Published May 8, 2020 | Version v1
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New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata

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Czepiński, Łukasz (2020): New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata. Acta Palaeontologica Polonica 65 (3): 481-497, DOI: 10.4202/app.00701.2019, URL: http://dx.doi.org/10.4202/app.00701.2019

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References

  • Alifanov, V.R. 2012. Order Lacertilia [in Russian]. In: E.N. Kurockin and A.V. Lopatin (eds.), Iskopaemye pozwonocnye Rossii i sopredel'nyh stran. Iskopaemye reptilii i pticy. Cast' 2, 7-136, GEOS, Moskva.
  • Arbour, V.M. and Currie, P.J. 2013. The taxonomic identity of a nearly complete ankylosaurid dinosaur skeleton from the Gobi Desert of Mongolia. Cretaceous Research 46: 24-30.
  • Arbour, V.M., Currie, P.J., and Badamgarav, D. 2014. The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia. Zoological Journal of the Linnean Society 172: 631-652.
  • Averianov, A.O. and Lopatin, A.V. 2020. An unusual new sauropod dinosaur from the Late Cretaceous of Mongolia. Journal of Systematic Palaeontology [published online http://doi.org/10.1080/14772019.2020. 1716402].
  • Averianov, A.O. and Sues, H.D. 2012. Correlation of Late Cretaceous continental vertebrate assemblages in Middle and Central Asia. Journal of Stratigraphy 36: 462-485.
  • Barsbold, R. 1983. Carnivorous dinosaurs from the Cretaceous of Mongolia [in Russian]. Sovmestnaa Sovetsko-Mongol'skaa Paleontologiceskaa Ekspeditcia, Trudy 19: 1-117.
  • Benton, J.M. 2000. Mongolian place names and stratigraphic terms. In: M.J. Benton, M.A. Shishkin, D.M. Unwin, and E.N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia, xxii-xxviii. Cambridge University Press, Cambridge.
  • Bohlin, B. 1953. Fossil reptiles from Mongolia and Kansu. The Sino-Swedish Expedition, Publication 37: 1-105.
  • Brown, D.B. and Schlaikjer, D.E.M. 1940. The structure and relationships of Protoceratops. Transactions of the New York Academy of Sciences 2: 99-100.
  • Burns, M.E., Currie, P.J., Sissons, R.L., and Arbour, V.M. 2011. Juvenile specimens of Pinacosaurus grangeri Gilmore, 1933 (Ornithischia: Ankylosauria) from the Late Cretaceous of China, with comments on the specific taxonomy of Pinacosaurus. Cretaceous Research 32: 174-186.
  • Burns, M.E., Tumanova, T.A., and Currie, P.J. 2015. Postcrania of juvenile Pinacosaurus grangeri (Ornithischia: Ankylosauria) from the Upper Cretaceous Alagteeg Formation, Alag Teeg, Mongolia: implications for ontogenetic allometry in ankylosaurs. Journal of Paleontology 89: 168-182.
  • Carpenter, K., Hayashi, S., Kobayashi, Y., Maryanska, T., Barsbold, R., Sato, K., and Obata, I. 2011. Saichania chulsanensis (Ornithischia, Ankylosauridae) from the Upper Cretaceous of Mongolia. Palaeontographica Abteilung A 294: 1-61.
  • Carr, T.D., Varricchio, D.J., Sedlmayr, J.C., Roberts, E.M., and Moore, J.R. 2017. A new tyrannosaur with evidence for anagenesis and crocodile-like facial sensory system. Scientific Reports 7: 44942.
  • Chen, X. 2017. The discovery and significance of the protoceratopsid dinosaur from the Lower Cretaceous of Ordos Region, Inner Mongolia. Acta Geoscientica Sinica 38: 154-158.
  • Chinzorig, T., Kobayashi, Y., Tsogtbaatar, K., Currie, P.J., Watabe, M., and Barsbold, R. 2017. First ornithomimid (Theropoda, Ornithomimosauria) from the Upper Cretaceous Djadokhta Formation of Togrogiin Shiree, Mongolia. Scientific Reports 7: 5835.
  • Clark, J.M., Norell, M.A., and Rowe, T. 2002. Cranial anatomy of Citipati osmolskae (Theropoda, Oviraptorosauria), and a reinterpretation of the holotype of Oviraptor philoceratops. American Museum Novitates 2002: 1-24.
  • Currie, P.J. 2016. Dinosaurs of the Gobi: Following in the footsteps of the Polish-Mongolian Expeditions. Palaeontologica Polonica 67: 83-100.
  • Currie, P.J., Badamgarav, D., Koppelhus, E.B., Sissons, R., and Vickaryous, M.K. 2011. Hands, feet, and behaviour in Pinacosaurus (Dinosauria: Ankylosauridae). Acta Palaeontologica Polonica 56: 489-505.
  • Czepinski, L. 2019. Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert. Historical Biology [published online http://doi.org/10.1080/0 8912963.2019.1593404].
  • Danilov, I.G., Sukhanov, V.B., and Syromyatnikova, E.V. 2013. Redescription of Zangerlia dzamynchondi (Testudines: Nanhsiungchelyidae) from the Late Cretaceous of Mongolia, with a reassessment of the phylogenetic position and relationships of Zangerlia. In: D.B. Brinkman, P.A. Holroyd, and J.D. Gardner (eds.), Morphology and Evolution of Turtles, 407-417. Springer, Dordrecht.
  • Dashzeveg, D., Dingus, L., Loope, D.B., Swisher, C.C., Dulam, T., and Sweeney, M.R. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates 2005: 1-31.
  • Dingus, L., Loope, D.B., Dashzeveg, D., Swisher, C.C., Minjin, C., Novacek, M.J., and Norell, M.A. 2008. The geology of Ukhaa Tolgod (Djadokhta Formation, Upper Cretaceous, Nemegt Basin, Mongolia). American Museum Novitates 2008: 1-40.
  • Dodson, P. 1976. Quantitative aspects of relative growth and sexual dimorphism in Protoceratops. Journal of Paleontology, 50: 929-940.
  • Dong, L., Xu, X., Wang, Y., and Evans, S.E. 2018. The lizard genera Bainguis and Parmeosaurus from the Upper Cretaceous of China and Mongolia. Cretaceous Research 85: 95-108.
  • Dzik, J. 2005. The chronophyletic approach: stratophenetics facing an incomplete fossil record. Special Papers in Palaeontology 73: 159-183.
  • Elzanowski,A. and Wellnhofer, P. 1992.A new link between theropods and birds from the Cretaceous of Mongolia. Nature 359: 821-823.
  • Farke, A.A. 2010. Evolution, homology, and function of the supracranial sinuses in ceratopsian dinosaurs. Journal of Vertebrate Paleontology 30: 1486-1500.
  • Funston, G.F., Currie, P.J., Ryan, M.J., and Dong, Z.M. 2019. Birdlike growth and mixed-age flocks in avimimids (Theropoda, Oviraptorosauria). Scientific Reports 9: 18816.
  • Funston, G.F., Mendonca, S.E., Currie, P.J., and Barsbold, R. 2018. Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin. Palaeogeography, Palaeoclimatology, Palaeoecology 494: 101-120.
  • Gao, K. and Hou, L. 1996. Systematics and taxonomic diversity of squamates from the Upper Cretaceous Djadochta Formation, Bayan Mandahu, Gobi Desert, People's Republic of China. Canadian Journal of Earth Sciences 33: 578-598.
  • Gao, K. and Norell, M.A. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 2000: 1-118.
  • Godefroit, P., Currie, P.J., Hong, L., Yong, S.C., and Dong, Z. 2008. A new species of Velociraptor (Dinosauria: Dromaeosauridae) from the Upper Cretaceous of northern China. Journal of Vertebrate Paleontology 28: 432-438.
  • Godefroit, P., Suberbiola, X.P., Li, H., and Dong, Z. 1999, A new species of the ankylosaurid dinosaur Pinacosaurus from the late cretaceous of inner Mongolia (P.R. China): Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la terre 69: 17-36.
  • Gradzinski, R., Kielan-Jaworowska, Z., and Maryanska, T. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27: 281-318.
  • Granger, W. and Gregory W.K. 1923. Protoceratops andrewsi, a pre-ceratopsian dinosaur from Mongolia. American Museum Novitates 72: 1-7.
  • Handa, N., Watabe, M., and Tsogtbaatar, K. 2012. New specimens of Protoceratops (Dinosauria: Neoceratopsia) from the Upper Cretaceous in Udyn Sayr, southern Gobi area, Mongolia. Paleontological Research 16: 179-199.
  • Hasegawa, H., Tada, R., Ichinnorov, N., and Minjin, C. 2009. Lithostratigraphy and depositional environments of the Upper Cretaceous Djadokhta Formation, Ulan Nuur basin, southern Mongolia, and its paleoclimatic implication. Journal of Asian Earth Sciences 35: 13-26.
  • Hone, D.W. and Naish, D. 2013. The "species recognition hypothesis" does not explain the presence and evolution of exaggerated structures in non-avialan dinosaurs. Journal of Zoology, 290: 172-180.
  • Hone, D.W., Wood, D., and Knell, R.J. 2016. Positive allometry for exaggerated structures in the ceratopsian dinosaur Protoceratops andrewsi supports socio-sexual signaling. Palaeontologia Electronica 19: 1-13.
  • Horner, J.R., Varricchio, D.J., and Goodwin, M.B. 1992. Marine transgressions and the evolution of Cretaceous dinosaurs. Nature 358: 59-61.
  • Hunt, G. and Rabosky, D.L. 2014. Phenotypic evolution in fossil species: pattern and process. Annual Review of Earth and Planetary Sciences 42: 421-441.
  • Ivakhnenko, M.F. [Ivahnenko, M.F.] and Kurzanov, S.M. 1988. Geological structure and age of localities Udan Sayr and Shara Tsav [in Russian]. Sovmestnaa Sovetsko-Mongol'skaa Paleontologiceskaa Ekspeditcia, Trudy 34: 100-105.
  • Jerzykiewicz, T. 2000. Lithostratigraphy and sedimentary settings of the Cretaceous dinosaur beds of Mongolia. In: M.J. Benton, M.A. Shishkin, and D.M. Unwin (eds.), The Age of Dinosaurs in Russia and Mongolia, 279-296, Cambridge University Press, Cambridge.
  • Jerzykiewicz, T., and Russell, D.A. 1991. Late Mesozoic stratigraphy and vertebrates of the Gobi Basin. Cretaceous Research 12: 345-377.
  • Jerzykiewicz, T., Currie, P.J., Eberth, D.A., Johnston, P.A., Koster, E.H., and Zheng, J.J. 1993. Djadokhta Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi. Canadian Journal of Earth Sciences 30: 2180- 2195.
  • Ji, S.A., Zhang, L., Lu, L., and Hao, J. 2017. The first discovery of the Late Cretaceous protoceratopsid fauna from Alxa, Inner Mongolia, China. Acta Geologica Sinica 91: 1908-1909.
  • Kielan-Jaworowska, Z., Hurum, J.H., and Badamgarav, D. 2003. An extended range of the multituberculate Kryptobaatar and distribution of mammals in the Upper Cretaceous of the Gobi Desert. Acta Palaeontologica Polonica 48: 273-278.
  • Kim, B., Yun, H., and Lee, Y-N. 2019. The postcranial skeleton of Bagaceratops (Ornithischia: Neoceratopsia) from the Baruungoyot Formation (Upper Cretaceous) in Hermiin Tsav of southwestern Gobi, Mongolia [in Korean]. Journal of the Geological Society of Korea 55: 179-190.
  • Knapp, A., Knell, R.J., Farke, A.A., Loewen, M.A., and Hone, D.W. 2018. Patterns of divergence in the morphology of ceratopsian dinosaurs: sympatry is not a driver of ornament evolution. Proceedings of the Royal Society B: Biological Sciences 285:20180312.
  • Kurzanov, S.M. 1992. A gigantic protoceratopsid from the Upper Cretaceous of Mongolia. Paleontological Journal 26: 103-116.
  • Lambert, O., Godefroit, P., Li, H., Shang, C.Y., and Dong, Z.M. 2001. A new species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P.R. China). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la terre 71: 5-28.
  • Lawson, R. and Lieb, C.S. 1990. Variation and hybridization in Elaphe bairdi (Serpentes: Colubridae). Journal of Herpetology 24: 280-292.
  • Lillegraven, J.A. and McKenna, M.C. 1986. Fossil mammals from the "Mesaverde" Formation (Late Cretaceous, Judithian) of the Bighorn and Wind River basins, Wyoming, with definitions of Late Cretaceous North American land-mammal "ages". American Museum Novitates 2840: 1-68.
  • Longrich, N.R., Currie, P.J., and Dong, Z. 2010. A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia. Palaeontology 53: 945-960.
  • Lucas, S.G. and Estep, J.W. 1998. Vertebrate biostratigraphy and biochronology of the Cretaceous of China. New Mexico Museum of Natural History and Science Bulletin 14: 1-20.
  • Makovicky, P.J. 2002. Taxonomic Revision and Phylogenetic Relationships of Basal Neoceratopsia (Dinosauria: Ornithischia). 288 pp. Ph.D. Thesis, Columbia University, New York, NY.
  • Makovicky, P.J. 2008. Telling time from fossils: a phylogeny-based approach to chronological ordering of paleobiotas. Cladistics 24: 350- 371.
  • Makovicky, P.J. and Norell, M.A. 2006. Yamaceratops dorngobiensis, a new primitive ceratopsian (Dinosauria: ornithischia) from the Cretaceous of Mongolia. American Museum Novitates 3530:1-42.
  • Makulbekov, N.M. and Kurzanov, S.M. 1986. The biogeographical relationships of the Late Cretaceous biota of Mongolia [in Russian]. Sovmestnaa Sovetsko-Mongol'skaa Paleontologiceskaa Ekspeditcia, Trudy 29: 106-112.
  • Maryanska, T. and Osmolska, H. 1975. Protoceratopsidae (Dinosauria) of Asia. Palaeontologia Polonica 33: 133-181.
  • McDonald, A.T. 2011. A subadult specimen of Rubeosaurus ovatus (Dinosauria: Ceratopsidae), with observations on other ceratopsids from the Two Medicine Formation. PloS one 6: e22710.
  • Morschhauser, E.M. 2012. The Anatomy and Phylogeny of Auroraceratops (Ornithischia: Ceratopsia) from the Yujingzi Basin of Gansu Province, China. 629 pp. PhD Thesis, University of Pennsylvania, Philadelphia.
  • Nabavizadeh, A. and Weishampel, D.B. 2016. The predentary bone and its significance in the evolution of feeding mechanisms in ornithischian dinosaurs. The Anatomical Record 299: 1358-1388.
  • Norell, M. and Makovicky, P.J. 1999. Important Features of the Dromaeosaurid Skeleton II: Information from Newly Collected Specimens of Velociraptor mongoliensis. American Museum Novitates 3282: 1-45.
  • Osmolska, H. 1980. The Late Cretaceous vertebrate assemblages of the Gobi Desert, Mongolia. Memoirs de la Societe Geologique de France 139: 145-150.
  • Padian, K. and Horner, J.R. 2011. The evolution of "bizarre structures" in dinosaurs: biomechanics, sexual selection, social selection or species recognition? Journal of Zoology 283: 3-17.
  • Pei, R., Norell, M.A., Barta, D.E., Bever, G.S., Pittman, M., and Xu, X. 2017. Osteology of a new Late Cretaceous troodontid specimen from Ukhaa Tolgod, Omnogovi Aimag, Mongolia. American Museum Novitates 2017: 1-48.
  • Rougier, G.W., Sheth, A.S., Spurlin, B.K., Bolortsetseg, M., and Novacek, M.J. 2016. Craniodental anatomy of a new Late Cretaceous multituberculate mammal from Udan Sayr, Mongolia. Palaeontologia Polonica 67: 197-248.
  • Scannella, J.B., Fowler, D.W., Goodwin, M.B., and Horner, J.R. 2014. Evolutionary trends in Triceratops from the Hell Creek formation, Montana. Proceedings of the National Academy of Sciences 111: 10245-10250.
  • Shuvalov, V.F. 2000. The Cretaceous stratigraphy and palaeobiogeography of Mongolia. In: M.J. Benton, M.A. Shishkin, and D.M. Unwin (eds.), The Age of Dinosaurs in Russia and Mongolia, 256-278. Cambridge University Press, Cambridge.
  • Shuvalov, V.F. [Suvalov, V.F.], and Chkhikvadze, V.M. [Chikvadze, V.M.] 1979. On stratigraphical and systematical position of some freshwater turtles from new Cretaceous localities in Mongolia [in Russian]. Sovmestnaa Sovetsko-Mongol'skaa Paleontologiceskaa Ekspeditcia, Trudy 8: 58-76.
  • Smith, J.M. 1966. Sympatric speciation. The American Naturalist 100: 637-650.
  • Storrs, G.W. and Efimov, M.B. 2000. Mesozoic crocodyliforms of northcentral Eurasia. In: M.J. Benton, M.A., Shishkin, D.M. Unwin, and E.N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia, 402-419. Cambridge University Press, Cambridge.
  • Szymura, J.M. 1993.Analysis of hybrid zones with Bombina. In: R.G. Harrison and R.G. Harrison (eds.), Hybrid Zones and the Evolutionary Process, 261-289. Oxford University Press, Oxford.
  • Tereschenko, V.S. 1996. A reconstruction of the locomotion of Protoceratops. Paleontological Journal 30: 232-245.
  • Tereschenko, V.S. 2007. Key to Protoceratopoid vertebrae (Ceratopsia, Dinosauria) from Mongolia. Paleontological Journal 41: 175-188.
  • Tereschenko, V.S. 2008. Adaptive features of protoceratopoids (Ornithischia: Neoceratopsia). Paleontological Journal 42: 273-286.
  • Tereschenko, V.S. 2018. On Polymorphism of Protoceratops andrewsi Granger et Gregory, 1923 (Protoceratopidae, Neoceratopsia). Paleontological Journal 52: 429-444.
  • Tong, H., Li, L., Jie, C., and Yi, L. 2016. New material of Jiangxichelys ganzhouensis Tong & Mo, 2010 (Testudines: Cryptodira: Nanhsiungchelyidae) and its phylogenetic and palaeogeographical implications. Geological Magazine 154: 456-464.
  • Tsogtbaatar, K. 2004. Fossil specimens prepared in Mongolian Paleontological center 1993-2001. Hayashibara Museum of Sciences Research Bulletin 2: 123-128.
  • Tsogtbaatar, K. and Chinzorig, T. 2010. Fossil specimens prepared in Mongolian Paleontological Center: 2002-2008 Hayashibara Museum of Sciences Research Bulletin 3: 155-166.
  • Tsogtbaatar, K., Weishampel, D.B., Evans, D.C., and Watabe, M. 2014. A new hadrosauroid (Plesiohadros djadokhtaensis) from the Late Cretaceous Djadokhtan Fauna of southern Mongolia. In: D.A. Eberth and D.C. Evans (eds.), Hadrosaurs, 108-135. Indiana University Press, Bloomington.
  • Tsuihiji, T., Barsbold, R., Watabe, M., Tsogtbaatar, K., Chinzorig, T., Fujiyama, Y., and Suzuki, S. 2014. An exquisitely preserved troodontid theropod with new information on the palatal structure from the Upper Cretaceous of Mongolia. Naturwissenschaften 101: 131-142.
  • Turner, A.H., Makovicky, P.J., and Norell, M.A. 2012. A review of dromaeosaurid systematics and paravian phylogeny. Bulletin of the American Museum of Natural History 371: 1-206.
  • Tverdochlebov, V.P. [Tverdohlebov, V.P.] and Zybin, J.I. 1974. Genesis of the Upper Cretaceous sediments with dinosaur remains at Tugrikin-us and Alag-Taag localities [in Russian]. Sovmestnaa Sovetsko-Mongol'skaa Paleontologiceskaa Ekspeditcia, Trudy 1: 314-319.
  • Varriale, F.J. 2011. Dental Microwear and the Evolution of Mastication in Ceratopsian Dinosaurs. 470 pp. Ph.D. Thesis, The Johns Hopkins University, Baltimore.
  • Watabe, M. and Suzuki, S. 2000. Report on the Japan-Mongolia Joint Paleontological expedition to the Gobi Desert, 1996. Hayashibara Museum of Natural Sciences Research Bulletin 1: 30-44.
  • Watabe, M. and Tsogtbaatar, K. 2004. Report on the Japan-Mongolia Joint Paleontological expedition to the Gobi Desert, 2000. Hayashibara Museum of Natural Sciences Research Bulletin 2: 45-67.
  • Watabe, M., Tsogtbaatar, K., Suzuki, S., and Saneyoshi, M. 2010. Geology of dinosaur-fossil-bearing localities (Jurassic and Cretaceous: Mesozoic) in the Gobi Desert: Results of the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin 3: 41-118.
  • Wilson, J.P., Ryan, M.J., and Evans, D.C. 2020.A new, transitional centrosaurine ceratopsid from the Upper Cretaceous Two Medicine Formation of Montana and the evolution of the "Styracosaurus -line" dinosaurs. Royal Society Open Science 7: 200284.