Figs 43, 53

Diagnosis. Ectoedemia caradjai can be separated from all other species in the suberis group by the white basal patch on the forewings, and in the male by the absence of androconial scales surrounding the hairpencil on the hindwing. In the male genitalia the truncate gnathos is diagnostic, the valvae resemble those of E. phaeolepis, and somewhat E. heckfordi (Fig. 53).

Biology. Hostplants: Quercus pubescens Willd., Q. petraea (Matt.) Liebl., Q. pyrenaica Willd., Q. frainetto Ten., Q. infectoria Olivier, Q. coccifera L. (Van Nieukerken 1985; Van Nieukerken et al. 2004a) Larvae from July to October. Adults from April to September. Since in Croatia from July larvae adults still emerged in July and from August larvae in September, we assume that there are at least three generations (at least partly) in southern Europe.

Distribution. Spain, Portugal: Van Nieukerken et al. 2004a; Czech Republic, Slovakia: Laštůvka and Laštůvka 1990b; Russia: Van Nieukerken et al. 2004b; Bulgaria: Beiger 1980, here confirmed by adults. One specimen from Portugal, listed by Van

Nieukerken et al. (2004a), was a misidentification for E. phaeolepis, see there. New records from Spain: Laštůvka and Laštůvka 2008 and below.

Material. Bulgaria: 2♁, Montana, 25.VI.1998, A. & Z. Laštůvka (AL). Croatia: 3♁, 5♀, Dalmatia, Orašac, 12.VII.2000 mines with larvae, 1♁, 2♀ ex l. VII.2000, 23.VIII.2001, mines with larvae, 3♀ ex l. IX.2001, 22.VII.2005, mines with larvae, 2♁ ex l. VII.2005, A. Laštůvka leg. (AL), 67♁, 17♀, Krk, various localities, 1976–2008, G. Baldizzone, E. Jäckh (GB, RMNH, USNM). Greece, Crete: 1♁, Lasithi, Kritsa, 15.IV.1989, R. Johansson (RJ). Greece, Kerkyra: leafmines, Kerkyra, Markos, 22.X.1987, J.H. Donner (RMNH); Greece, Lesvos: 1♁, Sikaminia, 520 m, 13.IX.2008, L. Kaila & J. Kullberg (MZH); 1♁, 2 km NNE Megalochori, 770 m, 11.IX.2008, L. Kaila & J. Kullberg (MZH). Greece, Samos: 1♀, Samos, Kokkari, 17. VI.1996, R. Sutter (RS). Greece, mainland: 1♁, Akhaia, Kalavrita, 23. VI.1996, A. & Z. Laštůvka (AL); 1♁, 1♀, Arkádia, Sapounakeika, 15. VI.1998, A. & Z. Laštůvka (AL); 1♁, Arkadia, Techniti L. Ladonas, 25. VI.1991, P. Grotenfelt (MZH); 3♁, 6♀, Evro, 35 km N Alexandropolis, Kirki, 500m, 20–21.VIII.1980, M. Fibiger, A. Moberg (NHRS, ZMUC); 1♁, Evros, Alexandropolis, Kirki 2, 27.VII.1985, P. Grotenfelt (MZH); 1♁, 1♀, Florina, Vatochorion, 17. VI. 8.VIII.1985, P. Grotenfelt (MZH, RMNH); 2♁, Fokida, Itea, 26. V.1999 (1♁), 6. VI.1999 (1♁), A. & Z. Laštůvka (AL); 3♁, 2♀, Fthiotida, Agios Haralambos, 21.VI.1997, 22.VI.1998, 27.V.1999, A. & Z. Laštůvka (AL); 1♁, 2♀, Ilia, Loutra Killinis, 9.VIII. 9.X.1980, 19.VI.1981, P. Grotenfelt (MZH); 1♁, 1♀, Ioannina, Pindos Konitsa, Pades, 7. VI.1981, P. Grotenfelt (MZH); 3♁, 2♀, Korinthia, Agios Vassilios, 17. VI.1996, A. & Z. Laštůvka (AL); 4♁, Lakonia, Apidia, 18.VI.1996, 15.VI.1997, A. & Z. Laštůvka (AL); 1♁, 1♀, Messinia, Exohori, 18.VI.1997, 2.VI.1999, A. & Z. Laštůvka (AL); 1♁, Messinia, Filiatra, 18.IX.1980, P. Grotenfelt (MZH); 5♁, 1♀, Messinia, Kardamili, 20.VI.1996, 19.VI.1997, 17.VI.1998, A. & Z. Laštůvka (AL); 2♁, Pieria, Leptokaria, 22.VI.1997, 23.VI.1998, A. & Z. Laštůvka (AL); 1♁, Preveza, Thesprotiko, 11. VI.1997, A. & Z. Laštůvka (AL); 1♁, 1♀, Trikala, Oxinia, 8. VI.1999, A. & Z. Laštůvka (AL); 3♁, 1♀, Serres, Kalókastro, 24. VI.1997, A. & Z. Laštůvka (AL); 1♀, Serres, Strimon, delta, asl, 16.VIII.1985, A. Moberg (NHRS); 1♁, Thesprotia, Kallithea, 9. VI.1998, A. & Z. Laštůvka (AL). Portugal: 3♁, 2♀, [Beira Baixa] Castelo Branco, Capinha, 12. VI.2009, A. & Z. Laštůvka (AL); 1♁, 2♀, Santarém, Valverde, 14.VI.2009, A. & Z. Laštůvka (AL). Spain: 1♁, Tarragona, Baix Camp, Prades, Col de les Masies, 980 m, 13.VII.2006, E. Requena & De-Gregorio (RMNH). Turkey: 1♀, Adana, Taurus, 4 km NE Feke, 1000 m, 7.IX.1983, G. Derra (GD); vacated mines, Antalya, Termessos, ruins, 890–1000 m, 2.III.2005, Quercus coccifera, E.J. van Nieukerken (RMNH).