Published June 16, 2021
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Table 1 in Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data
Authors/Creators
- 1. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA
- 2. Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA
- 3. Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA
- 4. Department of Entomology, University of Wisconsin, Madison, WI 53706, USA
- 5. Institute of Biosciences, University of Rostock, Universitaetsplatz 2, D- 18055 Rostock, Germany
- 6. Department of Entomology, Coleoptera, Stuttgart State Museum of Natural History, Rosenstein 1, 70191 Stuttgart, Germany
- 7. Department of Entomology, University of Arizona, Tucson, AZ 85721 - 0036, USA
- 8. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Berkeley, 142 Weill Hall # 3200, Berkeley, CA 94720, USA
- 9. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Santa Cruz, Long Marine Lab, 117 McAllister Way, Santa Cruz, CA 95060, USA
Description
Table 1. Genus-group names in supertribe Nebriitae. All published genus-group names are represented in this list in order of publication and include author, year of publication, type species and current taxonomic status (Ledoux and Roux 2005; Lorenz 2005). The “Type” column indicates if the type species for the name is represented (Y) or not (N) in our taxon sample. The “Div” column lists the number of known species for each valid genus-group name. The “Rep” column indicates how many species-group taxa are represented in our taxon sample, and the "% Rep" column indicates what percentage of the known diversity is represented in our taxon sample. The "Proposed status" column indicates the appropriate status of the names based on the results of our study, with any changes in status noted in the “Result” column.
| Genus group name | Type species | Current status | Type | Div | Rep | % Rep | Proposed status | Result |
|---|---|---|---|---|---|---|---|---|
| Leistus Frölich, 1799 | Leistus testaceus Frölich = Carabus ferrugineus Linnaeus | valid genus and subgenus | Y | 52 | 11 | 21 | valid genus and subgenus | |
| Pogonophorus Latreille, 1802 | Carabus spinibarbis Fabricius | valid subgenus of Leistus | N | 60 | 3 | 5 | valid subgenus of Leistus | |
| Nebria Latreille, 1802 | Carabus brevicollis Fabricius | valid genus and subgenus | Y | 109 | 6 | 6 | valid genus and subgenus | |
| Manticora Panzer, 1803 | Manticora pallipes Panzer = Leistus spinibarbis Fabricius | junior homonym of Manticora Fabricius 1792 | N | - | - | - | junior homonym of Manticora Fabricius 1792 | |
| Notiophilus Duméril, 1805 | Cicindela aquatica Linnaeus | valid genus | N | 66 | 4 | 6 | valid genus | |
| Alpaeus Bonelli, 1810 | Carabus hellwigii Panzer | subjective junior synonym of Nebria s. str. | N | - | - | - | subjective junior synonym of Nebria s. str. | |
| Pelophila Dejean, 1821 | Carabus borealis Paykull | valid genus | Y | 2 | 2 | 100 | valid genus | |
| Helobia Stephens, 1828 | Carabus brevicollis Fabricius | objective junior synonym of Nebria s. str. | Y | - | - | - | objective junior synonym of Nebria s. str. | |
| Eonebria Semenov & Znojko, 1828 | Nebria komarovi Semenov & Znojko | valid subgenus of Nebria | Y | 77 | 6 | 7 | valid subgenus of Nebria | |
| Opisthius Kirby, 1837 | Opisthius richardsoni Kirby | valid genus | Y | 1 | 1 | 100 | valid genus | |
| Harpazobia Gistel, 1856 | Carabus brevicollis Fabricius | objective junior synonym of Nebria s. str. | Y | - | - | - | objective junior synonym of Nebria s. str. | |
| Eurynebria Ganglbauer, 1891 | Carabus complanatus Linnaeus | valid subgenus of Nebria | Y | 1 | 1 | 100 | valid subgenus of Nebria | |
| Leistidius Daniel, 1903 | Leistus piceus Frölich | subjective junior synonym of Leistus s. str. | N | - | - | - | subjective junior synonym of Leistus s. str. | |
| Oreonebria Daniel, 1903 | Alpaeus castaneus Bonelli | valid subgenus of Nebria | Y | 26 | 9 | 35 | valid subgenus of Nebria | |
| Oreobius Daniel, 1903 | Leistus gracilis Fuss | subjective junior synonym of Pogonophorus | N | - | - | - | subjective junior synonym of Pogonophorus | |
| Nebriola Daniel, 1903 | Nebria laticollis Dejean | valid subgenus of Nebria | Y | 19 | 2 | 11 | valid subgenus of Nebria | |
| Epinebriola Daniel & Daniel, 1904 | Nebria oxyptera Daniel & Daniel | valid subgenus of Nebria | Y | 28 | 6 | 21 | valid subgenus of Nebria | |
| Chaetoleistus Semenov,1904 | Leistus relictus Semenov | subjective junior synonym of Pogonophorus | N | - | - | - | subjective junior synonym of Pogonophorus | |
| Acroleistus Reitter, 1905 | Leistus denticollis Reitter | subjective junior synonym of Leistus s. str. | N | - | - | - | subjective junior synonym of Leistus s. str. | |
| Euleistulus Reitter, 1905 | Leistus ellipticus Reitter = Leistus fulvus Chaudoir | subjective junior synonym of Leistus s. str. | Y | - | - | - | subjective junior synonym of Leistus s. str. | |
| Leistophorus Reitter, 1905 | Leistus fulvibarbis Dejean | subjective junior synonym of Leistus s. str. | Y | - | - | - | subjective junior synonym of Leistus s. str. | |
| Spelaeonebria Peyerimhoff, 1911 | Spelaeonebria nudicollis Peyerimhoff | valid subgenus of Nebria | Y | 2 | 1 | 50 | valid subgenus of Nebria | |
| Eurinoleistus Breit, 1914 | Leistus depressus Breit | subjective junior synonym of Pogonophorus | N | - | - | - | subjective junior synonym of Pogonophorus | |
| Paropisthius Casey, 1920 | Opisthius indicus Chaudoir | valid genus | Y | 5 | 4 | 80 | valid genus | |
| Patrobonebria Bänninger, 1923 | Nebria desgodinsi Oberthür | valid subgenus of Nebria | Y | 11 | 4 | 36 | subjective junior synonym of Epinebriola | New Synonymy |
| Psilonebria Andrewes, 1923 | Nebria superna Andrewes | valid subgenus of Nebria | Y | 4 | 2 | 50 | valid subgenus of Nebria | |
| Nebrileistus Bänninger, 1925 | Leistus ellipticus Wollaston | valid subgenus of Leistus | N | 2 | 1 | 50 | valid subgenus of Leistus | |
| Archastes Jedlička, 1935 | Archastes sterbai Jedlička | valid genus | N | 42 | 1 | 2 | valid subgenus of Nebria | New Status |
| Boreonebria Jeannel, 1937 | Carabus rufescens Ström = Carabus gyllenhali Schönherr | valid subgenus of Nebria | Y | 42 | 19 | 45 | valid subgenus of Nebria | |
| Eunebria Jeannel, 1937 | Carabus psammodes Rossi | valid subgenus of Nebria | Y | 54 | 15 | 28 | valid subgenus of Nebria | |
| Paranebria Jeannel, 1937 | Carabus lividus Linnaeus | valid subgenus of Nebria | Y | 3 | 2 | 67 | subjective junior synonym of Epinebriola | New Synonymy |
| Neonebria Hatch, 1939 | none designated | nomen nudum | - | - | - | - | nomen nudum | |
| Nebriorites Jeannel, 1941 | Alpaeus gagates Bonelli | valid subgenus of Nebria | Y | 2 | 1 | 50 | valid subgenus of Nebria | |
| Alpaeonebria Csiki, 1946 | Nebria fuscipes Fuss | valid subgenus of Nebria | N | 32 | 1 | 3 | subjective junior synonym of Nebria s. str. | New Synonymy |
| Nippononebria Uéno, 1955 | Nebria pusilla Uéno | valid subgenus of Nebria | N | 7 | 2 | 29 | valid genus and subgenus | New Status |
| Agonoamara Jedlička, 1962 | Agonoamara chujoi Jedlička = Nebria chalceola Bates | subjective junior synonym of Nippononebria | Y | - | - | - | subjective junior synonym of Nippononebria s. str. | |
| Evanoleistus Jedlička, 1965 | Leistus nepalensis Jedlička | valid subgenus of Leistus | N | 154 | 9 | 5 | valid subgenus of Leistus | |
| Neoleistus Erwin, 1970 | Leistus ferruginosus Mannerheim | valid subgenus of Leistus | Y | 3 | 3 | 100 | subjective junior synonym of Leistus s. str. | New Synonymy |
| Germaria Jeanne, 1972 | Nebria bremii Germar | junior homonym of Germaria Robineau-Desvoidy, 1830 | Y | - | - | - | junior homonym of Germaria Robineau-Desvoidy, 1830 | |
| Catonebria Shilenkov, 1975 | Carabus nitidulus Fabricius = Nebria banksii Crotch | valid subgenus of Nebria | Y | 42 | 39 | 93 | valid subgenus of Nebria | |
| Orientonebria Shilenkov, 1975 | Nebria coreica Solsky | valid subgenus of Nebria | Y | 1 | 1 | 100 | valid subgenus of Nebria | |
| Reductonebria Shilenkov, 1975 | Nebria ochotica Sahlberg | valid subgenus of Nebria | Y | 17 | 15 | 83 | valid subgenus of Nebria | |
| Sardoleistus Perrault, 1980 | Leistus sardous Baudi di Selve | valid subgenus of Leistus | Y | 1 | 1 | 100 | valid subgenus of Leistus | |
| Tetungonebria Shilenkov, 1982 | Nebria tetungi Shilenkov | subjective junior synonym of Eunebria | N | - | - | - | incertae sedis | |
| Asionebria Shilenkov, 1982 | Nebria roborowskii Semenov | valid subgenus of Nebria | Y | 9 | 3 | 33 | subjective junior synonym of Psilonebria | New Synonymy |
| Notiokasis Kavanaugh & Nègre, 1983 | Notiokasis chaudoiri Kavanaugh & Nčgre | valid genus | N | 1 | 0 | 0 | valid genus | |
| Archileistobrius Shilenkov & Kryzhanovskij, 1983 | Archileistobrius hwangtienyuni Shilenkov & Kryzhanovskij | valid genus | N | 1 | 0 | 0 | incertae sedis | |
| Germarina Jeanne, 1985 | Nebria bremii Germar | junior homonym of Germarina Mesnil 1963 | Y | - | - | - | junior homonym of Germarina Mesnil 1963 | |
| Himalayonebria Ledoux, 1985 | Nebria nouristanensis Ledoux | subjective junior synonym of Epinebriola | N | - | - | - | incertae sedis | |
| Pseudonebriola Ledoux & Roux, 1989 | Nebria saurica Shilenkov | valid subgenus of Nebria | N | 18 | 7 | 39 | subjective junior synonym of Boreonebria | New Synonymy |
| Latviaphilus Barsevskis, 1994 | Elaphrus biguttatus Fabricius | subjective junior synonym of Notiophilus | N | - | - | - | subjective junior synonym of Notiophilus | |
| Makarovius Barsevskis, 1994 | Notiophilus rufipes Curtis | subjective junior synonym of Notiophilus | N | - | - | - | subjective junior synonym of Notiophilus | |
| Sphodronebria Sciaky & Pavesi, 1994 | Nebria paradoxa Sciaky & Pavesi = Nebria tetungi Shilenkov | objective junior synonym of Tetungonebria | N | - | - | - | objective junior synonym of Tetungonebria | |
| Vancouveria Kavanaugh, 1995 | Nebria virescens Horn | valid subgenus of Nebria | Y | 3 | 3 | 100 | valid subgenus of Nippononebria | |
| Ledouxnebria Deuve, 1998 | Ledouxnebria brisaci Deuve | valid genus (fossil only) | - | 1 | - | - | valid genus (fossil only) | |
| Epispadias Ledoux & Roux, 1999 | Nebria janschneideri Ledoux & Roux | valid subgenus of Nebria | N | 2 | 1 | 50 | valid subgenus of Nebria | |
| Falcinebria Ledoux & Roux, 2005 | Nebria reflexa Bates | valid subgenus of Nebria | Y | 16 | 2 | 13 | valid subgenus of Nebria | |
| Nakanebria Ledoux & Roux, 2005 | Nebria kurosawai Nakane | valid subgenus of Nebria | N | 6 | 1 | 17 | valid subgenus of Nebria | |
| Sadonebria Ledoux & Roux, 2005 | Nebria sadona Bates | valid subgenus of Nebria | N | 17 | 4 | 24 | valid subgenus of Nebria | |
| Tyrrhenia Ledoux & Roux, 2005 | Carabus rubicundus Quesnel | valid subgenus of Nebria | Y | 18 | 2 | 11 | valid subgenus of Nebria | |
| Marggia Huber, 2014 | Nebria bremii Germar | valid subgenus of Nebria | Y | 2 | 1 | 50 | subjective junior synonym of Oreonebria | New Synonymy |
| Barbonebriola Huber & Schmidt, 2017 | Nebria barbata Andrewes | valid subgenus of Nebria | N | 6 | 1 | 17 | subjective junior synonym of Epinebriola | New Synonymy |
| Archaeonebria Kavanaugh & Schmidt, 2019 | Archaeonebria inexspectata Schmidt & Kavanaugh | valid genus (fossil only) | - | 1 | - | - | valid genus (fossil only) | |
| Parepinebriola subgen. nov. | Nebria delicata Huber & Schmidt | part of Epinebriola Daniel | Y | 5 | 5 | ? | new subgenus | New Subgenus |
| Insulanebria subgen nov. | Nebria snowi Bates | part of Reductonebria Shilenkov | Y | 2 | 2 | 100 | new subgenus | New Subgenus |
| Erwinebria subgen. nov. | Nebria sahlbergii Fisher von Waldheim | part of Reductonebria Shilenkov | Y | 21 | 20 | 95 | new subgenus | New Subgenus |
| Nivalonebria subgen. nov. | Nebria paradisi Darlington | part of Nakanebria Ledoux & Roux | Y | 2 | 2 | 100 | new subgenus | New Subgenus |
| Neaptenonebria subgen. nov. | Nebria ovipennis LeConte | part of Catonebria Shilenkov | Y | 6 | 6 | 100 | new subgenus | New Subgenus |
| Palaptenonebria subgen. nov. | Nebria mellyi Gebler | part of Catonebria Shilenkov | Y | 14 | 11 | 79 | new subgenus | New Subgenus |
Notes
Published as part of Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya & Chen, Jei-Ying, 2021, Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data, pp. 41 in ZooKeys 1044 on page 41, DOI: 10.3897/zookeys.1044.62245
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Additional details
Identifiers
References
- Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon
- Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera "Geadephaga":Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, [ii +] 530 pp.
- Fabricius, 1792. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom. I. Pars I. C. G. Proft, Hafniae, [xx +] 330 pp. https://doi.org/10.5962/bhl.title.122153