Biodiversity Literature Repository

Submit to community
Liberated Data
Published April 11, 2024 | Version v1
Journal article Open

Taxonomic and stratigraphic update of the material historically attributed to Megalosaurus from Portugal

Authors/Creators

Description

Malafaia, Elisabete, Mocho, Pedro, Escaso, Fernando, Narvaéz, Ivan, Ortega, Francisco (2024): Taxonomic and stratigraphic update of the material historically attributed to Megalosaurus from Portugal. Acta Palaeontologica Polonica 69 (2): 127-171, DOI: 10.4202/app.01113.2023, URL: http://dx.doi.org/10.4202/app.01113.2023

Files

source.pdf

Files (14.6 MB)

Name Size Download all
md5:978bd00aee38596d7a81c4ed6491c022
14.6 MB Preview Download

Linked records

Additional details

Identifiers

LSID
urn:lsid:plazi.org:pub:FF8BD00AEE38596DFF81FFEDFF91C022
URL
http://publication.plazi.org/id/FF8BD00AEE38596DFF81FFEDFF91C022

Has part
Taxonomic treatment: http://treatment.plazi.org/id/03B2A872EE315964FFE3F866FB14C09E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/03B2A872EE2C5974FF16FC01FD77C785 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/03B2A872EE215970FCA9FEBDFC55C29D (URL)
Taxonomic treatment: http://treatment.plazi.org/id/03B2A872EE255973FCA9FD0CFCA8C54E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/03B2A872EE1A5948FF16FA4CFDD0C78A (URL)
Figure: 10.5281/zenodo.12199494 (DOI)
Figure: 10.5281/zenodo.12199496 (DOI)
Figure: 10.5281/zenodo.12199498 (DOI)
Figure: 10.5281/zenodo.12199500 (DOI)
Figure: 10.5281/zenodo.12199502 (DOI)
Figure: 10.5281/zenodo.12199504 (DOI)
Figure: 10.5281/zenodo.12199506 (DOI)
Figure: 10.5281/zenodo.12199508 (DOI)
Figure: 10.5281/zenodo.12199510 (DOI)
Figure: 10.5281/zenodo.12199512 (DOI)
Figure: 10.5281/zenodo.12199514 (DOI)
Figure: 10.5281/zenodo.12199516 (DOI)
Figure: 10.5281/zenodo.12199518 (DOI)
Figure: 10.5281/zenodo.12199520 (DOI)
Figure: 10.5281/zenodo.12199522 (DOI)
Figure: 10.5281/zenodo.12199526 (DOI)
Figure: 10.5281/zenodo.12199528 (DOI)
Figure: 10.5281/zenodo.12199530 (DOI)
Figure: 10.5281/zenodo.12199532 (DOI)
Figure: 10.5281/zenodo.12199534 (DOI)
Figure: 10.5281/zenodo.12199538 (DOI)
Figure: 10.5281/zenodo.12199540 (DOI)
Figure: 10.5281/zenodo.12199542 (DOI)
Figure: 10.5281/zenodo.12199545 (DOI)
Figure: 10.5281/zenodo.12199549 (DOI)

References

  • Allain, R. 2001. Redescription de Streptospondylus altdorfensis, le dinosaure theropode de Cuvier, du Jurassique de Normandie. Geodiversitas 23: 349-367.
  • Allain, R. 2002. Discovery of megalosaur (Dinosauria, Theropoda) in the middle Bathonian of Normandy (France) and its implications for the phylogeny of basal Tetanurae. Journal of Vertebrate Paleontology 22: 548-563.
  • Allain, R. 2005. The postcranial anatomy of the megalosaur Dubreuillosaurus valesdunensis (Dinosauria Theropoda) from the middle Jurassic of Normandy, France. Journal of Vertebrate Paleontology 25: 850-858.
  • Allain, R. and Chure, D.J. 2002. Poekilopleuron bucklandii, the theropod dinosaur from the Middle Jurassic (Bathonian) of Normandy. Palaeontology 45: 1107-1121.
  • Allain, R. and Pereda Suberbiola, X. 2003. Dinosaurs of France. Comptes Rendus Palevol 2: 27-44.
  • Allain, R., Vullo, R., Rozada, L., Anquetin, J., Bourgeais, R., Goedert, J., Lasseron, M., Martin, J.E., Perez-Garcia, A., Peyre De Fabregues, C., Royo-Torres, R., Augier, D., Bailly, G., Cazes, L., Despres, Y., Gailliegue, A., Gomez, B., Goussard, F., Lenglet, T., Vacant, R., Mazan, and Tournepiche, J.-F. 2022. Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstatte (southwestern France): implications for continental faunal turnover at the J/K boundary. Geodiversitas 44: 683-752.
  • Allain, R., Xaisanavong, T., Richir, P., and Khentavong, B. 2012. The first definitive Asian spinosaurid (Dinosauria: Theropoda) from the early Cretaceous of Laos. Naturwissenschaften 99: 369-377.
  • Alonso, A. and Canudo, J.I. 2016. On the spinosaurid theropod teeth from the early Barremian (Early Cretaceous) Blesa Formation (Spain). Historical Biology 28: 823-834.
  • Alves, T.M., Gawthorpe, R.L., Hunt, D.W., and Monteiro, J.H. 2002. Jurassic tectono-sedimentary evolution of the Northern Lusitanian Basin (offshore Portugal). Marine and Petroleum Geology 19: 727-754.
  • Alves, T.M, Manuppella, G., Gawthorpe, R.L., Hunt, D.W., and Monteiro, J.H. 2003. The depositional evolution of diapir- and fault-bounded rift basins: examples from the Lusitanian Basin of West Iberia. Sedimentary Geology 162: 273-303.
  • Antunes, M.T. 1976. Dinossaurios eocretacicos de Lagosteiros, Ciencias da Terra UNL 1: 1-35.
  • Antunes, M.T. and Mateus, O. 2003. Dinosaurs of Portugal. Palevol 2: 77-95.
  • Antunes, M.T. and Pais, J. 1978. Notes on deposits from Taveiro stratigraphy, paleontology, age, paleoecology. Ciencias da Terra UNL 4: 109-128.
  • Antunes, M.T. and Sigogneau-Russell, D. 1991. New data on dinosaurs from the Upper Cretaceous of Portugal. Comptes Rendus de l'Academie des Scieces Paris 313 (Serie 11): 113-119.
  • Antunes, M.T. and Sigogneau-Russell, D. 1992. Fauna of small dinosaurs from the Portuguese late Cretaceous. Comunicacoes dos Servicos Geologicos de Portugal 78 (1): 49-62.
  • Azeredo, A.C., Cabral, M.C., Martins, M.J., Loureiro, I.M., and Ines, N. 2010. Stratigraphic study of a new outcrop of the Cabacos Formation (Oxfordian) in the Serra do Bouro region (Caldas da Rainha). Comunicacoes Geologicas 97: 5-22.
  • Azeredo,A.C., Ramalho, M.M., and Wright, P.V. 1998. The Middle-Upper Jurassic disconformity in the Lusitanian Basin, Portugal: preliminary facies analysis and evidence for palaeoclimatic fluctuation. Cuadernos de Geologia Iberica 24: 99-119.
  • Bakker, R.T., Siegwarth, J.D., Kralis, D., and Filla, J. 1992. Edmarka rex: A new, gigantic theropod dinosaur from the middle Morrison Formation, Late Jurassic of the Como Bluff outcrop region. Hunteria 2: 1-24.
  • Barbosa, B.P., Soares, A.F., Rocha, R.B., Manuppella, G., and Henriques, M.H. 2008. Geological map of Portugal at a scale of 1:50.000: explanatory news of the sheet 19-A (Cantanhede). Departamento de Geologia, Instituto Nacional de Engenharia, Tecnologia e Inovacao, Lisboa.
  • Bardet, N., Pereda Suberbiola, X., and Ruiz-Omenaca, J.I. 2008. Mesozoic marine reptiles from the Iberian Peninsula. Geo-Temas 10: 1245-1248.
  • Barker, C.T., Hone, D.W.E., Naish, D., Cau, A., Lockwood, J.A.F., Foster, B., Clarkin, C.E., Schneider, P., and Gostling, N.J. 2021. New spinosaurids from the Wessex Formation (Early Cretaceous, UK) and the European origins of Spinosauridae. Scientific Reports 11: 1-15.
  • Barrett, P.M., Benson, R.B.J., Rich, T.H. and Vickers-Rich, P. 2011. First spinosaurid dinosaur from Australia and the cosmopolitanism of Cretaceous dinosaur faunas. Biology Letters 7: 933-936.
  • Benson, R.B.J. 2008. A redescription of "Megalosaurus" hesperis (Dinosauria, Theropoda) from the Inferior Oolite (Bajocian, Middle Jurassic) of Dorset, United Kingdom. Zootaxa 1931: 57-67.
  • Benson, R.B.J. 2009. An assessment of variability in theropod dinosaur remains from the Bathonian (Middle Jurassic) of Stonesfield and New Park quarry, UK and taxonomic implications for Megalosaurus bucklandii and Iliosuchus incognitus. Palaeontology 52: 857-877.
  • Benson, R.B.J. 2010a. A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods. Zoological Journal of the Linnean Society 158: 882-935.
  • Benson, R.B.J. 2010b. The osteology of Magnosaurus nethercombensis (Dinosauria, Theropoda) from the Bajocian (Middle Jurassic) of the United Kingdom and a re-examination of the oldest records of tetanurans. Journal of Systematic Palaeontology 8: 131-146.
  • Benson, R.B.J., Barrett, P.M., Powell, H.P., and Norman, D.B. 2008. The taxonomic status of Megalosaurus bucklandii (Dinosauria, Theropoda) from the Middle Jurassic of Oxfordshire, UK. Palaeontology 51: 419-424.
  • Bittencourt, J.S. and Kellner, A.W.A. 2004. On a sequence of sacrocaudal theropod dinosaur vertebrae from the Lower Cretaceous Santana Formation, northeastern Brazil. Arquivos do Museu Nacional, Rio de Janeiro 62: 309-320.
  • Bonaparte, J.F. 1986. The dinosaurs (carnosaurs, allosaurids, sauropods, cetiosaurids) from the Middle Jurassic of Cerro Condor (Chubut, Argentina). Annales de Paleontologie 72: 247-386.
  • Britt, B. 1991. Theropods of Dry Mesa Quarry (Morrison Formation, Late Jurassic), Colorado, with emphasis on the osteology of Torvosaurus tanneri. In: B.J. Kowallis and K. Seeley (eds.), Young University Geology Studies 37, 1-72. Ocean Press, Provo.
  • Brusatte, S.L., Benson, R.B.J., and Hutt, S. 2008. The osteology of Neovenator salerii (Dinosauria: Theropoda) from the Wealden Group (Barremian) of the Isle of Wight. Monograph of the Palaeontographical Society 162: 1-75.
  • Brusatte, S., Carr, T.D., Erickson, G.M., Bever, G.S., and Norell, M.A. 2009. A long-snouted, multihorned tyrannosaurid from the Late Cretaceous of Mongolia. PNAS 106: 17261-17266.
  • Buckland, W. 1824. Notice on the Megalosaurus or great fossil lizard of Stonesjield. Transactions of the Geological Society of London 2-1: 390-396.
  • Buckley, L.G. and Currie, P.J. 2014. Analysis of intraspecific and ontogenetic variation in the dentition of Coelophysis bauri (Late Triassic), and implications for the systematics of isolated theropod teeth. In: S.G. Lucas, R. Sullivan, and L.H. Tanner (eds.), New Mexico Museum of Natural History & Science, Bulletin 63: 1-54.
  • Buffetaut, E. 2007. The spinosaurid dinosaur Baryonyx (Saurischia, Theropoda) in the Early Cretaceous of Portugal. Geology Magazine 144: 1021-1025.
  • Buffetaut, E. 2011.An early spinosaurid dinosaur from the Late Jurassic of Tendaguru (Tanzania) and the evolution of the spinosaurid dentition. Oryctos 10: 1-8.
  • Buffetaut, E. and Ingavat, R. 1986. Unusual theropod dinosaur teeth from the Upper Jurassic of Phu Wiang, northeastern Thailand. Revue de Paleobiology 5: 217-220.
  • Camarate Franca, J. and Zbyszewski, G. 1963. Geological map of Portugal at a scale of 1:50000: explanatory news of the sheet 26-B (Alcobaca). Servicos Geologicos de Portugal, Lisboa.
  • Camarate Franca, J., Zbyszewski, G., and Almeida, F.M. 1960. Geeologic map of Portugal at a scale of 1:50 000: explanatory news of the sheet 26-C (Peniche). Servicos Geologicos de Portugal, Lisboa.
  • Canale, J.I., Apesteguia, S., Gallina, P.A., Gianechini, F.A., and Haluza, A. 2016. The oldest theropods from the Neuquen Basin: Predatory dinosaur diversity from the Bajada Colorada Formation (Lower Cretaceous: Berriasian-Valanginian), Neuquen, Argentina. Cretaceous Research 71: 63-78.
  • Canale, J.I., Novas, F.E., Salgado, L., and Coria, R.A. 2014. Cranial ontogenetic variation in Mapusaurus roseae (Dinosauria: Theropoda) and the probable role of heterochrony in carcharodontosaurid evolution. Palaontologische Zeitschrift 89: 983-993.
  • Candeiro, C.R., Gil, L.M., and de Castro, P.E.P. 2018. Large-sized theropod Spinosaurus: an important component of the carnivorous dinosaur fauna in southern continents during the Cretaceous. Bulletin de la Societe Geologique de France, Earth Sciences Bulletin 189: 1-9.
  • Canudo, J.I. and Ruiz-Omenaca, J.I. 2003. Direct remains of theropod dinosaurs (excluding birds) in Spain. In: F.P. Lorent, M.M.R. Molina, P.R. Carrera (eds.), Dinosaurios y otros reptiles mesozoicos en Espana, Ciencias de la Tierra 26, 347-373. Universidad de La Rioja, Instituto de Estudios Riojanos, La Rioja.
  • Canudo, J.I., Gasulla, J.M., Gomez-Fernandez, D., Ortega, F., Sanz, J.L., and Yague, P. 2008. Primera evidencia de dientes aislados atribuidos a Spinosauridae (Theropoda) en el Aptiano inferior (Cretacico Inferior) de Europa: Formacion Arcillas de Morella (Espana). Ameghiniana 45: 649-662.
  • Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227-264.
  • Carpenter, K., Miles, C.A., and Cloward, K. 2001. New primitive stegosaur from the Morrison Formation, Wyoming. In: K. Carpenter (ed.), The Armored Dinosaurs, 55-75. Indiana University Press, Bloomington.
  • Carpenter, K., Miles, C., Ostrom, J.H., and Cloward, K. 2005. Redescription of the small maniraptoran theropods Ornitholestes and Coelurus from the Upper Jurassic Morrison Formation of Wyoming. In: K. Carpenter (ed.). The Carnivorous Dinosaurs, 49-71. Indiana University Press, Bloomington.
  • Carrano, M.T., Benson, R.B.J., and Sampson, S.D. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10: 211-300.
  • Carrington da Costa, J. 1940. A new chelonian fossil. Comunicacoes dos Servicos Geologicos de Portugal 21: 107-125.
  • Carvalho, A.M.G. and Santos, V.F. 1993. Dinosaur footprints from Sesimbra. Sesimbra Cultural 3: 10-14.
  • Charig, A.J. and Milner, A.C. 1997. Baryonyx walkeri, a fish-eating dinosaur from the Wealden of Surrey. Bulletin of the Natural History Museum. Geology Series 53: 11-70.
  • Choffat, P. 1904. The Cretaceous of the Arrabida and Ericeira regions. Comunicacoes de Comissao dos Servicos Geologicos de Portugal 6: 1-44.
  • Cobos, A. and Gasco, F. 2013. New vertebral remains of the stegosaurian dinosaur Dacentrurus from Riodeva (Teruel, Spain). Geogaceta 53: 17-20.
  • Cobos, A., Lockley, M.G., Gasco, F., Royo-Torres, R., and Alcala, L. 2014. Megatheropods as apex predators in the typically Jurassic ecosystems of the Villar del Arzobispo Formation (Iberian Range, Spain). Palaeogeography, Palaeoclimatology, Palaeoecology 399: 31-41.
  • Cobos, A., Royo-Torres, R., Luque, L., Alcala, L., and Mampel, L. 2010. An Iberian stegosaurs' paradise: the Villar del Arzobispo Formation (Tithonian-Berriasian) in Teruel (Spain). Palaeogeography, Palaeoclimatology, Palaeoecology 293: 223-236.
  • Cope, E.D. 1869. Synopsis of the extinct Batrachia, Reptilia and Aves of North America. Transactions of the American Philosophical Society 14 (1): 1-252.
  • Costa, F. and Mateus, O. 2019. Dacentrurine stegosaurs (Dinosauria): A new specimen of Miragaia longicollum from the Late Jurassic of Portugal resolves taxonomical validity and shows the occurrence of the clade in North America. PLoS ONE 14: e0224263.
  • Cuesta, E., Ortega, F., and Sanz, J.L. 2019. Axial osteology of Concavenator corcovatus (Theropoda; Carcharodontosauria) from the Lower Cretaceous of Spain. Cretaceous Research 95: 106-120.
  • Cuesta, E., Vidal, D., Ortega, F., and Sanz, J.L. 2018. The cranial osteology of Concavenator corcovatus (Theropoda; Carcharodontosauria) from the Lower Cretaceous of Spain. Cretaceous Research 91: 176-194.
  • Currie, P.J. 1995. New information on the anatomy and relationships of Dromaeosaurus albertensis (Dinosauria: Theropoda). Journal of Vertebrate Paleontology 15: 576-591.
  • Currie, P.J. and Zhao, X.-J. 1993. A new carnosaur (Dinosauria, Theropoda) from the Jurassic of Xinjiang, People's Republic of China. Canadian Journal of Earth Sciences 30: 2037-2081.
  • Currie, P.J., Rigby, J.K.J., and Sloan, R.E. 1990. Theropod teeth from the Judith River Formation of southern Alberta, Canada. In: K. Carpenter and P.J. Currie (eds.), Dinosaur Systematics: Approaches and Perspec- tives, 107-125. Cambridge University Press, Cambridge.
  • Dantas, P. 1990. Dinosaurs from Portugal. Gaia 2: 17-26.
  • Dantas, P., dos Santos, V.F., Lockley, M.G., and Meyer, C.A. 1994. Footprint evidence for limping dinosaurs from the Upper Jurassic of Portugal. Gaia 10: 43-48.
  • Dal Sasso, C., Maganuco, S., Buffetaut, E., and Mendez, M.A. 2005. New information on the skull of the enigmatic theropod Spinosaurus, with remarks on the size and affinities. Journal of Vertebrate Paleontology 25: 888-896.
  • Day, J.J. and Barrett, P.M. 2004. Material referred to Megalosaurus (Dinosauria: Theropoda) from the Middle Jurassic of Stonesfield, Oxfordshire, England: one taxon or two? Proceedings of the Geologists' Association 115: 359-366.
  • D'Emic, M.D. 2012. The early evolution of titanosauriform sauropod dinosaurs. Zoological Journal of the Linnean Society 166: 624-671.
  • Dinis, J.L., Rey, J., Cunha, P.P., Callapez, P., and Pena dos Reis, R. 2008. Stratigraphy and allogenic controls of the western Portugal Cretaceous: an updated synthesis. Cretaceous Research 29: 772-780.
  • Dollo, L. 1888. Iguanodontidae et Camptonotidae. Comptes Rendus de l'Academie des Sciences 106: 775-777.
  • Escaso, F. 2014. Evolutionary History of the Ornithischia (Dinosauria) From the Upper Jurassic of Portugal. 289 pp. Unpublished Ph.D. Thesis, Universidad Autonoma de Madrid, Madrid.
  • Escaso, F., Malafaia, E., Mocho, P., Narvaez, I., and Ortega, F. 2017. "Megalosaurus insignis" from Praia de Areia Branca (Lourinha, Portugal): is it Theropoda or Ornithopoda? In: A.B. Diaz, F. Barroso-Barcenilla, and P.C. Tonicher (eds.), Livro de Resumos da XXII Bienal da Real Sociedad Espanola de Historia Natural, 227. Real Sociedad Espanola de Historia Natural, Madrid.
  • Escaso, F., Ortega, F., Dantas, P., Malafaia, E., Silva, B., Gasulla, J.M., Mocho, P., Narvaez, I., and Sanz, J.L. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102-1112.
  • Escaso, F., Ortega, F., Dantas, P., Malafaia, E., Pimentel, N.L., Pereda-Suberbiola, X., Sanz, J.L., Kullberg, J.C., Kullberg, M.C., and Barriga, F. 2007. New evidence of shared dinosaur across Upper Jurassic Proto- North Atlantic: Stegosaurus from Portugal. Naturwissenschaften 94: 367-374.
  • Evers, S.W. and Wings, O. 2020. Late Jurassic theropod dinosaur bones from the Langenberg Quarry (Lower Saxony, Germany) provide evidence for several theropod lineages in the central European archipelago. PeerJ 8: e8437.
  • Evers, S.W., Rauhut, O.W.M., Milner, A.C., McFeeters, B., and Allain, R. 2015. A reappraisal of the morphology and systematic position of the theropod dinosaur Sigilmassasaurus from the "middle" Cretaceous of Morocco. PeerJ 3: e1323.
  • Figueiredo, S.D., de Carvalho, C.N., Cunha, P.P., and Sousa Carvalho, I.S. 2021. New dinosaur tracks from the Lower Barremian of Portugal (Areia do Mastro Formation, Cape Espichel). Journal of Geoscience and Environment Protection 9: 84-96.
  • Figueiredo, S., Rosina, P., and Figuti, L. 2015. Dinosaurs and other vertebrates from the Papo-Seco Formation (Lower Cretaceous) of southern Portugal. Journal of Iberian Geology 41: 301-314.
  • Figueiredo, S., Rosina, P., and Gomes, M. 2018. New remains of a avian dinosaur from the Lower Cretaceous of Portugal (Areias do Mastro- Sesimbra). Revista de Geo-Historia e Pre-Historia 2: 66-69.
  • Figueiredo, S., Strantzali, I.B., Gomes, M., Pimenta, R., and Santos, M. 2017. Preliminary data of new dinosaurs and turtles remains from the basal deposits of the Papo-Seco Formation in Areias do Mastro´s (Cabo Espichel, Portugal). Arnava 6 (1): 117-130.
  • Fitzinger, L. 1843. Systema Reptilium. Fasciculus primus Amblyglossae. Vindobonae. 106 + vi pp. Braumuller und Seidel, Viena.
  • Fraas, E. 1901. The sea crocodiles (Thalattosuchia n. g.) are a new saurian group from the Jurassic formation. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg 57: 409-418.
  • Fursich, F.T., Schneider, S., Werner, W., Lopez-Mir, B., and Pierce, C.S. 2021. Life at the continental-marine interface: palaeoenvironments and biota of the Alcobaca Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended. Palaeobiodiversity and Palaeoenvironments 102: 265-329.
  • Galton, P.M. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa. Postcranial skeleton. Palaontologische Zeitschrift 55: 271-312.
  • Galton P.M. 1991. Postcranial remains of the stegosaurian dinosaur Dacentrurus from the Upper Jurassic of France and Portugal. Geologica et Paleontologica 25: 299-327.
  • Galton, P.M. 1994. Notes on Dinosauria and Pterodactylia from the Cretaceous of Portugal. Neues Jahrbuch fur Geologie and Palaontologie Abhandlungen 194: 253-267.
  • Galton, P.M. 1996. Notes on Dinosauria from the Upper Cretaceous of Portugal. Neues Jahrbuch fur Geologie and Palaontologie Monatshefte 1996 (2): 83-90.
  • Galton, P.M. and Jensen, J.A. 1979. A new large theropod dinosaur from the Upper Jurassic of Colorado. Brigham Young University Geology Studies 26: 1-12.
  • Galton, P.M. and Upchurch, P. 2004. Stegosauria. In: D.B. Weishampel, P. Dodson, and H. Osmolska (ed.), The Dinosauria, 2nd Edition, 343-362. University of California Press, Berkeley.
  • Gasco, F., Cobos, A., Royo-Torres, R., Alcala, L. and Mampel, L. 2012. Theropod teeth diversity from Villar del Arzobispo Formation (Tithonian-Berriasian) at Riodeva (Teruel, Spain). Palaeobiodiversity and Palaeoenvironments 92: 273-286.
  • Gauthier, J. 1986. Saurischian monophyly and the origin of birds. In: K. Padian (ed.), The Origin of Birds and the Evolution of Flight. Memoirs of the California Academy of Sciences 8: 1-55.
  • Gerke, O. and Wings, O. 2016. Multivariate and cladistic analyses of isolated teeth reveal sympatry of theropod dinosaurs in the Late Jurassic of Northern Germany. PLoS ONE 11: e0158334.
  • Gianechini, F.A., Makovicky, P.J., and Apesteguia, S. 2011. The teeth of the unenlagiine theropod Buitreraptor from the Cretaceous of Patagonia, Argentina, and the unusual dentition of the Gondwanan dromaeosaurids. Acta Palaeontologica Polonica 56: 279-290.
  • Gilmore, C.W. 1909. Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and a description of two new species. Proceedings of the United States National Museum 36: 197-332.
  • Gilmore, C.W. 1914. Osteology of the armoured Dinosauria in the United States National Museum, with special reference to the genus Stegosaurus. Bulletin of the United States National Museum 89: 1-143.
  • Gilmore, C.W. 1920. Osteology of the carnivorous Dinosauria in the Unit- ed States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus. Bulletin of the United States National Museum 110: 1-154.
  • Goloboff, P.A. and Morales, M.E. 2023. TNT version 1.6, with a graphical interface for MacOS and Linux, including new routines in parallel. Cladistics 39: 144-153.
  • Hammer, O., Harper, D.A.T., and Ryan, P.D. 2001. Past: Paleontological Statistics Software Package for education and data analysis. Palaeontologia Electronica 4 (1): 1-9.
  • Hanson, M. and Makovicky, P.J. 2014. A new specimen of Torvosaurus tanneri originally collected by Elmer Riggs. Historical Biology 26: 775-784.
  • Harris, J.D. 1998. A reanalysis of Acrocanthosaurus atokensis, its phylogenetic status, and paleobiogeographic implications, based on a new specimen from Texas. Bulletin of the New Mexico Museum of Natural History & Science 13: 1-75.
  • Hay, O.P. 1930. Second Bibliography and Catalogue of the Fossil Vertebrata of North America. Volume 2. 990 pp. Carnegie Institute, Washington.
  • Hendrickx, C. and Mateus, O. 2014. Torvosaurus gurneyi n. sp., the largest terrestrial predator from Europe, and a proposed terminology of the maxilla anatomy in nonavian theropods. PLoS ONE 9: e88905.
  • Hendrickx, C., Mateus, O., and Araujo, R. 2015a. A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology 35: e982797.
  • Hendrickx, C., Mateus, O., and Araujo, R. 2015b. The dentition of megalosaurid theropods. Acta Palaeontologica Polonica 60: 627-642.
  • Hendrickx, C., Mateus, O., Araujo, R., and Choiniere, J. 2019. The distribution of dental features in non-avian theropod dinosaurs: Taxonomic potential, degree of homoplasy, and major evolutionary trends. Palaeontologia Electronica 22.3.74: 1-110.
  • Hendrickx, C., Stiegler, J., Currie, P.J., Han, F., Xu, X., Choiniere, J.N., and Wu, X.-C. 2020a. Dental anatomy of the apex predator Sinraptor dongi (Theropoda:Allosauroidea) from the Late Jurassic of China. Canadian Journal of Earth Sciences 57: 1127-1147.
  • Hendrickx, C., Tschopp, E. and Ezcurra, M.D. 2020b. Taxonomic identification of isolated theropod teeth: the case of the shed tooth crown associated with Aerosteon (Theropoda: Megaraptora) and the dentition of Abelisauridae. Cretaceous Research 108: 104312.
  • Hennig, E. 1915. Kentrosaurus aethiopicus, the stegosauridae of Tendaguru. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1915: 219-247.
  • Hill, G. 1988. The Sedimentology and Lithostratigraphy of the Upper Jurassic Lourinha Formation, Lusitanian Basin, Portugal. 292 pp. Unpublished Ph.D. Thesis, Open University, Milton Keynes.
  • Huberty, C.J. and Olejnik, S. 2006. Applied MANOVA and Discriminant Analysis. 2nd Edition. John Wiley & Sons, Hoboken.
  • Ibrahim, N., Sereno, P.C., Dal Sasso, C., Maganuco, S., Fabbri, M., Martill, D.M., Zouhri, S., Myhrvold, N., and Iurino, D.A. 2014. Semiaquatic adaptations in a giant predatory dinosaur. Science 345: 1613-1616.
  • Irmis, R.B. 2007. Axial skeleton ontogeny in the Parasuchia (Archosauria: Pseudosuchia) and its implications for ontogenetic determination in archosaurs. Journal of Vertebrate Paleontology 27: 350-361.
  • Isasmendi, E., Navarro-Lorbes, P., Saez-Benito, P., Viera, L.I., Torices, A., and Pereda-Suberbiola, X. 2023. New contributions to the skull anatomy of spinosaurid theropods: Baryonychinae maxilla from the Early Cretaceous of Igea (La Rioja, Spain). Historical Biology 35: 909-923.
  • Johnson, M.M., Young, M.T., and Brusatte, S.L. 2020. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 8: e9808.
  • Jonet, S. 1981. Contribution to the study of vertebrates from the Portuguese Cretaceous and especially from the Cenomanian of Extremadura. Comunicacoes dos Servicos Geologicos de Portugal 67: 10l-306.
  • Kellner, A.W.A., Azevedo, S.A.K., Machado, E.B., de Carvalho, L.B., and Henriques, D.D.R. 2011. A new dinosaur (Theropoda, Spinosauridae) from the Cretaceous (Cenomanian) Alcantara Formation, Cajual Island, Brazil. Anais da Academia Brasileira de Ciencias 83 (1): 99-108.
  • Kullberg, J.C., Rocha, R.B., Soares, A.F., Rey, J., Terrinha, P., Azeredo, A.C., Callapez, P., Duarte, L.V., Kullberg, M.C., Martins, L., Miranda, J.R., Alves, C., Mata, J., Madeira, J., Mateus, O., Moreira, M., and Nogueira, C.R. 2013. The Lusitanian Basin: stratigraphy, paleogeography and tectonics. In: R. Dias, A.Araujo, P. Terrinha, and J.C. Kullberg (eds.), Geologia de Portugal no contexto da Iberia, 989-1141. Escolar Editora, Lisboa.
  • Kullberg, J.C., Rocha, R.B., Soares, A.F., Rey, J., Terrinha, P., Callapez, P., and Martins, L. 2006. The Lusitanian Basin: stratigraphy, paleogeography and tectonics. In: R. Dias, A. Araujo, P. Terrinha, and J.C. Kullberg (eds.), Geologia de Portugal no contexto da Iberica, 317-368. Universidade de Evora, Evora.
  • Lapparent, A.F. and Zbyszewski, G. 1957. The dinosaurs from Portugal. Memoire Services Geologiques du Portugal 2 (nouvelle serie): 1-131.
  • Larson, D.W. 2008. Diversity and variation of theropod dinosaur teeth from the uppermost Santonian Milk River Formation (Upper Cretaceous), Alberta: a quantitative method supporting identification of the oldest dinosaur tooth assemblage in Canada. Canadian Journal of Earth Sciences 45: 1455-1468.
  • Leinfelder, R.R. 1987. Multifactorial control of sedimentation patterns in an ocean marginal basin: the Lusitanian Basin (Portugal) during the Kimmeridgian and Tithonian. Geologische Rundschau 76: 599-631.
  • Li, F., Peng, G., Yong, Y., Jiang, S., and Huang, D. 2009. A new carnosaur from the Late Jurassic of Qianwei, Sichuan, China. Acta Geologica Sinica 83: 1203-1214.
  • Lockley, M.G., Meyer, C.A., and dos Santos, V.F. 1994. Trackway evidence for a herd of juvenile sauropods from the Late Jurassic of Portugal. Gaia 10: 27-35.
  • Long, J.A. and McNamara, K.J. 1997. Heterochrony. In: P.J. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs, 311-317. Academic Press, San Diego.
  • Lubbe, T., Richter, U., and Knotschke, N. 2009. Velociraptorine dromaeosaurid teeth from the Kimmeridgian (Late Jurassic) of Germany. Acta Palaeontologica Polonica 54: 401-408.
  • Maddison, W.P. and Maddison, D.R. 2011. Mesquite: A Modular System for Evolutionary Analysis. Version 2.75 [available online, http://mesquiteproject.org].
  • Madsen, J.H., Jr. 1976a. Allosaurus fragilis: a revised osteology. Utah Geological Survey Bulletin 109: 1-163.
  • Madsen, J.H., Jr. 1976b. A second new theropod dinosaur from the Late Jurassic of east central Utah. Utah Geology 3 (1): 51-60.
  • Madsen, J.H., Jr. and Welles, S.P. 2000. Ceratosaurus (Dinosauria, Theropoda). A revised osteology. Miscellaneous Publication, Utah Geological Survey 00-2: 1-80.
  • Maidment, S.C.R., Brassey, C., and Barrett, P.M. 2015. The postcranial skeleton of an exceptionally complete individual of the plated dinosaur Stegosaurus stenops (Dinosauria: Thyreophora) from the Upper Jurassic Morrison Formation of Wyoming, U.S.A. PLoS ONE 10: e0138352.
  • Maidment, S.C.R., Chapelle, K.E.J., Bonsor, J.A., Button, D., and Barrett, P. 2022. Osteology and relationship of Cumnoria prestwichii (Ornithischia: Ornithopoda) from the Late Jurassic of Oxfordshire, UK. Monograph of the Palaeontographical Society 176: 1-55.
  • Maidment, S.C.R., Raven, T.J., Ouarhache, D., and Barrett, P.M. 2020. North Africa's first stegosaur: Implications for Gondwanan thyreophoran dinosaur diversity. Gondwana Research 77: 82-97.
  • Malafaia, E., Escaso, F., Mocho, P., Serrano-Martinez, A., Torices, A., Cachao, M., and Ortega, F. 2017a. Analysis of diversity, stratigraphic and geographical distribution of isolated theropod teeth from the Upper Jurassic of the Lusitanian Basin, Portugal. Journal of Iberian Geology 43: 257-291.
  • Malafaia, E., Gasulla, J.M., Escaso, F., Narvaez, I., and Ortega, F. 2020a. An update of the spinosaurid (Dinosauria: Theropoda) fossil record from the Lower Cretaceous of the Iberian Peninsula: distribution, diversity, and evolutionary history. Journal of Iberian Geology 46: 431-444.
  • Malafaia, E., Mocho, P., Escaso, F., and Ortega, F. 2020b. A new carcharodontosaurian theropod from the Lusitanian Basin: evidence of allosauroid sympatry in the European Late Jurassic. Journal of Vertebrate Paleontology 40 (1): e1768106.
  • Malafaia, E., Mocho, P., Escaso, F., and Ortega, F. 2017b. A juvenile allosauroid theropod (Dinosauria, Saurischia) from the Upper Jurassic of Portugal. Historical Biology 29: 654-676.
  • Malafaia, E., Mocho, P., Escaso, F., and Ortega, F. 2017c. New data on the anatomy of Torvosaurus and other remains of megalosauroid (Dinosauria, Theropoda) from the Upper Jurassic of Portugal. Journal of Iberian Geology 43: 33-59.
  • Malafaia, E., Ortega, F., Escaso, F. and Silva, B. 2015. New evidence of Ceratosaurus (Dinosauria: Theropoda) from the Late Jurassic of the Lusitanian Basin, Portugal. Historical Biology 27: 938-946.
  • Mannion, P.D. and Calvo, J.O. 2011. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 163: 155-181.
  • Mannion, P.D., Upchurch, P., Barnes, R.N., and Mateus, O. 2013. Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriformes. Zoological Journal of the Linnean Society 168: 98-206.
  • Mannion, P.D., Upchurch, P., Schwarz, D., and Wings, O. 2019. Taxonomic affinities of the putative titanosaurs from the Late Jurassic Tendaguru Formation of Tanzania: phylogenetic and biogeographic implications for eusauropod dinosaur evolution. Zoological Journal of the Linnean Society 185: 784-909.
  • Manuppella, G., Antunes, M.T., Costa Almeida, C.A., Azeredo, A.C., Barbosa, B., Cardoso, J.L., Crispim, J.A., Duarte, L.V., Henriques, M.H., Martins, L.T., Ramalho, M.M., Santos, V.F., and Terrinha, P. 2000. Geological Map of Portugal at Scale of 1:50 000: Explanatory Notice of the Sheet 27-A (Vila Nova de Ourem). Departamento de Geologia, Instituto Geologico e Mineiro, Lisboa.
  • Manuppella, G., Antunes, M.T., Pais, J., Ramalho, M.M., and Rey, J. 1999a. Geological Map of Portugal at Scale of 1:50 000: Explanatory Notice of the Sheet 30-A (Lourinha). Departamento de Geologia, Instituto Geologico e Mineiro, Lisboa.
  • Manuppella, G., Antunes, M.T., Pais, J., Ramalho, M.M., and Rey, J. 1999b. Geological Map of Portugal at Scale of 1:50 000: Explanatory Notice of the Sheet 38-B (Setubal). Departamento de Geologia, Instituto Geologico e Mineiro, Lisboa.
  • Manuppella, G., Zbyszewski, G., and Veiga Ferreira, O. 1978. Geological Map of Portugal at Scale of 1:50 000: Explanatory Notice of the Sheet 23-A (Pombal). Servicos Geologicos de Portugal, Lisboa.
  • Marsh, O.C. 1877a. New order of extinct Reptilia (Stegosauria) from the Jurassic of the Rocky Mountains. American Journal of Science 14 (Series 3): 513-514.
  • Marsh, O.C. 1877b. Notice of new dinosaurian reptiles from the Jurassic Formation. American Journal of Science 14 (Series 3): 514-516.
  • Marsh, O.C. 1878. Principal characters of American Jurassic dinosaurs. Part I. American Journal of Science 16 (Series 3): 411-416.
  • Marsh, O.C. 1881. Principal characters of American Jurassic dinosaurs. Part V. American Journal of Science 21 (Series 3): 417-423.
  • Marsh, O.C. 1884. Principal characters of American Jurassic dinosaurs. Part VIII. The order Theropoda. American Journal of Science 27 (Series 3): 329-340.
  • Martin, T. and Krebs, B. 2000. Guimarota. A Jurassic Ecosystem. 155 pp. Verlag Dr. Friedrich Pfeil, Munchen.
  • Mateus, O. 1998. Lourinhanosaurus antunesi, a new Upper Jurassic allosauroid (Dinosauria: Theropoda) from Lourinha, Portugal. Memorias da Academia das Ciencias de Lisboa 37: 111-124.
  • Mateus, O. 2005. Dinosaurs of the Upper Jurassic of Portugal, with Emphasis on Saurischians. 377 pp. Unpublished Ph.D. Thesis, Universidade Nova de Lisboa, Lisboa.
  • Mateus, O. and Antunes, M.T. 2000a. Ceratosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. In: Abstracts, 31st Inter- national Geological Congress. Rio de Janeiro.
  • Mateus, O. and Antunes, M.T. 2000b. Torvosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. In: Abstracts Book I Congresso Iberico de Paleontologia, XVI Jornadas de la Sociedad Espanola de Paleontologia, 115-117. Universidade de Evora, Evora.
  • Mateus, O. and Estraviz-Lopez, D. 2022. A new theropod dinosaur from the early Cretaceous (Barremian) of Cabo Espichel, Portugal: Implications for spinosaurid evolution. PLoS ONE 17: e0262614.
  • Mateus, O., Araujo, R., Natario, C., and Castanhinha, R. 2011. A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa 2827: 54-68.
  • Mateus, O., Dinis, J., and Cunha, P. P. 2017. Upper Jurassic to Lowermost Cretaceous of the Lusitanian Basin, Portugal-landscapes where dinosaurs walked. Ciencias da Terra 34: 618-634.
  • Mateus, O., Maidment, S.C.R., and Christiansen, N.A. 2009. A new longnecked "sauropod-mimic" stegosaur and the evolution of the plated dinosaurs. Proceedings of the Royal Society of London Series B 276: 1815-1821.
  • Mateus, O., Mannion, P.D., and Upchurch, P. 2014. Zby atlanticus, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 618-634.
  • Mateus, O., Walen, A., and Antunes, M.T. 2006. The large theropod fauna of the Lourinha Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of Allosaurus. New Mexico Museum of Natural History and Science Bulletin 36: 123-129.
  • Mocho, P., Royo-Torres, R., and Ortega, F. 2014. Phylogenetic reassessment of Lourinhasaurus alenquerensis, a basal Macronaria (Sauropoda) from the Upper Jurassic of Portugal. Zoological Journal of the Linnean Society 170: 875-916.
  • Mocho, P., Royo-Torres, R., and Ortega, F. 2017a. New data of the Portuguese brachiosaurid Lusotitan atalaiensis (Sobral Formation, Upper Jurassic). Historical Biology 29: 789-817.
  • Mocho, P., Royo-Torres, R., and Ortega, F. 2019. A new macronarian sauropod from the Upper Jurassic of Portugal. Journal of Vertebrate Paleontology 39 (1): e1578782.
  • Mocho, P., Royo-Torres, R., Escaso, F., Malafaia, E., de Miguel Chaves, C., Narvaez, I., Perez-Garcia, A., Pimentel, N., Silva, B.C., and Ortega, F. 2017b. Upper Jurassic sauropod record in the Lusitanian Basin (Portugal): Geographical and lithostratigraphical distribution. Palaeontologia Electronica 20.2.27A: 1-50.
  • Mocho, P., Royo-Torres, R., Malafaia, E., Escaso, F., and Ortega, F. 2016. Systematic review of Late Jurassic sauropods from the Museu Geologico collections (Lisboa, Portugal). Journal of Iberian Geology 42: 227-250.
  • Montenat, C., Guery, F., and Jamet, M. 1988. Mesozoic evolution of the Lusitanian Basin: comparison with the adjacent margin. In: G. Boillot, E.L. Winterer et al. (ed.), Proceedings of the Ocean Drilling Program, Scientific Results 103: 757-775.
  • Nopcsa, F. 1915. The dinosaurs of the Transylvanian parts of Hungary. Mitteilungen aus dem Jahrbuche der Kgl. Ungarischen Geologischen Reichanstalt 23: 1-24.
  • Novas, F.E., de Valais, S., Vickers-Rich, P., and Rich, T. 2005. A large Cretaceous theropod from Patagonia, Argentina, and the evolution of carcharodontosaurids. Naturwissenschaften 92: 226-230.
  • Oliveira, J.T., Pereira, H., Ramalho, M., and Antunes, M.T. 1992. Geological Map of Portugal, at the Scale of 1:500 000. Servicos Geologicos de Portugal, Lisboa.
  • Ortega, F., Escaso, F., Gasulla, J.M., Dantas, P., and Sanz, J.L. 2006. Dinosaurs of the Iberian Peninsula. Estudios Geologicos 62 (1): 219-240.
  • Ortega, F., Malafaia, E., Escaso, F., Perez-Garcia, A., and Dantas, P. 2009. Faunas of reptiles from the Upper Jurassic of Portugal. Paleolusitana 1: 43-56.
  • Osborn, H.F. and Mook, C.C. 1921. Camarasaurus, Amphicoelias, and other sauropods of Cope. Memoirs of the American Museum of Natural History New Series 3: 249-387.
  • Owen, R. 1842. Report on British fossil reptiles. Part II. Reports of the British Association for the Advancement of Sciences 11: 60-204.
  • Paul, G.S. 1988. Predatory Dinosaurs of the World: A Complete Illustrated Guide. 464 pp. Simon & Schuster, New York.
  • Pena dos Reis, R.P.B., Dinis, J.L., Cunha, P.M.R., and Trincao, P. 1996. Upper Jurassic sedimentary infill and tectonics of the Lusitanian Basin (western Portugal). In: A.C. Riccardi (ed.), Advances in Jurassic Research, GeoResearch Forum 1-2, 377-386. Transtec Publications, Zurich.
  • Pena dos Reis, R.P.B., Pimentel, N.L., and Garcia, A.J.V. 2011. The Lusitanian Basin, stratigraphic analysis and geodynamic evolution. Boletim de Geociencias Petrobras 19: 23-52.
  • Pena dos Reis, R.P.B., Proenca Cunha, C.P., Dinis, J.L., and Trincao, P.R. 2000. Geologic evolution of the Lusitanian Basin (Portugal) during the late Jurassic. GeoResearch Forum 6: 345-356.
  • Perez-Moreno, B.P., Chure, D.J., Pires, C., Silva, C.M.D., Santos, V.D., Dantas, P., Povoas, L., Cachao, M., Sanz, J.L., and Galopim de Carvalho, A.M. 1999. On the presence of Allosaurus fragilis (Theropoda: Carnosauria) in the Upper Jurassic of Portugal: first evidence of an intercontinental dinosaur species. Journal of the Geological Society 156: 449-452.
  • Rasmussen, E.S., Lomholt, S., Andersen, C., and Vejbaek, O.V. 1998. Aspects of the structural evolution of the Lusitanian Basin in Portugal and the shelf and slope area offshore Portugal.Tectonophysics 300: 199-225.
  • Rauhut, O.W.M. 2004. Provenance and anatomy of Genyodectes serus, a large-toothed ceratosaur (Dinosauria: Theropoda) from Patagonia. Journal of Vertebrate Paleontology 24: 894-902.
  • Rauhut, O.W.M. 2005. Osteology and relationships of a new theropod dinosaur from the Middle Jurassic of Patagonia. Palaeontology 48: 87-110.
  • Rauhut, O.W.M. 2011. Theropod dinosaurs from the Late Jurassic of Tendaguru (Tanzania). Special Papers in Palaeontology 86: 195-239.
  • Rauhut, O.W.M. and Carrano, M.T. 2016. The theropod dinosaur Elaphrosaurus bambergi Janensch, 1920, from the Late Jurassic of Tendaguru, Tanzania. Zoological Journal of the Linnean Society 178: 546-610.
  • Rauhut, O.W.M. and Lopez-Arbarello, A. 2009. Considerations on the age of the Tiouaren Formation (Iullemmeden Basin, Niger, Africa): Implications for Gondwanan Mesozoic terrestrial vertebrate faunas. Palaeogeography, Palaeoclimatology, Palaeoecology 271: 259-267.
  • Rauhut, O.W.M. and Pol, D. 2019. A probable basal allosauroid from the early Middle Jurassic Canadon Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs. Scientific Reports 9: 18826.
  • Rauhut, O.W.M., Foth, C., Tischlingerd, H., and Norell, M.A. 2012. Exceptionally preserved juvenile megalosauroid theropod dinosaur with filamentous integument from the Late Jurassic of Germany. PNAS 109: 11746-11751.
  • Rauhut, O.W.M., Hubner, T.R., and Lanser, K.-P. 2016. A new megalosaurid theropod dinosaur from the late Middle Jurassic (Callovian) of north-western Germany: Implications for theropod evolution and faunal turnover in the Jurassic. Palaeontologia Electronica 19.2.26A: 1-65.
  • Rauhut, O.W.M., Pinuela, L., Castanera, D., Garcia-Ramos, J.C., and Cela, I.S. 2018. The largest European theropod dinosaurs: remains of a gigantic megalosaurid and giant theropod tracks from the Kimmeridgian of Asturias, Spain. PeerJ 6: e4963.
  • Rauhut, O.W.M., Schwermann, A.H., Hubner, T.R., and Lanser, K.-P. 2020. The oldest record of the genus Torvosaurus (Theropoda: Megalosauridae) from the Callovian Ornatenton Formation of north-western Germany. Geologie und Palaontologie in Westfalen 93: 1-13.
  • Rey, J. 1972. Geological research on the Lower Cretaceous of Extremadura (Portugal). Memoires du Service geologique du Portugal 21: 1-477.
  • Rey, J. 1992. The Lower Cretaceous lithostratigraphic units of the Lisbon region. Comunicacoes dos Servicos Geologicos de Portugal 78: 103-24.
  • Rey, J. 1999. Lower Cretaceous Depositional Sequences in the Cascais Area. In: European Palaeontological Association Workshop, Field Trip A, 1-57. Lisboa.
  • Rey, J., Graciansky, P.C. de, and Jacquin, T. 2003. Depositional sequences in the Lower Cretaceous of the Lusitano Basin. Comunicacoes del Instituto Geologico e Minero 90: 15-42.
  • Ribeiro, A., Silva, J.B., Cabral, J., Dias, R., Fonseca, P., Kullberg, M.C., Terrinha, P., and Kullberg, J.C. 1996. Tectonics of the Lusitanian Basin. Final Report, Proj. MILUPOBAS, ICTE/GG/GeoFCUL. Lisboa.
  • Rocha, R.B. and Soares, A.F. 1984. Some thoughts on Jurassic sedimentation in the western Mesocenozoic rim of Portugal. Memorias e Noti- cias 97: 133-142.
  • Rocha, R.B., Manuppella, G., Mouterde, R., Ruget, C., and Zbyszewski, G. 1981. Geological Map of Portugal at the Scale of 1:50 000: Explanatory News of the Sheet 19-C (Figueira da Foz). Servicos Geologicos de Portugal, Lisboa.
  • Rocha, R.B., Marques, B.L., Kullberg, J.C., Caetano, P.C., Lopes, C., Soares, A.F., Duarte, L.V., Marques, J.F. and Gomes, C.R. 1996. The 1st and 2nd rifting phases of the Lusitanian Basin: stratigraphy, sequence analysis and sedimentary evolution. Final Report C.E.C. Proj. MILUPOBAS, Lisboa.
  • Rotatori, F.M., Moreno-Azanza, M., and Mateus, O. 2022. Reappraisal and new material of the holotype of Draconyx loureiroi (Ornithischia: Iguanodontia) provide insights on the tempo and modo of evolution of thumb-spiked dinosaurs. Zoological Journal of the Linnean Society 195: 125-156.
  • Royo-Torres, R., Cobos, A., and Alcala, L. 2006. A giant European dinosaur and a new sauropod clade. Science 314: 1925-1927.
  • Royo-Torres, R., Upchurch, P., Mannion, P.D., Mas, R., Cobos, A., Gasco, F., Alcala, L., and Sanz, J.L. 2014. The anatomy, phylogenetic relationships and stratigraphic position of the Tithonian-Berriasian Spanish sauropod dinosaur Aragosaurus ischialus. Zoological Journal of the Linnean Society 71: 623-655.
  • Ruiz-Omenaca, J.I., Pinuela, P., Garcia-Ramos, J.C., and Canudo, J.I. 2008. Carnivorous dinosaur teeth (Saurischia; Theropoda) from the Late Jurassic of Asturias, north Spain. In: Huerta Hurtado, P. and Fernandez-Baldor, F.T. (eds,), Actas de las IV Jornadas Internacionales sobre Paleontologia de Dinosaurios y su Entorno Salas de los Infantes, Burgos, 273-291. Colectivo Arqueologico y Paleontologico de Salas, Salas de los Infantes.
  • Sadleir, R., Barrett, P.M., and Powell, H.P. 2008. The anatomy and systematics of Eustreptospondylus oxoniensis, a theropod dinosaur from the middle Jurassic of Oxfordshire, England. Monograph of the Palaeontographical Society 160: 1-82.
  • Salgado, L., Coria, R.A., and Calvo, J.O. 1997. Evolution of titanosaurid sauropods. I: phylogenetic analysis based on the postcranial evidence. Ameghiniana 34: 3-32.
  • Samathi, A., Sander, P.M., and Chanthasit, P. 2021. A spinosaurid from Thailand (Sao Khua Formation, Early Cretaceous) and a reassessment of Camarillasaurus cirugedae from the Early Cretaceous of Spain. Historical Biology 33: 3480-3494.
  • Santos, V.F. 2003. Dinosaur Tracks in the Jurassic and Cretaceous of Portugal. Paleobiological and Paleoecological Considerations. 365 pp. Unpublished Ph.D. Thesis, Facultad de Ciencias, Universidad Autonoma de Madrid, Madrid.
  • Santos, V.F. 2008. Dinosaur Footprints from Portugal. 124 pp. Museu Nacional de Historia Natural, Lisboa.
  • Seeley, H.G. 1887. On the classification of the fossil animals commonly named Dinosauria. Proceedings of the Royal Society of London 43: 165-171.
  • Sauvage, H.E. 1896. Crocodilians and dinosaurians from the Mesozoic of Portugal. Bulletin de la Societe geologique de France 24: 46-49.
  • Sauvage, H.E. 1897-1898. Fossil vertebrates from Portugal. Contributions to the study of fish and reptiles from the Jurassic and Cretacic. Direction des Travaux Geologiques du Portugal 29: 1-46.
  • Sereno, P.C. 1998. A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen 210: 41-83.
  • Sereno, P.C., Beck, A.L., Dutheil, D.B., Gado, B., Larsson, H.C.E., Lyon, G.H., Marcot, J.D., Rauhut, O.W.M., Sadleir, R.W., Sidor, C.A., Varricchio, D.D., Wilson, G.P., and Wilson, J.A. 1998. A long-snouted predatory dinosaur from Africa and the evolution of spinosaurids. Science 282: 1298-1302.
  • Serrano-Martinez, A., Vidal, D., Sciscio, L., Ortega, F., and Knoll, F. 2015. Isolated theropod teeth from the Middle Jurassic of Niger and the early dental evolution of Spinosauridae. Acta Palaeontologica Polonica 61: 403-415.
  • Schade, M., Rauhut, O.W.M., Foth, C., Moleman, O., and Evers, S.W. 2023. A reappraisal of the cranial and mandibular osteology of the spinosaurid Irritator challengeri (Dinosauria: Theropoda). Palaeontologia Electronica 26: a17.
  • Schubert, B.W. and Ungar, P.S. 2005. Wear facets and enamel spalling in tyrannosaurid dinosaurs. Acta Palaeontologica Polonica 50: 93-99.
  • Smith, J.B., Lamanna, M.C., Mayr, H., and Lacovara, K.J. 2006. New information regarding the holotype of Spinosaurus aegyptiacus Stromer, 1915. Journal of Paleontology 80: 400-406.
  • Smith, J.B., Vann, D.R., and Dodson, P. 2005. Dental morphology and variation in theropod dinosaurs: implications for the taxonomic identification of isolated teeth. The Anatomical Record 285A: 699-736.
  • Soto, M. and Perea, D. 2008.A ceratosaurid (Dinosauria, Theropoda) from the Late Jurassic-Early Cretaceous of Urugay. Journal of Vertebrate Paleontology 28: 439-444.
  • Sues, H.-D., Frey, E., Martill, D.M., and Scott, D.M. 2002. Irritator challengeri, a spinosaurid (Dinosauria: Theropoda) from the Lower Cretaceous of Brazil. Journal of Vertebrate Paleontology 22: 535-547.
  • Taquet, P. and Welles, S.M. 1977. Redescription du crane de dinosaure theropode de Dives (Normandie). Annales de Paleontologie 63: 191- 206.
  • Taylor, A.M., Gowland, S., Leary, S., Keogh, K.J., and Martinius, A.W. 2014. Stratigraphical correlation of the Lourinha Formation in the Consolacao Sub-basin (Lusitanian Basin), Portugal. Geological Journal 49: 143-162.
  • Teixeira, C., Zbyszewski, G., Torres de Assuncao, C., and Manuppell, G. 1968. Geological Map of Portugal at the Scale of 1:50 000: Explanatory News of the Sheet 23-C (Leiria). Servicos Geologicos de Portugal, Lisboa.
  • Tschopp, E., Mateus, O., and Benson, R. 2015. A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ 3: e857.
  • Upchurch, P. 1995. Evolutionary history of sauropod dinosaurs. Philosophical Transactions of the Royal Society of London B 349: 365-390.
  • Upchurch, P., Barrett, P.M., and Dodson, P. 2004. Sauropoda. In: D.B. Weishampel, P. Dodson, and H. Osmolska (eds.), The Dinosauria, Second Editon, 259-322. University of California Press, Oakland.
  • Upchurch, P., Mannion, P.D., and Taylor, M.P. 2015. The anatomy and phylogenetic relationships of "Pelorosaurus" becklesii (Neosauropoda, Macronaria) from the Early Cretaceous of England. PLoS ONE 10: e0125819.
  • Verriere, A., Frobisch, N.B., and Frobisch, J. 2022. Regionalization, constraints, and the ancestral ossification patterns in the vertebral column of amniotes. Scientific Reports 12: 1-12.
  • Viera, L.I. and Torres, J.A. 1995. The presence of Baryonyx walkeri (Saurischia, Theropoda) in the Weald of La Rioja (Spain). Nota previa. MUNIBE 47: 57-61.
  • Vullo, R., Abit, D., Ballevre, M., Billon-Bruyat, J.-P., Bourgeais, R., Buffetaut, E., Daviero-Gomez, V., Garcia, G., Gomez, B., Mazin, J.-M., Morel, S., Neraudeau, D., Pouech, J., Rage, J.-C., Schnyder, J., and Tong, H. 2014. Palaeontology of the Purbeck-type (Tithonian, Late Jurassic) bonebeds of Chassiron (Oleron Island, western France). Comptes Rendus Palevol 13: 421-441.
  • Walker,A.D. 1970.A revision of the Jurassic reptile Hallopus victor (Marsh), with remarks on the classification of crocodiles. Philosophical Transactions of the Royal Society of London 257 (Series B): 323-372.
  • Wilson, J.A. 2002. Sauropod dinosaur phylogeny: critique and cladistic analysis. Zoological Journal of the Linnean Society 136: 217-276.
  • Wilson, J.A. and Sereno, P.C. 1998. Early evolution and higher-level phylogeny of sauropod dinosaurs. Journal of Vertebrate Paleontology 18 (Supplement 2): 1-68.
  • Wilson, R.C.L. 1988. Mesozoic development of the Lusitanian Basin, Portugal. Revista de la Sociedad Geologica Espanola 1: 393-407.
  • Wilson, R.C.L. 1975. Atlantic opening and Mesozoic continental margin basins of Iberia. Earth and Planetary Science Letters 25: 33-43.
  • Young, C.M.E., Hendrickx, C., Challands, T.J., Foffa, D., Ross, D.A., Butler, I.B., and Brusatte, S.L. 2019. New theropod dinosaur teeth from the Middle Jurassic of the Isle of Skye, Scotland. Scottish Journal of Geology 55: 7-19.
  • Young, M.T. and Andrade, M.B. 2009. What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia, Metriorhynchidae) from the Upper Jurassic of Bayern, Germany. Zoological Journal of the Linnean Society 157: 551-585.
  • Young, M.T., Andrade, M.B., Cornee, J.-J., Steel, L., and Foffa, D. 2014. Re-description of a putative Early Cretaceous "teleosaurid" from France, with implications for the survival of metriorhynchids and teleosaurids across the Jurassic-Cretaceous Boundary. Annales de Paleontologie 100: 165-174.
  • Zbyszewski, G. 1946. Bones of Omosaurus discovered near Baleal (Peniche). Comunicacoes dos Servicos Geologicos de Portugal 28: 135-144.
  • Zbyszewski, G. and Moitinho de Almeida, F. 1960. Geological Map of Portugal at the Scale of 1:50 000: Explanatory News of the Sheet 26-D (Caldas da Rainha). Servicos Geologicos de Portugal, Lisboa.
  • Zinke, J. 1998. Small theropod teeth from the Upper Jurassic coal mine of Guimarota (Portugal). Palaontologische Zeitschrift 72: 179-189.
  • Zittel, K.A. 1887-1890. Handbuch der Palaontologie 1. Abteilung: Palao- zoologie 3. 900 pp. Druck und Verlag von R. Oldenbourg, Munchen.