/108:38-40.
- Lobl I, Klausnitzer B, Hartmann M et al. The silent extinction of species and taxonomists -an appeal to science policymakers and legislators. Diversity 2023;15:1053. https://doi.org/10.3390/d15101053
- Mallet J, Willmott K. Taxonomy: renaissance or tower of babel? Trends in Ecology and Evolution 2003;18:57-9. https://doi.org/10.1016/ s0169-5347(02)00061-7
- May RM. Taxonomy as destiny. Nature 1990;347:129-30. https://doi. org/10.1038/347129a0
- Mayr E. Principles of Systematic Zoology. New York: McGraw-Hill, 1969.
- Mayr E, Ashlock PD. Principles of Systematic Zoology, 2nd edn. New York: McGraw-Hill, 1991.
- MillerJY.Castniidae.In:HeppnerJB (ed.),AtlasofNeotropicalLepidoptera. Checklist: Part 2. Hyblaeoidea-Pyraloidea-Tortricoidea. Gainesville: Association for Tropical Lepidoptera/Scientific Publishers, 1995, 133-7, 176-7.
- Neave SA. Nomenclator Zoologicus. A List of the Names of Genera and Subgenera in Zoology from the Tenth Edition of Linnaeus 1758 to the end of 1935. London: The Zoological Society of London, 1939-40.
- Neave SA. Nomenclator Zoologicus. (Vol. 6 edited by M.A. Edwards & A.T. Hopwood; Vol. 7 by M.A. Edwards & H.G. Vevers). London: The Zoological Society of London, 1950-75.
- Ohl M. The Art of Naming. (Translated by Elisabeth Lauffer). Cambridge, MA: The MIT Press, 2018, xv+294.
- Okano K. Descriptons of four new butterflies of the genus Delias (Lep.: Pieridae), with some notes on Delias. Tokurana 1989:14:1-6.
- Pape T. Species can be named from photos. Nature 2016;537:307. https://doi.org/10.1038/537307b
- Peggie D, Vane-Wright RI, Yata O. An illustrated checklist of the pierid butterflies of northern and central Maluku (Indonesia). Butterflies (Tokyo) 1995;11:23-47.
- Polaszek A, Wilson EO. Sense and stability in animal names. Trends in Ecology and Evolution 2005;20:421-2. https://doi.org/10.1016/j. tree.2005.05.016
- deQueirozK.Speciesconceptsandspeciesdelimitation.SystematicBiology 2007;56:879-86. https://doi.org/10.1080/10635150701701083
- Reissinger E.Zur Taxonomie und Systematik der Gattung Catasticta Butler (Lepidoptera, Pieridae). Entomologische Zeitschrift 1972;82:97-124.
- Rheindt FE, Bouchard P, Pyle RL et al. Tightening the requirements for species diagnoses would help integrate DNA-based descriptions in taxonomic practice. PLoS Biology 2023;21:1-13.
- Robbins RK, Lamas G. Nomenclatural changes in the neotropical Eumaeini (Lepidoptera: Lycaenidae: Theclinae). Revista Brasileira de Zoologia 2002;19:197-214. https://doi.org/10.1590/ s0101-81752002000500015
- Robbins RK, Nicolay S. Taxonomy and nomenclature of Strymon istapa and S. columella (Lycaenidae: Theclinae: Eumaeini). Journal of the Lepidopterists Society 1999;52:318-27.
- Rocha LA, Aleixo A, Allen G et al. Specimen collection: an essential tool. Science 2014;344:814-5. https://doi.org/10.1126/ science.344.6186.814
- Rosenberg G, Krell F-T, Pyle R. Call to register new species in ZooBank. Nature 2012;491:40. https://doi.org/10.1038/491040b
- Saigusa T. A study on new subspecies of the tribe Theclini from eastern Asia (Lepidoptera, Lycaenidae).Zephyrus Researches 1993;1:12-22.
- Saito K, Seki Y. Four new taxa of Lycaenidae from South-East Asia (Lepidoptera). Butterflies (Tokyo) [2004];38:44-7.
- Schenk ET, McMasters JH.Procedure in Taxonomy (Revised Edition by A.M. Keen and S.W.Muller). Stanford, CA: Standford University Press, 1948.
- Scott P, Rines R. Naming the Loch Ness monster. Nature 1975;258:466- 8. https://doi.org/10.1038/258466a0
- Slapeta J. Ten simple rules for describing a new (parasite) species. International Journal for Parasitology: Parasites and Wildlife 2013;2:152-4.
- Smart P. The International Butterfly Book. London: Salamander Books, 1975, 275.
- Stradomsky BV. A molecular phylogeny of subfamily Polyommatinae (Lepidoptera: Lycaenidae). Caucasian Entomological Bulletin 2016;12:145-56. https://doi.org/10.23885/1814-3326-2016-12-1- 145-156
- Tautz D, Arctander P, Minelli A et al. A plea for DNA taxonomy. Trends in Ecology and Evolution 2003;18:70-4. https://doi.org/10.1016/ s0169-5347(02)00041-1
- Taxonomy Decadal Plan Working Group. Discovering Diversity: A Decadal Plan for Taxonomy and Biosystematics in Australia and New Zealand 2018-2028. Canberra and Wellington: Australian Academy of Science and Royal Society Te Aparangi, 2018.
- Thiele KR, Conix S, Pyle RL et al. Towards a global list of accepted species I. Why taxonomists sometimes disagree, and why this matters. Organisms Diversity and Evolution 2021;21:615-22. https://doi. org/10.1007/s13127-021-00495-y
- Thorpe SE. Is photography-based taxonomy really inadequate, unnecessary, and potentially harmful for biological sciences? A reply to Ceriaco et al. (2016). Zootaxa 2017;4226:449-50.
- Toussaint EFA, Braby MF, Muller CJ et al. Molecular phylogeny, systematics and generic classification of the butterfly subfamily Trapezitinae (Lepidoptera: Papilionoidea: Hesperiidae). Zoological Journal of the Linnean Society 2022;195:1407-21.
- Vane-Wright RI. Taxonomy, methods of. In: Levin SA (ed.), Encyclopedia of Biodiversity,2nd edn.Waltham,MA:Academic Press,2013, 97-111.
- Vane-Wright RI, Whalley PES. Linnaeus' fabulous butterfly. The Linnean 1985;1:19-24.
- Viloria AL, Pyrcz TW, Wojtusiak J et al. A brachypterous butterfly? Proceedings of the Royal Society of London (B) (Supplement) Biology Letters 2003;270:21-4.
- Wall F. A new krait from Oudh (Bungarus walli). Journal of the Bombay Natural History Society 1907;17:608-11.
- Wang Y-H, Hsiao Y-W, Lee K-H et al. Acoustic differentiation and behavioral response reveals cryptic species within Buergeria treefrogs (Anura, Rhacophoridae) from Taiwan. PLoS ONE 2017;12:1-23.
- Wang Y-H, Hsiao Y-W, Lee K-H et al. Correction: acoustic differentiation and behavioral response reveals cryptic species within Buergeria treefrogs (Anura, Rhacophoridae) from Taiwan. PLoS ONE 2018;13:e0193040. https://doi.org/10.1371/journal.pone.0193040
- Wang Y-H, Hsiao Y-W, Lee K-H et al. Renaming a new Buergeria species from Taiwan -correction of a nomenclatural act not available to ICZN. Nature Conservation Quaerterly 2020;111:4-13.
- Wheeler QD, Raven PH,Wilson EO.Taxonomy:impediment or expedient? Science 2004;303:285. https://doi.org/10.1126/science.303.5656.285
- Wiens JJ. Species delimitation: new approaches for discovering diversity. Systematic Biology 2007;56:875-8. https://doi. org/10.1080/10635150701748506
- Will KW, Mishler BD, Wheeler QD.The perils of DNA barcoding and the need for integrative taxonomy. Systematic Biology 2005;54:844-51. https://doi.org/10.1080/10635150500354878
- Williams D, Wuster W, Fry BG. The good, the bad and the ugly:Australian snake taxonomists and a history of the taxonomy of Australia's venomous snakes. Toxicon 2006;48:919-30. https://doi.org/10.1016/j. toxicon.2006.07.016
- Wilson EO. Taxonomy as a fundamental discipline. Philosophical Transactions of the Royal Society of London, Series B: Biological Sciences 2004;359:739. https://doi.org/10.1098/rstb.2003.1440
- Winston JE.Describing Species.Practical Taxonomic Procedure for Biologists. New York: Columbia University Press, 1999.
- Winston JE.Twenty-first century biological nomenclature -the enduring power of names. Integrative and Comparative Biology 2018;58:1122- 31. https://doi.org/10.1093/icb/icy060
- Wojtusiak J. North Andean cloud forest and subparamo species of the genus Catasticta: C. chrysolopha Kollar C. spectrum Reissinger and C. abdita new species. Lambillionea 1998;98:595-9.
- Wuster W, Kaiser H. Bungled Bungarus: lessons from a venomous snake complex illustrate why taxonomic decisions belong in taxonomycompetent journals. Zootaxa 2023;5297:139-43. https://doi. org/10.11646/zootaxa.5297.1.9
- Wuster W,Bush B, Keogh JS et al. Taxonomic contributions in the 'amateur' literature: comments on recent descriptions of new genera and species by Raymond Hoser.Litteratura Serpentium 2001;21:67-79, 86-91.
- Wuster W, Thomson SA, O'Shea M et al. Confronting taxonomic vandalism in biology: conscientious community self-organization can preserve nomenclatural stability. Biological Journal of the Linnean Society 2021;133:645-70.
- Yagishita A,Nakano S, Morita S.An Illustrated List of the Genus Delias Hubner of the World. Tokyo:Khepera Publishers, 1993, xiv+384, xiv+410+vi pls.
- Yago M. A new species of Horaga Moore (Lepidoptera, Lycaenidae, Lycaeninae, Theclini), with a key to the species of the subtribe Horagina. Transactions of the Lepidopterological Society of Japan 2004;55:13-25.
- Yeates DK. Viva La Revolucion! Designing the digital renaissance in zoological taxonomy. Australian Journal of Entomology 2009;48:189-93. https://doi.org/10.1111/j.1440-6055.2009.00703.x
- Yeates DK, Seago A, Nelson L et al. Integrative taxonomy, or iterative taxonomy? Systematic Entomology 2011;36:209-17. https://doi. org/10.1111/j.1365-3113.2010.00558.x
- Zhang Z-Q. A new era in zoological nomenclature and taxonomy: ICZN accepts e-publication and launches ZooBank. Zootaxa 2012;3450:8. https://doi.org/10.11646/zootaxa.3450.1.2