Published September 28, 2022
| Version v1
Journal article
Restricted
Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification
Authors/Creators
- 1. Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. & fataahzarei@gmail.com; https://orcid.org/0000-0001-5552-4301
- 2. Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. & hresmaeili@shirazu.ac.ir; https://orcid.org/0000-0002-9736-397X
- 3. Inland Waters Aquaculture Research Center, Iranian Fisheries Sciences Research Institute, Agricultural Research, Education and Extension Organization, Bandar Anzali, Iran.
- 4. Natural History Museum Rijeka, Lorenzov prolaz 1, HR-51000 Rijeka, Croatia.
- 5. SNSB-Bavarian State Collection of Zoology, Department of Ichthyology, Münchhausenstr. 21, 81247 München, Germany. schliewen@snsb.de; https://orcid.org/0000-0003-0187-4319
- 6. National Museum of Natural History, Department of Vertebrate Zoology, Smithsonian Institution, Washington D.C, USA. StepienCA@si.edu; https://orcid.org/0000-0002-5544-4333
Description
Zarei, Fatah, Esmaeili, Hamid Reza, Abbasi, Keyvan, Kovačić, Marcelo, Schliewen, Ulrich K., Stepien, Carol A. (2022): Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification. Zootaxa 5190 (2): 151-193, DOI: 10.11646/zootaxa.5190.2.1
Files
Restricted
The record is publicly accessible, but files are restricted. <a href="https://zenodo.org/account/settings/login?next=https://zenodo.org/records/7120060">Log in</a> to check if you have access.
Linked records
Additional details
References
- Abbasi, K. (2017) Fishes of Guilan. Iliya Culture Publication, Rasht, 208 pp. [in Persian]
- Abdoli, A., Coad, B.W. & Naderi, M. (2000) First record of Rhinogobius similis, Gill 1895 in Iran. Iranian Scientific Fisheries Journal, 9, 73-76.
- Ahnelt, H., Abdoli, A., Naderi, M. & Coad, B.W. (2000) Anatirostrum profundorum: a rare deep-water gobiid species from the Caspian Sea. Cybium, 24, 139-159.
- Ahnelt, H., Bianco, P.G. & Schwammer, H. (1995) Systematics and zoogeography of Knipowitschia caucasica (Teleostei: Gobiidae) based on new records from the Aegean Anatolian area. Ichthyological Exploration of Freshwaters, 6, 49-60.
- Ahnelt, H., Coad, B.W., Abdoli, A. & Zirkohy, H.P. (2007) Gobiid fishes of the genera Chasar, Mesogobius and Neogobius (Teleostei: Gobiidae) from Iran (South Caspian Basin). Zoology in the Middle East, 41, 55-62. https://doi.org/10.1080/09397140.2007.10638227
- Ahnelt, H. & Holcik, J. (1996) Distribution of two species of the genus Neogobius (Pisces: Gobiidae) in the catchment area of the southern Caspian Sea. Acta Universitatis Carolinae Biologica, 40, 99-114.
- Aliev, D.S., Sukhanova, A.I. & Shakirova, F.M. (1988) Fishes of the Inland Waters of Turkmenistan. Ylym, Ashkhabad, 142 pp. [in Russian]
- Beling, D. & Iljin, B. (1927) Benthophiloides brauneri ng, n. sp., ein fur das Schwarzmeerbassin neuer Vertreter der Familie der Gobiidae. Travaux de la Station biologique du Dniepre, Kiev, 2, 309-325. [in German]
- Berg, L.S. (1927) Notes on the Caspian Benthophilus (Gobiidae). In: Festschrift fur Prof. N. M. Knipowitsch 1885-1925. s.n., Moskwa, pp. 331-344.
- Berg, L.S. (1949) Freshwater Fishes of the USSR and Adjacent Countries. Vol. III. Program for Scientific Translations, Jerusalem, 250 pp.
- Bogutskaya, N. (2020a) Benthophilus baeri. The IUCN Red List of Threatened Species, 2020, e.T156755278A156755320. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156755278A156755320.en
- Bogutskaya, N. (2020b) Benthophilus casachicus. The IUCN Red List of Threatened Species, 2020, e.T156755653A156755659. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156755653A156755659.en
- Bogutskaya, N. (2020c) Benthophilus ctenolepidus. The IUCN Red List of Threatened Species, 2020, e.T156755702A156755728. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156755702A156755728.en
- Bogutskaya, N. (2020d) Benthophilus grimmi. The IUCN Red List of Threatened Species, 2020, e.T156756575A156756602. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156756575A156756602.en
- Bogutskaya, N. (2020e) Benthophilus kessleri. The IUCN Red List of Threatened Species, 2020, e.T156756692A156756750. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156756692A156756750.en
- Bogutskaya, N. (2020f) Benthophilus leptocephalus. The IUCN Red List of Threatened Species, 2020, e.T156756871A156756908. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156756871A156756908.en
- Bogutskaya, N. (2020g) Benthophilus leptorhynchus. The IUCN Red List of Threatened Species, 2020, e.T156757038A156757112. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156757038A156757112.en
- Bogutskaya, N. (2020h) Benthophilus pinchuki. The IUCN Red List of Threatened Species, 2020, e.T156755866A156755972. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156755866A156755972.en
- Bogutskaya, N. (2020i) Benthophilus spinosus. The IUCN Red List of Threatened Species, 2020, e.T156758702A156758728. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156758702A156758728.en
- Bogutskaya, N. (2020j) Benthophilus svetovidovi. The IUCN Red List of Threatened Species, 2020, e.T156758973A156759008. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156758973A156759008.en
- Bogutskaya, N. (2020k) Neogobius caspius. The IUCN Red List of Threatened Species, 2020, e.T156761995A156762095. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156761995A156762095.en
- Bogutskaya, N. (2020l) Ponticola bathybius. The IUCN Red List of Threatened Species, 2020, e.T156761759A156761841. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156761759A156761841.en
- Bogutskaya, N. (2021) Benthophilus ragimovi. The IUCN Red List of Threatened Species, 2021, e.T159639612A159639639. https://doi.org/10.2305/IUCN.UK.2021-1.RLTS.T159639612A159639639.en
- Bogutskaya, N.G., Kijashko, P.V., Naseka, A.M. & Orlova, M.I. (2013) Identification Keys for Fish and Invertebrates of the Caspian Sea. Vol. 1. Fish and Molluscs. KМK Scientific Press Ltd., St. Petersburg and Moscow, 543 pp. [in Russian]
- Boldyrev, V.S. & Bogutskaya, N.G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18, 31-96.
- Brauner, A. (1898) Zur Ichtyofauna des Dnjeprs. Wiestnik Rybopromyschlennosti, 450 pp. [in German]
- Coad, B.W. (2008) Fresh water fishes of Iran. Available from: http://www.briancoad.com/main.asp?page=welcome.asp (accessed 15 May 2022)
- Coad, B.W. & Abdoli, A. (2000) Rhinogobius cf. similis Gill, 1859, a goby new to the fish fauna of Iran and the problem of alien invasions. Zoology in the Middle East, 20, 55-59. https://doi.org/10.1080/09397140.2000.10637812
- De Filippi, F. (1863) Nuove o poco note specie di animali vertebrati raccolti in un viaggio in Persia, nell'estate dell'anno 1862. Archivio per la Zoologia, l'Anatomia e la Fisiologia, 2, 377-394. [in Italian]
- De Filippi, F. (1865) Note de un Viaggio in Persia Nei 1862. G. Daelli & C. Editori, Milano, 396 pp. [in Italian]
- Eagderi, S., Nikmehr, N. & Poorbagher, H. (2020) Ponticola patimari sp. nov.(Gobiiformes: Gobiidae) from the southern Caspian Sea basin, Iran. FishTaxa, 17, 22-31.
- Economidis, P.S. & Miller, P.J. (1990) Systematics of freshwater gobies from Greece (Teleostei: Gobiidae). Journal of Zoology, 221, 125-170. https://doi.org/10.1111/j.1469-7998.1990.tb03781.x
- Esmaeili, H.R. & Abbasi, K. (2021) Checklist of fishes of the Caspian Sea basin: land of wetlands. In: Jawad, L.A. (Ed.), Southern Iraq's Marshes. Springer, Cham, pp. 319-349.
- Esmaeili, H.R., Coad, B.W., Gholamifard, A., Nazari, N. & Teimory, A. (2010) Annotated сhecklist of the freshwater fishes of Iran. Zoosystematica Rossica, 19, 361-386.
- Esmaeili, H.R., Sayyadzadeh, G., Eagderi, S. & Abbasi, K. (2018) Checklist of freshwater fishes of Iran. FishTaxa, 3, 1-95.
- Freyhof, J. (2014) Ponticola cyrius. The IUCN Red List of Threatened Species, 2014, e.T19513722A19849649. https://doi.org/10.2305/IUCN.UK.2014-1.RLTS.T19513722A19849649.en
- Freyhof, J. (2020) Knipowitschia bergi. The IUCN Red List of Threatened Species, 2020, e.T135608A137270791. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T135608A137270791.en
- Freyhof, J. & Kottelat, M. (2008a) Babka gymnotrachelus. The IUCN Red List of Threatened Species, 2008, e.T188118A8643960. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T188118A8643960.en
- Freyhof, J. & Kottelat, M. (2008b) Benthophiloides brauneri. The IUCN Red List of Threatened Species, 2008, e.T2761A9477974. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T2761A9477974.en
- Freyhof, J. & Kottelat, M. (2008c) Benthophilus granulosus. The IUCN Red List of Threatened Species, 2008, e.T135682A4180004. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135682A4180004.en
- Freyhof, J. & Kottelat, M. (2008d) Benthophilus leobergius. The IUCN Red List of Threatened Species, 2008, e.T135680A4178654. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135680A4178654.en
- Freyhof, J. & Kottelat, M. (2008e) Benthophilus macrocephalus. The IUCN Red List of Threatened Species, 2008, e.T135656A4172098. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135656A4172098.en
- Freyhof, J. & Kottelat, M. (2008f) Benthophilus mahmudbejovi. The IUCN Red List of Threatened Species, 2008, e.T135654A4171635. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135654A4171635.en
- Freyhof, J. & Kottelat, M. (2008g) Knipowitschia caucasica. The IUCN Red List of Threatened Species, 2008, e.T11030A3240732. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T11030A3240732.en
- Freyhof, J. & Kottelat, M. (2008h) Knipowitschia longecaudata. The IUCN Red List of Threatened Species, 2008, e.T135509A4135081. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135509A4135081.en
- Freyhof, J. & Kottelat, M. (2008i) Neogobius melanostomus. The IUCN Red List of Threatened Species, 2008, e.T14524A4442374. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T14524A4442374.en
- Freyhof, J. & Kottelat, M. (2008j) Neogobius pallasi. The IUCN Red List of Threatened Species, 2008, e.T135596A4156361. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T135596A4156361.en
- Freyhof, J. & Kottelat, M. (2008k) Ponticola gorlap. The IUCN Red List of Threatened Species, 2008, e.T188114A8642335. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T188114A8642335.en
- Freyhof, J. & Kottelat, M. (2008l) Ponticola syrman. The IUCN Red List of Threatened Species, 2008, e.T188116A8643348. https://doi.org/10.2305/IUCN.UK.2008.RLTS.T188116A8643348.en
- Freyhof, J. & Naseka, A.M. (2007) Proterorhinus tataricus, a new tubenose goby from Crimea, Ukraine (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18, 325-334.
- Fricke, R., Eschmeyer, W.N. & Van der Laan, R. (2022) Eschmeyer's catalog of fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. Available from: http://researcharchive.calacademy.org/research/ ichthyology/catalog/fishcatmain.asp (accessed 15 May 2022)
- Iljin, B.S. (1928) Two new genera and a new species of Gobiidae from the Caspian sea. Reports of the Astrakhan Scientific Fishery Station, 3, 1-13.
- Iljin, B.S. (1936) A new goby from the Caspian Sea Gobius nonultimus sp. n (Pisces, Gobiidae). Doklady Akademii Nauk SSSR, Series A, 337-339. [in Russian]
- Iljin, B.S. (1941) Asra turcomanus, a new genus and species of gobies (Gobiidae) from the Caspian Sea. Biology Bulletin of the Academy of Sciences of the USSR, 3, 385-390.
- IUCN. (2021) The IUCN red list of threatened species. version 2021-3. International Union for Conservation of Nature, Gland, Switzerland. Available from: https://www.iucnredlist.org (accessed 14 June 2022)
- Jouladeh-Roudbar, A., Ghanavi, H.R. & Doadrio, I. (2020) Ichthyofauna from Iranian freshwater: annotated checklist, diagnosis, taxonomy, distribution and conservation assessment. Zoological Studies, 59, 21. https://doi.org/10.6620/ZS.2020.59-21
- Karsiotis, S.I., Brown, J.E., Pierce, L.R. & Stepien, C.A. (2012) Salinity tolerance of the invasive round goby: experimental implications for seawater ballast exchange and spread to North American estuaries. Journal of Great Lakes Research, 38, 121-128. https://doi.org/10.1016/j.jglr.2011.12.010
- Kessler, K.F. (1874) A description of fishes belonging to the families common to both the Black and the Caspian seas. Travaux de la Societe des Naturalistes de St. Petersbourg, 5, 191-324. [in French]
- Kessler, K.T. (1877) The Aralo-Caspian Expedition. IV. Fishes of the Aralo-Caspio-Pontine Ichthyological Region. Naturgeschichte der Fische Islands, St. Petersburg, 360 pp.
- Knipovich, N.M. (1906) General Review of the Caspian Expedition of 1904. [Reports of the Caspian Expedition in the Year 1904]. Glavnoye Upravleniye Zemleustroystva I Zemledeliya, St. Peterburg, 113 pp. [in Russian]
- Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland & Freyhof, Berlin, Germany.
- Kovacic, M. (2020) Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification. Zootaxa, 4877, 75-101. https://doi.org/10.11646/zootaxa.4877.1.3
- Kovacic, M., Esmaeili, H.R., Zarei, F., Abbasi, K. & Schliewen, U.K. (2021) A new species of Tadpole goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea. Zootaxa, 4980 (1), 45-63. https://doi.org/10.11646/zootaxa.4980.1.3
- Kuliev, Z.M. (1989) Ryby Zaliva Kirova Kaspiiskogo Morya (Sistematika, Biologiya, Promysel). Akademiya Nauk Azerbaidzhanskoi SSR, Instituta Zoologii, Baku, 183 pp. [in Russian]
- Larson, H. (2019a) Knipowitschia iljini. The IUCN Red List of Threatened Species, 2019, e.T123435142A123494817. https://doi.org/10.2305/IUCN.UK.2019-3.RLTS.T123435142A123494817.en
- Larson, H. (2019b) Mesogobius nonultimus. The IUCN Red List of Threatened Species, 2019, e.T123435177A123494822. https://doi.org/10.2305/IUCN.UK.2019-3.RLTS.T123435177A123494822.en
- Mamilov, N. (2020a) Anatirostrum profundorum. The IUCN Red List of Threatened Species, 2020, e.T156754995A156755086. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156754995A156755086.en
- Mamilov, N. (2020b) Benthophiloides turcomanus. The IUCN Red List of Threatened Species, 2020, e.T156755140A156755201. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156755140A156755201.en
- Mamilov, N. (2020c) Ponticola goebelii. The IUCN Red List of Threatened Species, 2020, e.T156763140A156763185. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T156763140A156763185.en
- Miller, P.J. (1986) Gobiidae. In: Whitehead, P.J.P., Bauchot, M.-L., Hureau, J.-C., Nielsen, J. & Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and the Mediterranean. Vol. 3. UNESCO, Paris, pp. 1019-1085.
- Miller, P.J. (2003) The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, Germany.
- Miller, P.J. (2004a) The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH Wiebelsheim, Verlag fur Wissenschaft und Forschung. [unknown pagination]
- Miller, P.J. (2004b) Knipowitschia longecaudata (Kessler, 1877). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 383-397.
- Miller, P.J. (2004c) Knipowitschia Iljin, 1927. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 331-337.
- Miller, P.J. & Pinchuk, V.I. (2004a) Hyrcanogobius bergi Iljin, 1928. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 432-438.
- Miller, P.J. & Pinchuk, V.I. (2004b) Knipowitschia iljini Berg, 1931. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 375-382.
- Miller, P.J., Vasil'eva, E.V. & Economou,A.N. (2004) Knipowitschia caucasica (Berg, 1916). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2 AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 3343-3364.
- Mousavi-Sabet, H. (2021) Dams and their impacts on fishes in Iran. In: Jawad, L.A. (Ed.), Tigris and Euphrates Rivers: Their Environment from Headwaters to Mouth. Aquatic Ecology Series, Springer, Cham, pp. 401-420.
- Naseka, N.M. & Bogutskaya, N.G. (2009) Fishes of the Caspian Sea: zoogeography and updated check-list. Zoosystematica Rossica, 18, 295-317.
- Navozov, N.P. (1912) Materials for an ichthyofauna of the Ural River basin. Vestnik Ryby, 1912, 252-283. [in Russian]
- Neilson, M.E. & Stepien, C.A. (2009) Escape from the Ponto-Caspian: evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei). Molecular Phylogenetics and Evolution, 52, 84-102. https://doi.org/10.1016/j.ympev.2008.12.023
- Neilson, M.E. & Stepien, C.A. (2011) Historic speciation and recent colonization of Eurasian monkey gobies (Neogobius fluviatilis and N. pallasi) revealed by DNA sequences, microsatellites, and morphology. Diversity and Distributions, 17, 688-702. https://doi.org/10.1111/j.1472-4642.2011.00762.x
- Ohayon, J.L. & Stepien, C.A. (2007) Genetic and biogeographic relationships of the racer goby Neogobius gymnotrachelus (Gobiidae: Teleostei) from exotic and native Eurasian locations. Journal of Fish Biology, 71, 360-370. https://doi.org/10.1111/j.1095-8649.2007.01659.x
- Panin, G.N., Mamedov, R.M. & Mitrofanov, I.V. (2005) Present State of the Caspian Sea. Nauka, Moscow, 356 pp. [in Russian]
- Pinchuk, V.I. (1977) Classification of gobies of the genera Gobius Linne (home species), Neogobius Iljin and Mesogobius Bleeker. Voprosy Ikhtiologii, 17, 587-596. [in Russian]
- Pinchuk, V.I. & Miller, P.J. (2003a) Benthophilus pinchuki Ragimov, 1982. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 246-250.
- Pinchuk, V.I. & Miller, P.J. (2003b) Neogobius caspius (Eichwald, 1831). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 173-180.
- Pinchuk, V.I. & Miller, P.J. (2004a) Anatirostrum profundorum (Berg, 1927). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 274-278.
- Pinchuk, V.I. & Miller, P.J. (2004b) Benthophiloides brauneri Beling & Iljin, 1927. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 153-159.
- Pinchuk, V.I. & Miller, P.J. (2004c) Benthophilus ctenolepidus Kessler, 1877. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 187-191.
- Pinchuk, V.I. & Miller, P.J. (2004d) Mesogobius nonultimus (Iljin, 1936). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 132-137.
- Pinchuk, V.I., Vasil'eva, E.D. & Miller, P.J. (2003a) Neogobius ratan (Nordmann, 1840). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 357-369.
- Pinchuk, V.I., Vasil'eva, E.D. & Miller, P.J. (2003b) Neogobius syrman (Nordmann, 1840). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 377-397.
- Pinchuk, V.I., Vasil'eva, E.D. & Miller, P.J. (2004a) Benthophilus baeri Kessler, 1877. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 175-181.
- Pinchuk, V.I., Vasil'eva, E.D. & Miller, P.J. (2004b) Chasar bathybius (Kessler, 1877). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe Vol. 8/II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 97-105.
- Pinchuk, V.I., Vasil'eva, E.D., Vasil'ev, V.P. & Miller, P.J. (2003c) Neogobius fluviatilis (Pallas, 1814). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 222-252.
- Pinchuk, V.I., Vasil'eva, E.D., Vasil'ev, V.P. & Miller, P.J. (2003d) Neogobius melanostomus (Pallas, 1814). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 293-345.
- Pinchuk, V.I., Vasileva, E.D., Vasilev, V.P. & Miller, P.J. (2003e) Neogobius gymnotrachelus (Kessler, 1857). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 265-279.
- Ragimov, D.B. (1965) On the distribution of gobiids at the western coast of the central and southern Caspian. Doklady Akademii Nauk Azerbaijanskoy SSSR, 21, 47-50. [in Russian]
- Ragimov, D.B. (1967) On the systematics of fishes belonging to the genus Gobius in the Caspian Sea. In: Biological Productivity of the Kurinsk-Caspian Fishing Region. Akademii Nauk Azerbaijanskoy SSSR. Zook, Baku, pp. 252-277. [in Russian]
- Ragimov, D.B. (1985) Some Caspian species of the genus Benthophilus. Voprosy Ichtiologii, 25, 917-924. [in Russian]
- Sadeghi, R., Esmaeili, H.R., Zarei, F., Esmaeili, A. & Abbasi, K. (2019) The taxonomic status of an introduced freshwater goby of the genus Rhinogobius to Iran (Teleostei: Gobiidae). Zoology in the Middle East, 65, 51-58. https://doi.org/10.1080/09397140.2018.1540149
- Sakai, H., Ikoma, K., Frolov, S.V., Yamazaki, Y., Takahashi, H. & Ida, H. (2000) Morphological features of a Russian freshwater goby, Rhinogobius lindbergi (Pisces: Gobiidae), and its genetic relationships to Japanese species. Biogeography, 2, 51- 61.
- Sal'nikov, V.B. (1995) Possible changes in the composition of the ichthyofauna after completion of the Karakum Canal in Turkmenistan. Voprosy Ichtiologii, 35, 108-121. [in Russian]
- Sanzo, L. (1911) Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi. Mittheilungen aus der Zoologischen Station zu Neapel, 20, 251-328. [in Italian]
- Shakirova, F.M. & Sukhanova, A.I. (1994) Iktiofauna Turkmenistana (sostav i rasprostranenie). Izvestiya Akademii Nauk Turkmenistana, Seriya Biologicheskikh Nauk, 3, 35-45. [in Russian]
- Stepien, C.A. & Tumeo, M.A. (2006) Invasion genetics of Ponto-Caspian gobies in the Great Lakes: a 'cryptic' species, absence of founder effects, and comparative risk analysis. Biological Invasions, 8, 61-78. https://doi.org/10.1007/s10530-005-0237-x
- Suvorov, E.N. (1948) Fishery Water Bodies of the USSR. Izdatel'stvo Leningradskogo Gosudarsvennogo Universiteta, Leningrad, 238 pp. [in Russian]
- Suzuki, T., Shibukawa, K., Senou, H. & Chen, I.-S. (2016) Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, 63, 227-238. https://doi.org/10.1007/s10228-015-0494-3
- Tajbakhsh, F., Rajabi-Maham, H., Abdoli, A., Stepien, C.A. & Kiabi, B.H. (2022) DNA sequence support for re-classification of the endemic south Caspian Sea deepwater goby as Neogobius bathybius (from Ponticola; Perciformes: Gobiidae) and recent population expansion of a continuous population. Ichthyology and Herpetology, 110, 13-21. https://doi.org/10.1643/i2020015
- Tajbakhsk, F., Stepien, C.A., Abdoli, A., Tabatabaei, N. & Kiabi, B.H. (2018) Geometric morphometric and meristic analysis of the deepwater goby, Ponticola bathybius (Kessler, 1877) (Teleostei: Gobiidae) in the Iranian waters of the Caspian Sea. Iranian Journal of Ichthyology, 5, 64-73.
- Vasil'eva, E.D., Mousavi-Sabet, H. & Vasil'ev, V.P. (2015) Ponticola iranicus sp. nov. (Actinopterygii: Perciformes: Gobiidae) from the Caspian Sea basin. Acta Ichthyologica et Piscatoria, 45, 189-197. https://doi.org/10.3750/AIP2015.45.2.09
- Vasil'eva, E.D. & Kuga, T.I. (2008) Gobies of the genus Rhinogobius (Gobiidae) of Primorye and water bodies of Central Asia and Kazakhstan: II. Comparative craniological analysis of gobies introduced to Central Asia. Journal of Ichthyology, 48, 29-36. https://doi.org/10.1134/S0032945208010037
- Vasil'eva, E.D., Schwarzhans, W.W., Medvedev, D.A. & Vasil'ev, V.P. (2016) Cryptic species of Ponto-Caspian bighead goby of the genus Ponticola (Gobiidae). Journal of Ichthyology, 56, 1-18. https://doi.org/10.1134/S003294521601015X
- Vasil'eva, E.D. & Vasil'ev, V.P. (2003a) Neogobius cyrius (Kessler, 1874). In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 203-211.
- Vasil'eva, E.D. & Vasil'ev, V.P. (2003b) Neogobius gorlap Iljin in Berg, 1949. In: Miller, P.J. (Ed.), The Freshwater Fishes of Europe. Vol. 8/I Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 253-264.
- Zarei, F., Esmaeili, H.R., Kovacic, M., Schliewen, U.K. & Abbasi, K. (2022a) Ponticola hircaniaensis sp. nov., a new and critically endangered gobiid species (Teleostei: Gobiidae) from the southern Caspian Sea basin. Zootaxa, 5154 (4), 401- 430. https://doi.org/10.11646/zootaxa.5154.4.1
- Zarei, F., Esmaeili, H.R., Sadeghi, R., Schliewen, U.K., Kovacic, M., Abbasi, K. & Gholamhosseini, A. (2022b) An integrative insight into the diversity, distribution and biogeography of the freshwater endemic clade of the Ponticola syrman group (Teleostei: Gobiidae) in the Caucasus biodiversity hotspot. Ecology & Evolution, 12. https://doi.org/10.1002/ece3.9300
- Zarei, F., Esmaeili, H.R., Schliewen, U.K. & Abbasi, K. (2022c) Taxonomic diversity and distribution of the genus Proterorhinus (Teleostei: Gobiidae) in the Caucasus biodiversity hotspot with conservation implications. Aquatic Conservation: Marine and Freshwater Ecosystems, 32, 129-138. https://doi.org/10.1002/aqc.3728
- Zarei, F., Esmaeili, H.R., Schliewen, U.K., Abbasi, K. & Sayyadzadeh, G. (2021) Mitochondrial phylogeny, diversity, and ichthyogeography of gobies (Teleostei: Gobiidae) from the oldest and deepest Caspian sub-basin and tracing source and spread pattern of an introduced Rhinogobius species at the tricontinental crossroad. Hydrobiologia, 848, 1267-1293. https://doi.org/10.1007/s10750-021-04521-0