Distribution. SE Bolivia (departments of Tarija and Santa Cruz), N Argentina (provinces of Jujuy, Salta, Formosa, and Chaco), Brazil (W of the Rio Parana— through the N of Mato Grosso State into SW Goias, and Mato Grosso do Sul states), and Paraguay (E of the Rio Paraguay as far as the mouth of the Rio Parana). Its range to the west in Bolivia is poorly known; it would seem that it is absent from the Bolivian Chaco as it is from the adjacent Paraguayan Chaco west of the Rio Paraguay. There is no evidence to date that the Yungas populations in SE Bolivia and NW Argentina are continuous with the population in Brazil and Paraguay to the E.