Published September 15, 2025
| Version v1
Journal article
Open
The threat-work: a network of potential threats to soil biodiversity
Authors/Creators
- 1. NOrthern Alberta InstitUte OF TechnOlOgy, 11762 - 106 Street, EdmOntOn, AB, Canada, T5G 2R1.
- 2. FOOd and AgricUltUre OrganizatiOn OF the United NatiOns, Viale delle Terme di Caracalla, ROme, 00153, Italy.
- 3. Wageningen UR, RadiX bUilding, DrOevendaalsesteeg 1, 6708 PB, Wageningen, P.O.BOX 100, 6700 AC Wageningen,
- 4. Embrapa FlOrestas, Estrada da Ribeira Km 111, COlOmbO, PR, 83411-000, Brazil.
Description
Lindo, Zoë, Battigelli, Jeff, Parnell, J. Jacob, Ruiter, Peter de, Brown, George G., Barreto, Carlos (2025): The threat-work: a network of potential threats to soil biodiversity. Soil Organisms 97 (6): 31-46, DOI: 10.25674/418, URL: http://dx.doi.org/10.25674/418
Files
source.pdf
Files
(661.3 kB)
| Name | Size | Download all |
|---|---|---|
|
md5:beea1639ec61a5d193209ba1989e432d
|
661.3 kB | Preview Download |
Linked records
Additional details
Identifiers
- LSID
- urn:lsid:plazi.org:pub:FFEA1639EC61FFD19320FFA1FF9E432D
References
- Addison, J. A. (2009). Distribution and impacts of invasive earthworms in Canadian forest ecosystems. Biological Invasions, 11, 59-79.
- Adhikari, K., & Hartemink, A. E. (2016). Linking soils to ecosystem services -A global review. Geoderma, 262, 101- 111.
- Anderson, J. M. (1975). The enigma of soil animal species diversity. In J. Vanek (Ed.), Progress in Soil Zoology (pp. 51- 58). Springer. https://doi.org/10.1007/978-94-010-1933-0_5
- Andriuzzi, W. S., Pulleman, M. M., Schmidt, O., Brussaard, L., & Faber, J. H. (2015). Anecic earthworms (Lumbricus terrestris) alleviate negative effects of extreme rainfall events on soil and plants in field mesocosms. Plant and Soil, 397, 103-113. https://doi.org/10.1007/s11104-015-2604-4
- Anthony, M. A., Bender, S. F., & van der Heijden, M. G. (2023). Enumerating soil biodiversity. Proceedings of the National Academy of Sciences, 120, e2304663120.
- Barreto, C., & Lindo, Z. (2018). Drivers of decomposition and the detrital invertebrate community differs across a hummock-hollow microtopology in boreal peatlands. Ecoscience, 25, 39-48. https://doi.org/10.1080/11956860.20 17.1412282
- Barreto, C., & Lindo, Z. (2022). Response of soil biodiversity to global change. Pedobiologia, 90, 150792. https://doi. org/10.1016/j.pedobi.2022.150792
- Barreto, C. R. A., Rillig, M., & Lindo, Z. (2020). Addition of polypropylene and polyester in soil affects decomposition rates but not microarthropod communities. Soil Organisms, 92(2), 109-119. https://doi.org/10.25674/so92iss2pp109
- Barreto, C., Conceicao, P. H. S., de Lima, E. C. A., Stievano, L. C., Zeppelini, D., Kolka, R. K., Hanson, P. J., & Lindo, Z. (2023). Large-scale experimental warming reduces soil faunal biodiversity through peatland drying. Frontiers in Environmental Science, 11, 1153683.
- Barreto, C., Buchkowski, R. W., & Lindo, Z. (2024). Restructuring of soil food webs reduces carbon storage potential in boreal peatlands. Soil Biology and Biochemistry, 193, 109413. https://doi.org/10.1016/j.soilbio.2024.109413
- Beaumelle, L., Thouvenot, L., Hines, J., Jochum, M., Eisenhauer, N., & Phillips, H. R. P. (2021). Soil fauna diversity and chemical stressors: A review of knowledge gaps and roadmap for future research. Ecography, 44, 845-859.
- Biswas, B., Qi, F., Biswas, J. K., Wijayawardena, A., Khan, M. A. I., & Naidu, R. (2018). The fate of chemical pollutants with soil properties in the climate change paradigm - A review. Soil Systems, 2(3), 51.
- Blankinship, J. C., Niklaus, P. A., & Hungate, B. A. (2011). A meta-analysis of responses of soil biota to global change. Oecologia, 165(3), 553-565. https://doi.org/10.1007/s00442- 011-1909-0
- Bodner, G., Zeiser, A., Keiblinger, K., Rosinger, C., Winkler, S. K., Stumpp, C., & Weninger, T. (2023). Managing the pore system: Regenerating the functional pore spaces of natural soils by soil-health oriented farming systems. Soil & Tillage Research, 234, 105862. https://doi.org/10.1016/j. still.2023.105862
- Brousseau, P.-M., Chauvat, M., De Almeida, T., & Forey, E. (2021). Invasive knotweed modifies predator -prey interactions in the soil food web. Biological Invasions, 23, 1987-2002. https://doi.org/10.1007/s10530-020-02431-0
- Brussaard, L., de Ruiter, P. C., & Brown, G. G. (2007). Soil biodiversity for agricultural sustainability. Agriculture, Ecosystems & Environment, 121(3), 233-244. https://doi. org/10.1016/j.agee.2006.12.013
- Buchkowski,R.W., Barreto,C.,&Lindo,Z.(2023).soilfoodwebs: An R package for analyzing and simulating nutrient fluxes through food webs. European Journal of Soil Biology, 119, 103556. https://doi.org/10.1016/j.ejsobi.2023.103556
- Cadel, M., Cousin, I., & Therond, O. (2023). Relationships between soil ecosystem services in temperate annual field crops: A systematic review. Science of the Total Environment, 902, 165930. https://doi.org/10.1016/j.scitotenv.2023.165930
- Camenzind, T., Mason-Jones, K., Mansour, I., Rillig, M. C., & Lehmann, J. (2023). Formation of necromass-derived soil organic carbon determined by microbial death pathways. Nature Geoscience, 16, 115-122. https://doi.org/10.1038/ s41561-022-01109-w
- Carrascosa, M., Sanchez-Moreno, S., & Alonso-Prados, J. L. (2015). Effects of organic and conventional pesticides on plant biomass, nematode diversity and the structure of the soil food web. Nematology, 17(1), 11-26. https://doi. org/10.1163/15685411-00002837
- Coleman, D. C. (2008). From peds to paradoxes: Linkages between soil biota and their influences on ecological processes. Soil Biology and Biochemistry, 40(2), 271-279. https://doi. org/10.1016/j.soilbio.2007.08.005
- Cotrufo, F. M., Wallenstein, M. D., Boot, C. M., Denef, K., & Paul, E. (2013). The Microbial Efficiency-Matrix Stabilization (MEMS) framework integrates plant litter decomposition with soil organic matter stabilization. Global Change Biology, 19(4), 988-995. https://doi.org/10.1111/gcb.12113
- Cramer, G. R., Lauchli, A., & Polito, V. S. (1985). Displacement of Ca2+ by Na+ from the plasmalemma of root cells. Plant Physiology, 79(1), 207-211. https://doi.org/10.1104/ pp.79.1.207
- de Castro, Berdugo, M., Eldridge, D.J., Eisenhauer, N., Singh, B.K., Cui, H., Abades, S., Alfaro, F.D., Bamigboye, A.R., Bastida, F., et al. (2021). Local stability properties of complex, species-rich soil food webs with functional block structure. Ecology and Evolution, 11(23), 16070-16081. https://doi. org/10.1002/ece3.8303
- de Vries, F. T., & Caruso, T. (2016). Eating from the same plate? Revisiting the role of labile carbon inputs in the soil food web. Soil Biology and Biochemistry, 102, 4-9. https://doi. org/10.1016/j.soilbio.2016.06.023
- Delgado-Baquerizo, M., Reich, P. B., Trivedi, C., Eldridge, D. J., Abades, S., Alfaro, F. D., ... & Singh, B. K. (2020). Multiple elements of soil biodiversity drive ecosystem functions across biomes. Nature Ecology & Evolution, 4, 210-220. https://doi. org/10.1038/s41559-019-1084-y
- Du, Y., Guo, S., Wang, R., Song, X., & Ju, X. (2023). Soil pore structure mediates the effects of soil oxygen on the dynamics of greenhouse gases during wetting-drying phases. Science of the Total Environment, 895, 165192. https://doi.org/10.1016/j. scitotenv.2023.165192
- FAO, ITPS, GSBI, SCBD, & EC. (2020). State of knowledge of soil biodiversity - Status, challenges and potentialities: Report 2020. FAO. https://doi.org/10.4060/cb1928en
- Gardi, C., Jeffery, S., & Saltelli, A. (2013). An estimate of potential threats levels to soil biodiversity in EU. Global Change Biology, 19(5), 1538-1548. https://doi.org/10.1111/ gcb.12159
- Geisen, S., Wall, D. H., & van der Putten, W. H. (2019). Challenges and opportunities for soil biodiversity in the Anthropocene. Current Biology, 29(19), R1036-R1044. https://doi.org/10.1016/j.cub.2019.08.007
- Giller, P. S. (1996). The diversity of soil communities: The 'poor man's tropical rainforest'. Biodiversity and Conservation, 5(2), 135-168. https://doi.org/10.1007/BF00055827
- Gonzalez, J. M., & Aranda, B. (2023). Microbial growth under limiting conditions -Future perspectives. Microorganisms, 11(7), 1641.https://doi.org/10.3390/microorganisms11071641
- Gonzalez, R., Kavouras, J., Barragan, V., Bernhard, E., Dawalibi, S., Diaz, M., ... & Pawlica, N. (2021). Use the metabolic fingerprint in microbial communities to evaluate the anthropogenic impact on soils. In FAO (Ed.), Keep soil alive, protect soil biodiversity - Global Symposium on Soil Biodiversity 19-22 April 2021: Proceedings. FAO. https:// www.fao.org/documents/card/en/c/cb1928en
- Gregory, A. S., Ritz, K., McGrath, S. P., Quinton, J. N., Goulding, K. W. T., Jones, R. J. A., ... & Whitmore, A. P. (2015). A review of the impacts of degradation threats on soil properties in the UK. Soil Use and Management, 31(1), 1-15. https://doi.org/10.1111/sum.12118
- Guerra, C. A., Heintz-Buschart, A., Sikorski, J., Chatzinotas, A., Guerrero-Ramirez, N., Cesarz, S., Beaumelle, L., Rillig, M.C., Maestre, F.T., Delgado-Baquerizo, M., et al. (2020a). Blind spots in global soil biodiversity and ecosystem function research. Nature Communications, 11, 3870. https://doi. org/10.1038/s41467-020-17688-2
- Guerra, C. A., Rosa, I. M. D., Valentini, E., Wolf, F., Filipponi, F., Karger, D. N., ... & Eisenhauer, N. (2020b). Global vulnerability of soil ecosystems to erosion. Landscape Ecology, 35, 823-842. https://doi.org/10.1007/s10980-020- 00992-9
- Guerra, C. A., Berdugo, M., Eldridge, D. J., Eisenhauer, N., Singh, B. K., Cui, H., ... & Delgado-Baquerizo, M. (2022). Global hotspots for soil nature conservation. Nature, 610(7930), 693-698. https://doi.org/10.1038/s41586-022- 05399-z
- Haddaway, N. R., Hedlund, K., Jackson, L. E., Katterer, T., Lugato, E., Thomsen, I. K., ... & Soderstrom, B. (2015). What are the effects of agricultural management on soil organic carbon in boreo-temperate systems? Environmental Evidence, 4, 23. https://doi.org/10.1186/s13750-015-0049-0
- Han, L., Chen, L., Feng, Y., Kuzyakov, Y., Chen, Q., Zhang, S., ... & Rillig, M. C. (2024). Microplastics alter soil structure and microbial community composition. Environment International, 185, 108508. https://doi.org/10.1016/j. envint.2024.108508
- Hayne, R. J., & Swift, R. S. (1990). Stability of soil aggregates in relation to organic constituents and soil water content. Journal of Soil Science, 41(1), 73-83. https://doi. org/10.1111/j.1365-2389.1990.tb00045.x
- Hopple, A. M., Wilson, R. M., Kolton, M., Zalman, C. A., Chanton, J. P., Kostka, J., ... & Bridgham, S. D. (2020). Massive peatland carbon banks vulnerable to rising temperatures. Nature Communications, 11, 2373. https://doi. org/10.1038/s41467-020-16311-8
- Hunt, H. W., Coleman, D. C., Ingham, E. R., Ingham, R. E., Elliott, E. T., Moore, J. C., ... & Morley, C. R. (1987). The detrital food web in a shortgrass prairie. Biology and Fertility of Soils, 3(1-2), 57-68. https://doi.org/10.1007/ BF00260580
- Husson, O. (2013). Redox potential (Eh) and pH as drivers of soil/plant/microorganism systems: A transdisciplinary overview pointing to integrative opportunities for agronomy. Plant and Soil, 362, 389-417. https://doi.org/10.1007/s11104- 012-1429-7
- Intergovernmental Panel on Climate Change (IPCC). (2022). Climate Change 2022: Impacts, Adaptation and Vulnerability. Cambridge University Press. https://doi. org/10.1017/9781009325844
- Jeffery, S., & Gardi, C. (2010). Soil biodiversity under threat: A review. Acta Societatis Zoologicae Bohemicae, 74, 7-12.
- Jeffery, S., Gardi, C., & Jones, A. (2010). European atlas of soil biodiversity. Publications Office of the European Union. https://data.europa.eu/doi/10.2788/94222
- Johnston, A. E., Poulton, P. R., & Coleman, K. (2009). Soil organic matter: Its importance in sustainable agriculture and carbon dioxide fluxes. In D. L. Sparks (Ed.), Advances in Agronomy (Vol. 101, pp. 1-57). Academic Press. https:// doi.org/10.1016/S0065-2113(08)00801-8
- Jones, S. K., Estrada-Carmona, N., Juventia, S. D., Dulloo, M. E., Laporte, M. A., Villani, C., & Remans, R. (2021). Agrobiodiversity Index scores show agrobiodiversity is underutilized in national food systems. Nature Food, 2(10), 712-723. https://doi.org/10.1038/s43016-021-00342-7
- Junting, Y., Xiaosong, L., Bo, W., Junjun, W., Bin, S., Changzhen, Y., & Zhihai, G. (2021). High spatial resolution topsoil organic matter content mapping across desertified land in Northern China. Frontiers in Environmental Science, 9, 668912. https://doi.org/10.3389/fenvs.2021.668912
- Klaminder, J., Krab, E. J., Larsbo, M., Jonsson, H., Fransson, J., & Koestel, J. (2023). Holes in the tundra: Invasive earthworms alter soil structure and moisture in tundra soils. Science of the Total Environment, 859, 160125. https://doi. org/10.1016/j.scitotenv.2022.160125
- Kronzucker, H. J., Coskun, D., Schulze, L. M., Wong, J. R., & Britto, D. T. (2013). Sodium as nutrient and toxicant. Plant and Soil, 369, 1-23. https://doi.org/10.1007/s11104-013- 1801-2
- Lal, R. (2004). Soil carbon sequestration to mitigate climate change. Geoderma, 123, 1-22. https://doi.org/10.1016/j. geoderma.2004.01.032
- Lancaster, S. H., Hollister, E. B., Senseman, S. A., & Gentry, T. J. (2010). Effects of repeated glyphosate applications on soil microbial community composition and the mineralization of glyphosate. Pest Management Science, 66(1), 59-64. https:// doi.org/10.1002/ps.1831
- Langmaid, K. K. (1964). Some effects of earthworm invasion in virgin podzols. Canadian Journal of Soil Science, 44, 34-37.
- Larsen, T., Schjonning, P., & Axelsen, J. (2004). The impact of soil compaction on euedaphic Collembola. Applied Soil Ecology, 26, 273-281.
- Li, J., Ramirez, G. H., Kiani, M., Quideau, S., Smith, E., Janzen, H., Larney, F., & Puurveen, D. (2018). Soil organic matter dynamics in long-term temperate agroecosystems: Rotation and nutrient addition effects. Canadian Journal of Soil Science, 98, 232-245.
- Liang, C., Amelung, W., Lehmann, J., & Kastner, M. (2019). Quantitative assessment of microbial necromass contribution to soil organic matter. Global Change Biology, 25, 3578-3590.
- Lindo, Z. (2015). Warming favours small-bodied organisms through enhanced reproduction and compositional shifts in belowground systems. Soil Biology and Biochemistry, 91, 271-278.
- Ma, X., Ni, X., Guo, Z., Zou, X., Chen, J., Shen, W., & Kuzyakov, Y. (2023). Nitrogen addition influences fine root growth and mycorrhizal symbiosis formation in trees with contrasting root morphology. Applied Soil Ecology, 189, 104987.
- Markkula, I., Cornelissen, J. H. C. R., & Aerts, R. (2019). Sixteen years of simulated summer and winter warming have contrasting effects on soil mite communities in a sub- Arctic peat bog. Polar Biology, 42, 581-591.
- McGill, W. B., & Spence, J. R. (1985). Soil fauna and soil structure: Feedbacks between size and architecture. Quaestiones Entomologicae, 21, 645-654.
- Meehan, M. L., Turnbull, K. F., Sinclair, B. J., & Lindo, Z. (2022). Predators minimize energy costs, not maximize energy gains when feeding under warming: Evidence from a microcosm experiment. Functional Ecology, 36, 2279-2288.
- Minasny, B., Malone, B. P., McBratney, A. B., Angers, D. A., Arrouays, D., Chambers, A., Chaplot, V., Chen, Z. S., Cheng, K., Das, B. S. et al. (2017). Soil carbon 4 per mille. Geoderma, 292, 59-86.
- Newbold, T., Hudson, L. N., Hill, S. L. L., & Purvis, A. (2015). Global effects of land use on local terrestrial biodiversity. Nature, 520, 45-50. https://doi.org/10.1038/nature14324
- Newman, M. M., Hoilett, N., Lorenz, N., Dick, R. P., Liles, M. R., Ramsier, C., & Kloepper, J. W. (2016). Glyphosate effects on soil rhizosphere-associated bacterial communities. Science of the Total Environment, 543, 155-160. https://doi. org/10.1016/j.scitotenv.2015.11.008
- Nicolescu, V.-N., Redei, K., Vor, T., Bastien, J.-C., Brus, R., Bencat, T., Dodan, M., Cvjetkovic, B., Andrasev, S., La Porta, N., Lavnyy, V., Petkova, K., Peric, S., Bartlett, D., Hernea, C., Pastor, M., Mataruga, M., Podrazsky, V., Sfecla, V., & Stefancik, I. (2020). A review of black walnut (Juglans nigra L.) ecology and management in Europe. Trees, 34, 1087-1112.
- Oades, J. M. (1984). Soil organic matter and structural stability: Mechanisms and implications for management. Plant and Soil, 76, 319-337.
- Omuto, C. T., Kome, G. K., Ramakhanna, S. J., Muzira, N. M., Ruley, J. A., Jayeoba, O. J., … & Nyamai, M. (2024). Trend of soil salinization in Africa and implications for agro-chemical use in semi-arid croplands. Science of the Total Environment, 951, 175503.
- Orgiazzi, A., Bardgett, R. D., & Barrios, E. (2016a). Global Soil Biodiversity Atlas. European Commission.
- Orgiazzi, A., Panagos, P., Yugini, Y., Dunbar, M. B., Gardi, C., Montanarella, L., & Ballabio, C. (2016b). A knowledge-based approach to estimating the magnitude and spatial patterns of potential threats to soil biodiversity. Science of the Total Environment, 545-546, 11-20.
- Pahalvi, H. N., Rafiya, L., Rashid, S., Nisar, B., & Kamili, A. N. (2021). Chemical fertilizers and their impact on soil health. In Dar et al. (Eds.), Microbiota and Biofertilizers (Vol. 2, pp. 1-20). Springer. https://doi.org/10.1007/978-3-030-61010- 4_1
- Palacino, B., Ascaso, S., Valero, A., & Valero, A. (2024). Regeneration costs of topsoil fertility: An exergy indicator of agricultural impacts. Journal of Environmental Management, 369, 122297.
- Pereira, M. C., O'Riordan, R., & Stevens, C. (2021). Urban soil microbial community and microbial-related carbon storage are severely limited by sealing. Journal of Soils and Sediments, 21, 1455-1465.
- Philippot, L., Chenu, C., Kappler, A., Rillig, M. C., & Fierer, N. (2024). The interplay between microbial communities and soil properties. Nature Reviews Microbiology, 22, 226-239.
- Porre, R. J., van Groenigen, J. W., De Deyn, G. B., de Goede, R. G. M., & Lubbers, I. M. (2016). Exploring the relationship between soil mesofauna, soil structure and N2O emissions. Soil Biology and Biochemistry, 96, 55-64.
- Potapov, A. M., Drescher, J., Darras, K., Wenzel, A., Janotta, N., Nazarreta, R., Kasmiatun, Laurent, V., Mawan, A., Utari, E. H., et al. (2024). Rainforest transformation reallocates energy from green to brown food webs. Nature, 627, 116-127.
- Potapov, A. M., Klarner, B., Sandmann, D., Widyastuti, R., & Scheu, S. (2019). Linking size spectrum, energy flux and trophic multifunctionality in soil food webs of tropical landuse systems. Journal of Animal Ecology, 88, 1845-1859.
- Powlson, D. S., Gregory, P. J., Whalley, W. R., Quinton, J. N., Hopkins, D. W., Whitmore, A. P., Hirsch, P. R., & Goulding, K. W. T. (2011). Soil management in relation to sustainable agriculture and ecosystem services.Food Policy, 36, S72-S87.
- Qiu, L., Zhang, Q., Zhu, H., Reich, P. B., Banerjee, S., van der Heidjen, M. G. A., Sadowsky, M. J., Ishii, S., Jia, X., Shao, M., Liu, B., Jiao, H., Li, H., & Wei, X. (2021). Erosion reduces soil microbial diversity, network complexity and multifunctionality. ISME, 15, 2474-2489.
- Reinhart, K. O., & Callaway, R. M. (2006). Soil biota and invasive plants. New Phytologist, 170, 445-457. https://doi. org/10.1111/j.1469-8137.2006.01715.x
- Rezapour, S., Nouri, A., Asadzadeh, F., Barin, M., Erpul, G., Jagadamma, S., & Qin, R. (2023). Combining chemical and organic treatments enhances remediation performance and soil health in saline-sodic soils. Communications Earth & Environment, 4, 285. https://doi.org/10.1038/s43247-023- 00948-6
- Rillig, M. C., Masahiro, R., Lehmann, A., Aguilar-Trigueros, C. A., Buchert, S., Wulf, A., Iwasaki, A., Roy, J., & Yang, G. (2019). The role of multiple global change factors in driving soil functions and microbial biodiversity. Science, 366, 886- 890. https://doi.org/10.1126/science.aay2832
- Rinke, A., Kuhry, P., & Dethloff, K. (2008). Importance of a soil organic layer for Arctic climate: A sensitivity study with an Arctic RCM. Geophysical Research Letters, 35, L13709.
- Rousk, J., Smith, A. R., & Jones, D. L. (2013). Investigating the long-term legacy of drought and warming on the soil microbial community across five European shrubland ecosystems. Global Change Biology, 19, 3872-3884.
- Scalenghe, R., & Marsan, F.A. (2009). The anthropogenic sealing of soils in urban areas. Landscape and Urban Planning, 90, 1-10.
- Semple, K. T., Morriss, A. W. J., & Paton, G. I. (2003). Bioavailability of hydrophobic organic contaminants in soils: Fundamental concepts and techniques for analysis. European Journal of Soil Science, 54, 809-818. https://doi.org/10.1046/ j.1351-0754.2003.0564.x
- Six, J., Bossuyt, H., Degryze, S., & Denef, K. (2004). A history of research on the link between (micro)aggregates, soil biota, and soil organic matter dynamics. Soil and Tillage Research, 79, 7-31.
- Soana, E., Vincenzi, F., Colombani, N., Mastrocicco, M., Fano, E. A., & Castaldelli, G. (2022). Soil denitrification, the missing piece in the puzzle of nitrogen budget in lowland agricultural basins. Ecosystems, 25, 633-647.
- Solly, E. F., Weber, V., Zimmermann, S., Walthert, L., Hagedorn, F., & Schmidt, M. W. I. (2020). A critical evaluation of the relationship between the effective cation exchange capacity and soil organic carbon content in Swiss Forest soils. Frontiers in Forests and Global Change, 3, 98. https://doi.org/10.3389/ ffgc.2020.00098
- Sperow, M. (2020). What might it cost to increase soil organic carbon using no-till on U.S. cropland? C arbon Balance and Management, 15, 26. https://doi.org/10.1186/s13021-020- 00162-3
- Standen, K. M., & Baltzer, J. L. (2021). Permafrost condition determines plant community composition and communitylevel foliar functional traits in a boreal peatland. Ecology and Evolution, 11, 10133-10146. https://doi.org/10.1002/ ece3.7818
- Stevenson, F. J. (1982). Humus chemistry. Wiley.
- Tibbett, M., Gil-Martinez, M., Fraser, T., Green, I. D., Duddigan, S., de Oliveira, V. H., Raulund-Rasmussen, K., Sizmur, T. A., & Diaz, A. (2019). Long-term acidification of pH neutral grasslands affects soil biodiversity, fertility and function in a heathland restoration. Catena, 180, 401-415.
- Tibbett, M., Fraser, T. D., & Duddigan, S. (2020). Identifying potential threats to soil biodiversity. PeerJ, 8, e9271. https:// doi.org/10.7717/peerj.9271
- Turnbull, M. S., & Lindo, Z. (2015). Combined effects of abiotic factors on Collembola communities reveal precipitation may act as a disturbance. Soil Biology and Biochemistry, 82, 36- 43.
- van der Putten, W., Klironomos, J., & Wardle, D. (2007). Microbial ecology of biological invasions. ISME Journal, 1, 28-37. https://doi.org/10.1038/ismej.2007.9
- Wagg, C., Bender, S. F., Widmer, F., & Van Der Heijden, M. G. (2014). Soil biodiversity and soil community composition determine ecosystem multifunctionality. Proceedings of the National Academy of Sciences, 111(14), 5266-5270.
- Wall, D. H., Bardgett, R. D., Behan-Pelletier, V., Herrick, J. E., Jones, T. H., Six, J., ... & Ritz, K. (Eds.). (2012). Soil ecology and ecosystem services. OUP Oxford.
- Wan, B., Liu, T., Gong, X., Zhang, Y., Li, C., Chen, X., Hu, F., Griffiths, B. S., & Liu, M. (2022). Energy fluxes across multitrophic levels drive ecosystem multifunctionality: Evidence from nematode food webs. Soil Biology and Biochemistry, 169, 108656.
- Wang, B., An, S., Liang, C., Liu, Y., & Kuzyakov, Y. (2021). Microbial necromass as the source of soil organic carbon in global ecosystems. Soil Biology and Biochemistry, 162, 108422.
- Wei, Z., Wu, S., Zhou, S., & Lin, C. (2013). Installation of impervious surface in urban areas affects microbial biomass, activity (potential C mineralisation), and functional diversity of the fine earth. Soil Research, 51(1), 59-67.
- Wu, H., Cui, H., Fu, C., Li, R., Qi, F., Liu, Z., Yang, G., Xiao, K., & Qiao, M. (2024). Unveiling the crucial role of soil microorganisms in carbon cycling: A review. Science of the Total Environment, 909, 168627.
- Zhang, L., Zhao, Z., Jiang, B., Baoyin, B., Cui, Z., Wang, H., Li, Q., & Cui, J. (2024). Effects of long-term application of nitrogen fertilizer on soil acidification and biological properties in China: A meta-analysis. Microorganisms, 12(7), 1683.
- Zhang, H., Wang, L., Fu, W., Xu, C., Zhang, H., Xu, X., Ma, H., Wang, J., & Zhang, Y. (2024). Soil acidification can be improved under different long-term fertilization regimes in a sweetpotato -wheat rotation system. Plants, 13(13), 1740.
- Zhang, Z., Qiao, M., Li, D., Yin, H., & Liu, Q. (2016). Do warming-induced changes in quantity and stoichiometry of root exudation promote soil N transformations via stimulation of soil nitrifiers, denitrifiers and ammonifiers? European Journal of Soil Biology, 74, 60-68.
- Zheng, J., Guo, R., Li, D., Zhang, J., & Han, S. (2017). Nitrogen addition, drought and mixture effects on litter decomposition and nitrogen immobilization in a temperate forest. Plant and Soil, 416, 165-179.