Figs 40C 1 –C 3, 4Y, 5, 7

* Pleurotoma concinna n.f.— Handmann 1883: 171 [non Pleurotoma concinna Scacchi, 1836].

Perronea [sic] (Perronea) [sic] concinna Handmann — Mariani & Papp 1966: 144, pl. 1, figs 11–12 [non Pleurotoma concinna Scacchi, 1836].

Type material. Neotype: NHMW 2021 /0158/0001, SL: 23.2 mm, MD: 9.0 mm, St. Veit an der Triesting (Austria), figs 40C 1 –C 3. The syntypes from St. Veit an der Triesting (Austria) have been stored in the Kollegium Kalksburg in Vienna, a Catholic private school. One syntype has been illustrated in Mariani & Papp (1966, pl. 1, fig. 11) but this specimen is lost. Therefore, we designate a neotype from the type locality.

Type locality. St. Veit an der Triesting (Austria), Vienna Basin.

Type stratum. Sandy nearshore equivalent of the Baden Formation.

Age. Middle Badenian (late Langhian).

Etymology. In honor of Gerhard Wanzenböck (Gainfarn, Austria), fossil collector and friend of the first author.

Revised description. Shell medium-sized, moderately broad fusiform with relatively short, conical spire. Protoconch largely destroyed, terminating with broad, strongly convex whorl. Teleoconch of seven whorls. First three teleoconch whorls faintly concave with narrow subsutural spiral cord and close-set, strongly opisthocline, comma-shaped axial riblets; no suprasutural cord. Sculpture changing on fourth teleoconch whorl; subsutural cord becomes moderately broad, rounded collar, whorl profile weakly concave below, bearing six narrow, flattened spiral cords. Suture narrowly incised, superficial, linear. Last whorl ~70% of total height, ovate. Subsutural collar broad, rounded, weakly swollen, sharply delimited. Subsutural ramp, shoulder and base not delimited. Profile below collar evenly rounded, strongly constricted at base. Siphonal fasciole poorly developed. Fine spiral cords over ramp, obsolete at mid-whorl, reappearing on abapical half of base, strengthening slightly towards and over fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately narrow and deep, asymmetrically U-shaped with apex mid-ramp. Siphonal canal long, straight, narrow. Columella weakly excavated, very slightly twisted at fasciole. Columellar and parietal callus thickened, well demarcated from base, forming broad rim.

Discussion. Handmann (1883) was unaware that his Pleurotoma concinna was variously preoccupied by Scacchi, (1836: 12), Adams (1852:140) and Dunker (1857: 356) (see Tucker 2004: 227). Therefore, we propose Clavatula wanzenboecki as replacement name. Perrona wanzenboecki nov. nom. is unique amongst its congeners in not having a suprasutural band developed and not having any of the major cords on the last whorl developed, so that the last whorl profile is evenly rounded. It is reminiscent of Tomellana dulaii nov. nom. and Megaclavatula nemethi (Kovács & Vicián, 2021) in general shape, but is much smaller. In addition, T. dulaii has a slightly angulated subsutural collar and M. nemethi is slightly more slender and has tripartite early teleoconch whorl sculpture. Tomellana jouannetii (Des Moulins, 1842) is also very similar, but is much larger, has a higher spire and lacks prominent spiral cords below the subsutural collar (see Peyrot 1931, pl. 8, figs 43, 61, 62, 66, 71). Moreover, both of Tomellana species compared above lack the comma-shaped axial riblets on the neanic whorls, typical of Perrona.

Paleoenvironment. Deposits and faunas from St. Veit an der Triesting represent an intertidal lagoon with mudflats and scattered oyster beds (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: St. Veit an der Triesting (Austria) (Mariani & Papp 1966).