Published December 21, 2023 | Version v1

Belohina inexpectata Paulian 1958

  • 1. Viale Venezia 45, 25123 Brescia, Italy - alberto. ballerio. bs @ aballerio. it
  • 2. Department of Zoology, Denver Museum of Nature & Science, 2001 Colorado Boulevard, Denver, Colorado 80205 - 5798, U. S. A. - frank. krell @ dmns. org
  • 3. Parc Botanique et Zoologique de Tsimbazaza, Antananarivo B. P 4096, Madagascar - jasrandri. herpbzt @ gmail. com
  • 4. Department of Entomology, University of California, Riverside, CA, 92521, U. S. A. - hawksdc @ gmail. com
  • 5. Department of Entomology, Washington State University, Pullman, WA 99164 - 6382, U. S. A. - e. murray @ wsu. edu
  • 6. Systematics Research Collections, University of Nebraska State Museum, W 436 Nebraska Hall, Lincoln, NE 68588 - 0546, U. S. A. - mjpaulsen @ unl. edu
  • 7. Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum National d'Histoire Naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, 75005 Paris, France - jerome. sueur @ mnhn. fr
  • 8. Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, Saint Petersburg, 199397, Russia - frolov @ scarabaeoidea. com * Corresponding author

Description

Belohina inexpectata Paulian, 1958

(Fig. 3)

Etymology: not provided by the author, but clearly from Latin inexpectatus, a, um (unexpected), due to the unexpected discovery of this strange beetle.

Type locality: road between Beloha and Tsihombe, 25 km from Beloha (Paulian only mentions “road between Beloha and Tsihombe”, which is the only locality reported on the labels of the holotype, but the paratype in MNHB bears a label indicating the exact distance from Beloha, a detail that has greatly facilitated the first author’s field research).

Material examined: Holotype (female): Madagascar Sud, entre Beloha et Tsihombe, III.57, R. J. E. [probably Jean Elie Randiamasy] / Belohina inexpectata n. g. n. sp. R Paulian det., 1957 / Holotype [pinned, in good condition]; one paratype (male): 25 km de Beloha Rte de Tsihombe III.57 Jean Elie / Belohina inexpectata n. g. n. sp. / Institut Scientifique Madagascar / Paratype [pinned, in good condition] (both holotype and paratype in MNHN).

Further material examined: Madagascar, forêt sèche d’Antreaky (Beloha Androy district), 25° 08’ 09’’ S; 45° 09’ 58’’ E, 150 m a.s.l., 1.III.2008, J. E. Randrianirina leg. (6 specimens: 3 males and 1 female dissected) (ABCB, DMNS, UPSA, PBZT). Madagascar, road Tsiombe-Beloha (Beloha Androy district), 25 km from Beloha, 25° 08’ 11’’ S; 45° 13’ 16’’ E, 96 m a.s.l., 10.XII.2007 A. Ballerio & J. E. Randrianirina leg. (one almost complete dead specimen and many fragments) (ABCB, BMNH, UFMT, UNSM, ZIN).

Notes

Published as part of Ballerio, Alberto, Krell, Frank-Thorsten, Randrianirina, Jasmin E., Hawks, David C., Murray, Elizabeth A., Paulsen, Matthew J., Sueur, Jérôme, Mon-Treuil, Olivier & Frolov, Andrey V., 2023, Rediscovery of the enigmatic Madagascan endemic Belohina inexpectata Paulian, 1958, with notes on its morphology and phylogenetic position (Coleoptera, Scarabaeoidea: Belohinidae), pp. 139-160 in Fragmenta entomologica (Frag. Entomol., Roma) (Frag. Entomol., Roma) 55 (2) on page 143, DOI: 10.13133/2284-4880/1539, http://zenodo.org/record/12763089

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References

  • Paulian R. 1958. Nouveau Coleoptere Scarabeide de Madagascar, type d'une nouvelle sous-famille. Le Naturaliste Malgache, 10: 39 - 46.