Dugastella nitida (A. Milne-Edwards, 1879), n. comb.

(Figs 1-7)

Caridina nitida A. Milne-Edwards, 1879: 77-78.

“penaeid shrimps” – Crosnier in Nury 1988: 305, pl. 6, figs 4, 7. — Nury & Thomassin 1994: 103; Gaudant et al. 2018: 464,475, fig.14C, D.

TYPE MATERIAL. — Neotype (designated herein) • France. 1 specimen; near Aix-en-Provence, “Insect bed” of Les Figons; Late Oligocene, late Chattian; MNHN.F.A59323 (Figs 1A, B; 2A, D; 3B, also figured by Gaudant et al. 2018: fig. 14C).

TYPE LOCALITY. — Aix-en-Provence, France.

TYPE AGE. — Late Oligocene (late Chattian).

ADDITIONAL MATERIAL. — Same locality data as neotype; MNHN.F.A59324 (also figured by Gaudant et al. 2018: fig. 14D), series of eight specimens numbered from MNHN.F.A90841 to MNHN.F.A90848.

COMPARATIVE MATERIAL. — Dugastella marocana Bouvier, 1912: Settat spring, Chaouia, Morocco, 1913, P. Pallary coll., MNHN-IU-2016-11857 (ex MNHN-Na-11925).

AMENDED DESCRIPTION

Small sized shrimp (total body length 17-27 mm). Rostrum compressed, acute, slightly curved upwards distally with a weakly convex part near its basis, bearing on its dorsal margin 18-26 moveable teeth arranged in a continuous series beginning anteriorly to the orbit and reaching the apex, leaving an unarmed proximal portion of the rostrum, and 14-18 serrations on its ventral margin (Fig.2D, E). Rostrum overreaching scaphocerite and about as long as carapace (0.9-1.1 times carapace length). Carapace smooth, without grooves. In the best-preserved specimens, supra-orbital tooth visible on the carapace.Antennal tooth fused with lower orbital angle. Pterygostomial angle acute and slightly produced anteriorly. Eyes well developed with peduncle reaching half of basal segment of antennular peduncle.Cornea not preserved. Flagellum of the antenna at least as long as the body. Long scaphocerite (4.7 times as long as wide).Antennules rarely preserved but longer than carapace.Long stylocerite,almost reaching first segment of antennular peduncle.Basal segment about as long as second and third segments combined.Mouthparts visible in some specimens: mandibles robust, with the incisor process bearing at least one denticle, lacking a palp (Fig.4 A-D), and first maxilla composed of a distal endite narrower at its base and with a somewhat straight inner margin on its distal end,a broad round proximal endite and a palp (Fig. 4E, F). Third maxilliped long. First and second pereiopods with small chela, dactylus about as long as palma. Pereiopods 3 to 5 similar, propodus 2-4 times as long as dactylus, few spiniform setae on dactylus (Fig. 5E, F), the distal one being the largest. Pleopods not preserved in our additional specimens.Long pleon with typical caridean hump on the third segment.Sixth somite 0.6-0.7times as long as carapace, 1.3-2 times as long as fifth somite, 0.8 times as long as telson. Telson 2.8 times as long as wide, with three pairs of dorsolateral spiniform setae, with 6 distal setae and an acute projection on its posterior margin (Fig. 5A, B). Uropodal exopod with diaeresis bearing a single large spiniform seta (Fig. 5C, D). Endopod slightly shorter or as long as exopod (Fig. 5A, B).