Type:— COLOMBIA. Santander: vicinity of Tona, Eastern Cordillera, 17 February 1927, E. P. Killip et A. C. Smith 19749 (holotype: US!; isotypes: A!, GH-34535! GH-46047!).

Terrestrial trees, 7–25 m tall. Leaf blade 11–18 cm long, 6.0– 9.5 cm wide, elliptic, chartaceous, apex acute to acuminate, base cuneate, puberulent adaxially, hispid or puberulent abaxially, secondary veins 13–16 pairs; petiole 2.0–4.0 cm long, glabrous, periderm flaking off; stipules 3.0–6.0 cm long, caducous, hispid. Syconia solitary; peduncle 3.0–7.0 mm long, puberulent; lateral bracts absent; receptacle 1.7–2.5 cm in diam. when dry, muricate, hispid, apex rounded (Fig. 9A).

Distribution and habitat in Colombia: — Antioquia, Caquetá, Cundinamarca, Magdalena, Nariño and Tolima in montane forests of the Andes at elevations between 1500–2300 m (Fig. 8).

Selected specimens: — COLOMBIA. Antioquia: Caldas, Reserva Alto de San Miguel, cuenca alta del Río Medellín, Roldán 2680 (HUA); Medellín, San Cristóbal, Benitez 1310 (JAUM); Medellín, Corregimiento Santa Elena, SE de Medellín, vía Rionegro, Callejas 7364 (HUA, NY); Yarumal, Alberto 1501 (US).—Caquetá: San Vicente del Caguán, Cuenca del Río Pato, Mendoza 5085 (HUA).—Cundinamarca: 3 Km No de Alban en la carretera a Villeta. Barclay 3672 (COL, US); Albán, vereda Java, Granjas del Padre Luna, Suarez 3 (COL); Albán, Granja del Padre Luna, Valencia 110 (COL); Albán, Granjas del Padre Luna, orillas del Río Dulce, Bernal 3178 (COL); Fusagasugá, García-Barriga 11963 (COL, US); Santandercito, Uribe 1048 (JAUM); Sasaíma, San Bernardo, García-Barriga 11587 (COL); Sasaíma, San Bernardo, quebrada de La Maria y Río Dulce, García-Barriga 17584 (COL, US, NY); Carretera a Cachipay, Gutiérrez 143 (COL, US, GH, NY).—Huila: San Agustín, Schultes 5281 (COL, US, GH, K); San Agustín, Parque Arqueológico, Romero-Castañeda 6569 (COL); sudeste de Santa Ana, Little 7250 (F, US).—Magdalena: Sierra Nevada de Santa Marta, este de Quebrada Indiana, Kirkbride 2208, 2380 (COL, US).—Nariño: Túquerres, entre Balalaika y Yascual, caminho, Mora 402 (COL).—Tolima: Cordillera central, Río Ata, Nevado del Huila, Jr.Little 8752 (COL, F, US).

Notes:— Ficus santanderana was considered a synonym of F. tonduzii by DeWolf (1965), but Ficus santanderana differs by its leaf blade width (6.0–9.5 vs. 10–18 cm), length of the secondary veins (2.5–5.5 vs. 6.5–11.5 cm), petiole length (2.0–4.0 vs. 5.0– 9.5 cm), and peduncle length (3.0–7.0 vs. 1.0–2.0 mm). Ficus santanderana was considered a synonym of F. mutisii by Berg (2009), but differs by basal lateral veins (extending to 1/8 the length of leaf vs. 1/ 16 in F. mutisii), and receptacle (1.7–2.5 vs. 0.8–1.2 cm diam.).