Ptychadena keilingi (Monard, 1937)

Keiling’s Ridged Frog (Fig. 31; Map 29)

Material (38 specimens): PEM A12428, Cuiva River bridge on EN250, -11.98345° 17.72367°, 1,267m asl; PEM A12477–8, INBAC (no number x2), Cuanavale River source lake, -13.08537° 18.89098°, 1,360 m asl; PEM A12491, Dala River, near Samanunga village, -12.93170° 18.81458°, 1,363 m asl; PEM A12559–61, Cuito River source lake, -12.68935° 18.36012°, 1,431 m asl; PEM A12646, outlet of Cuito River source lake, -12.70453° 18.35445°, 1,429 m asl; PEM A12752–9, INBAC: WC-4797, (no number x2), Cuando River source, -13.00346° 19.12751°, 1,353 m asl; PEM A12782, Quembo River, trap 1, -13.13592° 19.04417°, 1,369 m asl; PEM A12822– 4, Cuiva River source, -12.66856° 18.35307°, 1,433 m asl; PEM A12894–6, INBAC: WC-4691, INBAC: (no number x1), Quembo River source lake, -13.13624° 19.04591°, 1,366 m asl; PEM A13818–19, en route to Cuemba Village, -11.96587° 17.76176°, 1,302 m asl; PEM A14694, Lungwebungu River camp, right bank past first oxbow, -12.57956° 18.67761°, 1,295 m asl; PEM A14736, Quembo River, walk back from small waterfall, -13.52988° 19.28340°, 1,242 m asl; PEM A14780, Quembo River bridge camp, trap 1, -13.52801° 19.28147°, 1,236 m asl; PEM A14781, INBAC: WC-6962, Quembo River right side tributary (Micongo River) past village, -13.51877° 19.28487°, 1,248 m asl; PEM A14894, wetland near old quarry east of Quemba, -12.16960° 18.22965°, 1,353 m asl. Additional specimens (2 specimens): SAIAB 204561 (2 specimens), Quembo River source lake, -13.13611° 19.04500°, 1,367 m asl. Description: Medium sized Ptychadena; elongated body; longitudinal dorsal ridges present; rostrum protruding well beyond lower jaw and forming a very pointed and elongated keratinized protrusion; long toes; reduced webbing, with up to four phalanges of longest toe free of webbing. Dorsum colorful; red infusions on the flanks; black facial mask, from snout through eye to forearm; broad light vertebral band present; thin light line on dorsal tibia; top of thighs with light and dark bands; ventrum immaculate, with black spots on chest in some specimens; black spot in front of arm that connects with lower jaw; lower jaw with spots; back of thighs with irregular longitudinal black and yellow stripes. Adult females (n = 19) varied from 24.6–39.9 (34.8) mm (largest female: PEM A12755); adult males (n = 14) varied from 25.3–31.5 (27.9) mm (largest male: PEM A14894). Habitat and natural history notes: Found in the flooded margins of rivers and lakes. Comments: Originally described from Dala, in north-eastern Angola (Laurent 1964). It is only known from northeastern Angola (Laurent 1964; Marques et al. 2018), western Zambia (Channing 2001), and southwestern DRC (Channing and Rödel 2019). This is the southernmost record for this species, and a new species record for southeastern Angola and the Okavango River basin.