Fig. 2. Phenology of dung beetle species that occurred only in spring and early summer. Symbols indicate the presence of a species in each sample. Sampling started 26 March 2012 (before the first A on the date of collection axis) and ended 16 May 2013 (after the last M on the same axis). Circles show the presence of a beetle species on the sheep farm on the respective collection date, while squares show the presence of a beetle species on the cattle farm on the respective collection date. Presence of a species at the sheep farm is indicated below the labeled presence of the species at the cattle farm.