Biodiversity Literature Repository
Published December 31, 2022 | Version v1
Journal article Open

A revision of the genus Mesobuthus Vachon 1950 with a description of 14 new species (Scorpiones Buthidae)

Creators

Description

Kovařík, František, Fet, Victor, Gantenbein, Benjamin, Graham, Matthew R., Aydin, Ersen, Yağmur, Šťáhlavský, František, Nikita, Poverennyi, Novruzov, Nizami E. (2022): A revision of the genus Mesobuthus Vachon 1950 with a description of 14 new species (Scorpiones Buthidae). Euscorpius 348: 1-189, DOI: 10.5281/zenodo.7162849

Files

source.pdf

Files (64.0 MB)

Name Size Download all
md5:7211ad771066f11d9344573321570887
64.0 MB Preview Download

Linked records

Additional details

Cites
Publication: 10.1093/bioinformatics/btu531 (DOI)
Publication: 10.1038/s41598-021-91001-z (DOI)
Publication: 10.1093/sysbio/syy032 (DOI)
Publication: 10.1101/2021.10.31.466646 (DOI)
Has part
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1060F11B90C4515E2495F8EE (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1075F10E93F454FA2331FB02 (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:F24CCA40-7CB7-4CB2-B55B-CCF8F03825E4 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1072F105909455BE242EF852 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F107FF10093FA57D922ACFEC5 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F107BF13F908D565E24EBFAA5 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1044F137900B527E24C5FE05 (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:77238E6D-756F-47BF-9D3D-4BB5E1F82E56 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F104CF1329096569E2593F785 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1056F12893B857D9239DFA6D (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:D625796C-166A-4037-900D-62A53BE66C33 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1053F127925451272462FAE5 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F105CF15D90D552BE239BFA65 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1026F15D93E2513E2254F88E (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:A1C3DA05-7ADF-4C85-A317-21DB661EF581 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1022F15A9089567E21E1F865 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1021F14C907B57D9236EF9A4 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1037F14C93F5517E2257F8CE (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:E81C1A94-70F4-4176-9255-F717E063D1C1 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1031F140924653DD230CF965 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F103BF17F9337503E243BFC85 (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:6560F74F-3E28-44EA-ADC7-4CBE8367E659 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1004F17590F6541E2494FE24 (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:B2043A81-30E4-4EBC-A0A8-66A7E20DBF56 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F100EF17090F056FE2524FA65 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F100BF17090CC513E25B2F94E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1015F165907354FE2342F861 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F101EF165901D531E2494FAAE (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:349B1277-250E-47A1-AFB8-5FFF16D9EA4F (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F1018F19E9245553E2253FA8A (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10E5F19093F45204206AF87B (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:82742C1C-C052-445A-9C19-6672CC79DBD9 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10EBF18F908C531024E4FBA5 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10F4F18F9032537E2495FACE (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:2CBE605F-A7E2-4274-B390-26FDD500BFF1 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10FEF183908E57D92396FA06 (URL)
Taxonomic treatment: urn:lsid:zoobank.org:act:1B02D85C-F08F-4A07-9550-70505A41A6CA (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10F8F18393DE529F2254F9E0 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10C5F1B9901F559E2300FC25 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/8E28D50F10C2F1B7924754FE230BF91C (URL)
Figure: 10.5281/zenodo.7162855 (DOI)
Figure: 10.5281/zenodo.7162857 (DOI)
Figure: 10.5281/zenodo.7162859 (DOI)
Figure: 10.5281/zenodo.7162861 (DOI)
Figure: 10.5281/zenodo.7162863 (DOI)
Figure: 10.5281/zenodo.7162867 (DOI)
Figure: 10.5281/zenodo.7162871 (DOI)
Figure: 10.5281/zenodo.7162873 (DOI)
Figure: 10.5281/zenodo.7162877 (DOI)
Figure: 10.5281/zenodo.7162882 (DOI)
Figure: 10.5281/zenodo.7162884 (DOI)
Figure: 10.5281/zenodo.7162888 (DOI)
Figure: 10.5281/zenodo.7162890 (DOI)
Figure: 10.5281/zenodo.7162892 (DOI)
Figure: 10.5281/zenodo.7162894 (DOI)
Figure: 10.5281/zenodo.7162896 (DOI)
Figure: 10.5281/zenodo.7162898 (DOI)
Figure: 10.5281/zenodo.7162900 (DOI)
Figure: 10.5281/zenodo.7162902 (DOI)
Figure: 10.5281/zenodo.7162906 (DOI)
Figure: 10.5281/zenodo.7162908 (DOI)
Figure: 10.5281/zenodo.7162910 (DOI)
Figure: 10.5281/zenodo.7162912 (DOI)
Figure: 10.5281/zenodo.7162914 (DOI)
Figure: 10.5281/zenodo.7162916 (DOI)
Figure: 10.5281/zenodo.7162918 (DOI)
Figure: 10.5281/zenodo.7162920 (DOI)
Figure: 10.5281/zenodo.7162922 (DOI)
Figure: 10.5281/zenodo.7162924 (DOI)
Figure: 10.5281/zenodo.7162926 (DOI)
Figure: 10.5281/zenodo.7162928 (DOI)
Figure: 10.5281/zenodo.7162930 (DOI)
Figure: 10.5281/zenodo.7162932 (DOI)
Figure: 10.5281/zenodo.7162934 (DOI)
Figure: 10.5281/zenodo.7162936 (DOI)
Figure: 10.5281/zenodo.7162938 (DOI)
Figure: 10.5281/zenodo.7162942 (DOI)
Figure: 10.5281/zenodo.7162944 (DOI)
Figure: 10.5281/zenodo.7162946 (DOI)
Figure: 10.5281/zenodo.7162948 (DOI)
Figure: 10.5281/zenodo.7162950 (DOI)
Figure: 10.5281/zenodo.7162956 (DOI)
Figure: 10.5281/zenodo.7162960 (DOI)
Figure: 10.5281/zenodo.7162962 (DOI)
Figure: 10.5281/zenodo.7162966 (DOI)
Figure: 10.5281/zenodo.7162968 (DOI)
Figure: 10.5281/zenodo.7162970 (DOI)
Figure: 10.5281/zenodo.7162976 (DOI)
Figure: 10.5281/zenodo.7162978 (DOI)
Figure: 10.5281/zenodo.7162980 (DOI)
Figure: 10.5281/zenodo.7162982 (DOI)
Figure: 10.5281/zenodo.7162984 (DOI)
Figure: 10.5281/zenodo.7162990 (DOI)
Figure: 10.5281/zenodo.7162992 (DOI)
Figure: 10.5281/zenodo.7163004 (DOI)
Figure: 10.5281/zenodo.7163006 (DOI)
Figure: 10.5281/zenodo.7163010 (DOI)
Figure: 10.5281/zenodo.7163014 (DOI)
Figure: 10.5281/zenodo.7163016 (DOI)
Figure: 10.5281/zenodo.7163018 (DOI)
Figure: 10.5281/zenodo.7163020 (DOI)
Figure: 10.5281/zenodo.7163024 (DOI)
Figure: 10.5281/zenodo.7163467 (DOI)
Figure: 10.5281/zenodo.7163030 (DOI)
Figure: 10.5281/zenodo.7163034 (DOI)
Figure: 10.5281/zenodo.7163036 (DOI)
Figure: 10.5281/zenodo.7163038 (DOI)
Figure: 10.5281/zenodo.7163040 (DOI)
Figure: 10.5281/zenodo.7163042 (DOI)
Figure: 10.5281/zenodo.7163044 (DOI)
Figure: 10.5281/zenodo.7163046 (DOI)
Figure: 10.5281/zenodo.7163049 (DOI)
Figure: 10.5281/zenodo.7163051 (DOI)
Figure: 10.5281/zenodo.7163055 (DOI)
Figure: 10.5281/zenodo.7163058 (DOI)
Figure: 10.5281/zenodo.7163064 (DOI)
Figure: 10.5281/zenodo.7163066 (DOI)
Figure: 10.5281/zenodo.7163070 (DOI)
Figure: 10.5281/zenodo.7163072 (DOI)
Figure: 10.5281/zenodo.7163074 (DOI)
Figure: 10.5281/zenodo.7163076 (DOI)
Figure: 10.5281/zenodo.7163078 (DOI)
Figure: 10.5281/zenodo.7163082 (DOI)
Figure: 10.5281/zenodo.7163086 (DOI)
Figure: 10.5281/zenodo.7163088 (DOI)
Figure: 10.5281/zenodo.7163090 (DOI)
Figure: 10.5281/zenodo.7163094 (DOI)
Figure: 10.5281/zenodo.7163096 (DOI)
Figure: 10.5281/zenodo.7163098 (DOI)
Figure: 10.5281/zenodo.7163102 (DOI)
Figure: 10.5281/zenodo.7163104 (DOI)
Figure: 10.5281/zenodo.7163108 (DOI)
Figure: 10.5281/zenodo.7163111 (DOI)
Figure: 10.5281/zenodo.7163113 (DOI)
Figure: 10.5281/zenodo.7163117 (DOI)
Figure: 10.5281/zenodo.7163121 (DOI)
Figure: 10.5281/zenodo.7163123 (DOI)
Figure: 10.5281/zenodo.7163125 (DOI)
Figure: 10.5281/zenodo.7163127 (DOI)
Figure: 10.5281/zenodo.7163129 (DOI)
Figure: 10.5281/zenodo.7163131 (DOI)
Figure: 10.5281/zenodo.7163133 (DOI)
Figure: 10.5281/zenodo.7163135 (DOI)
Figure: 10.5281/zenodo.7163137 (DOI)
Figure: 10.5281/zenodo.7163139 (DOI)
Figure: 10.5281/zenodo.7163143 (DOI)
Figure: 10.5281/zenodo.7163145 (DOI)
Figure: 10.5281/zenodo.7163147 (DOI)
Figure: 10.5281/zenodo.7163151 (DOI)
Figure: 10.5281/zenodo.7163155 (DOI)
Figure: 10.5281/zenodo.7163157 (DOI)
Figure: 10.5281/zenodo.7163159 (DOI)
Figure: 10.5281/zenodo.7163161 (DOI)
Figure: 10.5281/zenodo.7163165 (DOI)
Figure: 10.5281/zenodo.7163167 (DOI)
Figure: 10.5281/zenodo.7163169 (DOI)
Figure: 10.5281/zenodo.7163171 (DOI)
Figure: 10.5281/zenodo.7163173 (DOI)
Figure: 10.5281/zenodo.7163175 (DOI)
Figure: 10.5281/zenodo.7163177 (DOI)
Figure: 10.5281/zenodo.7163179 (DOI)
Figure: 10.5281/zenodo.7163181 (DOI)
Figure: 10.5281/zenodo.7163185 (DOI)
Figure: 10.5281/zenodo.7163187 (DOI)
Figure: 10.5281/zenodo.7163189 (DOI)
Figure: 10.5281/zenodo.7163191 (DOI)
Figure: 10.5281/zenodo.7163193 (DOI)
Figure: 10.5281/zenodo.7163195 (DOI)
Figure: 10.5281/zenodo.7163197 (DOI)

References

  • ADILARDI, R. S., A. A. OJANGUREN-AFFILASTRO, D. A. MARTI & L. M. MOLA. 2020. Chromosome puzzle in the southernmost populations of the medically important scorpion Tityus bahiensis (Perty 1833) (Buthidae), a polymorphic species with striking structural rearrangements. Zoologischer Anzeiger, 288: 139-150.
  • AITCHISON, J. E. T. 1888. The botany of the Afghan Delimitation Commission. Transactions of Linnean Society of London, Botany, 3(1): 1-139.
  • ANIKIN, V. V. 1997. [Rare species of Insecta and Arachnida in marginal structures of the steppe zone of the Saratov Province]. Pp. 28-29 in: Problemy izuchenii a kraevykh struktur biotsenozov [Problems of study of marginal structures]. Saratov (in Russian).
  • ANIKIN, V. V. 2006. [Rare and endangered species of terrestrial invertebrates in the Saratov Province]. Povolzhskii ekologicheskii zhurnal [Povolzhye Ecological Journal], Vyp. Spets. [Special Issue]: 47-56 (in Russian).
  • ANIKIN, V. V. & E. A. KIREEV. 1998. [Ecological-floristic description of the habitats of some rare species of insects and arachnids of the Lower Volga region]. Pp. 106-107 in: Materials of the International Research-Practical Conference "Problemy sokhraneniia bioraznoobraziia aridnykh regionov Rossii" [Problems of Conservation of Biodiversity of the Arid Regions of Russia]. Volgograd, 11-17 September 1998 (in Russian).
  • ANIKIN, V. V. & N. M. POVERENNYI. 2017. [Establishing the taxonomic status of scorpions of the genus Mesobuthus (Arachnida: Scorpiones) from the Lower Povolzhye based on coxI DNA sequence analysis]. Entomologicheskie i parazitologicheskie issledovaniia v Povolzhye [Entomological and Parasitological Research in Povolzhye], 14: 20-24 (in Russian).
  • ANIKIN, V. V. & A. S. SAZHNEV. 2016. [Terrestrial imvertebrate species recommended for the new edition of the Red Data Book of the Saratov Province]. Ivzestiia Saratovskogo universiteta. Nov. ser. Khimiia. Biologiia. Ekologiia [Proceedings of the Saratov University. New Ser. Chemistry, Biology, Ecology], 16(3): 313-317 (in Russian).
  • ATAMURADOV, K. I. 1994. Paleogeography of Turkmenistan. Pp. 49-64 in: Fet, V. & K. I. Atamuradov (eds), Biogeography and Ecology of Turkmenistan. Kluwer Academic Publishers, Dordrecht.
  • BARAHOEI, H., S. NAVIDPOUR, M. ALIABADIAN, R. SIAHSARVIE & O. MIRSHAMSI. 2020. Scorpions of Iran (Arachnida: Scorpiones): Annotated checklist, delta database and identification key. Journal of Insect Biodiversity and Systematics, 6(4): 375-474.
  • BIRULA, A. A. 1896. Miscellanea scorpiologica. I. Zur Synonymie der russischen Skorpione. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St.- Petersbourg, 1: 229-245.
  • BIRULA, A. A. 1900a. Beitrage zur Kenntniss der Scorpionenfauna Ost-Persiens. (1. Beitrag). Bulletin de l'Academie Imperiale des Sciences de St.-Petersbourg, (5), 12(4): 355-375.
  • BIRULA, A. A. 1900b. Scorpiones mediterranei Musei Zoologici mosquensis. Izvestiya Imperatorskago Obshchestva Lyubitelei Prirody, Istorii, Antropologii i Etnografii (Societas Caesarea Amicorum Rerum Naturalium, Anthropologiae Ethnographiae Universitatis Moscoviensis), 98, 3(1): 8-20 (in Russian).
  • BIRULA, A. A. 1903. Beitrage zur Kenntniss der Scorpionenfauna Ost-Persiens. (Zweiter Beitrag). Bulletin de l'Academie Imperiale des Sciences de St.-Petersbourg, (5), 19(2): 67-80.
  • BIRULA, A. A. 1904a. Miscellanea scorpiologica. VI. Ueber die Buthus -Arten des Centralasiens nebst ihrer geographischen Verbreitung. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St.- Petersbourg, 9: 20-27.
  • BIRULA, A. A. 1904b. Miscellanea scorpiologica. VII. Synopsis der russischen Skorpione. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St.- Petersbourg, 9: 28-38.
  • BIRULA, A. A. 1905a. Beitrage zur Kenntniss der Scorpionenfauna Persiens (Dritter Beitrag). Bulletin de l'Academie Imperiale des Sciences de St.-Petersbourg, 23: 119-148.
  • BIRULA, A. A. 1905b. Miscellanea Scorpiologica. VIII. Bemerkunger ueber die Skorpione-Sammlung des Kaukasischer Museum zu Tiflis. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St.- Petersbourg, 10: 119-131.
  • BIRULA, A. A. 1911a. Arachnologische Beitrage. I. Zur Scorpionen- und Solifugen-Fauna des Chinesischen Reichs. Revue Russe d'Entomologie, 11: 195-201.
  • BIRULA, A. A. 1911b. Miscellanea scorpiologica. IX. Ein Beitrag zur Kenntniss der Skorpionenfauna des Russischen Reiches und der angrenzender Lander. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg, 16: 161-179.
  • BIRULA, A. A. 1912. Ein Beitrag zur Kenntniss der Skorpionenfauna des Kaukasuslander. Izvestiya Kavkazskago muzeya (Mitteilungen des Kaukasischen Museums), 7(1): 117-127. [BIRULA, A. A.] BYALYNITSKII-BIRULYA, A. A. 1917a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskago Muzeya (Memoires du Musee du Caucase), Tiflis: Imprimerie de la Chancellerie du Comite pour la Transcaucasie, A(5), 253 pp. (in Russian; published August 1917) (in Russian). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Jerusalem: Israel Program for Scientific Translations, 170 pp. [BIRULA, A. A.] BYALYNITSKII-BIRULYA, A. A. 1917b. Faune de la Russie et des pays limitrophes fondee principalement sur les collections du Musee Zoologique de l'Academie des Sciences de Russie. Arachnides (Arachnoidea). Petrograd, 1(1): xx, 227 (in Russian). English translation: Byalynitskii-Birulya, A. A. 1965. Fauna of Russia and Adjacent Countries. Arachnoidea. Vol. I. Scorpions. Jerusalem: Israel Program for Scientific Translations, xix, 154 pp.
  • BIRULA, A. A. 1918. Miscellanea scorpiologica. XI. (Materiaux pour servir a la scorpiofaune de laMesopotamie inferieure, du Kurdistan et de la Perse septentrionale). Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de Russie, 27: 1-44 (in Russian).
  • BIRULA, A. A. 1925. Skorpiologische Beitrage. 10. Zur geographischen Verbreitung zweier weitverbreiteter Skorpionen-Arten Palaearcticums. Zoologischer Anzeiger, 63: 62-67.
  • BIRULA, A. A. 1927. Zoologische Ergebnisse der von P. K. Kozlov in den Jahren 1925-1926 ausgefuhrten Expedition nach der Mongolei. I. Skorpione und Solifugen. Annuaire du Musee Zoologique de l'Academie des Sciences d'URSS, 28: 201-218.
  • BIRULA, A. A. 1928. K voprosu o nizhnem techenii Volgi kak zoogeograficheskoi granitse [On the Lower Volga as a zoogeographic border]. Doklady Akademii Nauk SSSR 1928: 338-340 (in Russian).
  • BRAGINA, T. M. & E. A. YAGMUR. 2014. New data on the distribution of Mesobuthus eupeus thersites (C.L. Koch, 1839) (Scorpiones: Buthidae) in Central Kazakhstan. Serket, 14(1):1-5.
  • BRYSON, R. W., L. PRENDINI, W. E. SAVARY & P. B. PEARMAN. 2014. Caves as microrefugia: Pleistocene phylogeography of the troglophilic North American scorpion Pseudouroctonus reddelli. BMC Evolutionary Biology, 14(1): 1-17.
  • CRUCITTI, P. & D. CICUZZA. 2000. Gli Scorpioni del Parco Nazionale del Monte Nemrut (Turchia sud-orientale) (Scorpiones). Memorie della Societa entomologica italiana, 78(2): 275-294.
  • CRUCITTI, P. & D. CICUZZA. 2001a. Ricerche bioecologiche su Mesobuthus eupeus del Monte Ararat (Turchia orientale) (Scorpiones: Buthidae). Boll. Soc. Entomol. Ital. 133(1): 3-11.
  • CRUCITTI, P. & D. CICUZZA. 2001b. Scorpions of Anatolia: Ecological patterns. Pp. 225-234 in Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society.
  • CRUCITTI, P. & V. VIGNOLI. 2002. Gli Scorpioni (Scorpiones) dell'Anatolia sud-orientale (Turchia). Bolletino del Museo Regionale di Scienze Naturali, Torino, 19(2): 433-480.
  • DERMITZAKIS, D. M. & D. J. PAPANIKOLAOU. 1981. Paleogeography and geodynamics of the Aegean region during the Neogene. Annales Geologique des Pays Hellenique, 30: 245-289.
  • DRUMMOND, A. J., M. A. SUCHARD, D. XIE & A. RAMBAUT. 2012. Bayesian phylogenetics with BEAUti and the BEAST 1.7. Molecular Biology and Evolution, 29: 1969-1973.
  • EDGAR, R. C. 2004. MUSCLE: multiple sequence alignment with high accuracy and high throughput. Nucleic Acids Research, 32: 1792-1797.
  • FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali "Enrico Caffi", 13(1988): 73-171.
  • FET, V. 1994. Fauna and zoogeography of scorpions (Arachnida: Scorpiones) in Turkmenistan. Pp. 525-534 in Fet, V. & K. I. Atamuradov (eds.). Biogeography and Ecology of Turkmenistan. Monographiae Biologicae 72. Dordrecht: Kluwer Academic Publishers.
  • FET, V. 2010. Scorpions of Europe. Acta Zoologica Bulgarica, 62: 3-12.
  • FET, V., B. GANTENBEIN, A. V. GROMOV, G. LOWE & W. R. LOURENCO. 2003. The first molecular phylogeny of Buthidae (Scorpiones). Euscorpius, 4: 1-10.
  • FET, V., F. KOVARIK, B. GANTENBEIN & M. R. GRAHAM. 2021. Three new cryptic species of Olivierus (Scorpiones: Buthidae) from Kazakhstan and Uzbekistan. Zootaxa, 5006(1): 54-72.
  • FET, V., F. KOVARIK, B. GANTENBEIN, R. C. KAISER, A. K. STEWART & M. R. GRAHAM. 2018. Revision of the Mesobuthus caucasicus complex from Central Asia, with descriptions of six new species (Scorpiones: Buthidae). Euscorpius, 255: 1-77.
  • FET, V. & G. LOWE. 2000. Family Buthidae. Pp. 54-286 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder (eds.). Catalog of the Scorpions of the World (1758-1998). New York: New York Entomological Society, 690 pp.
  • FET, V., M. E. SOLEGLAD & S. L. ZONSTEIN. 2011. The genus Akrav Levy, 2007 (Scorpiones: Akravidae) revisited. Euscorpius, 134: 1-49.
  • FET, V., M. E. SOLEGLAD, S. L. ZONSTEIN, I. NAAMAN, S. LUBATON, B. LANGFORD & A. FRUMKIN. 2017. The second record of a relict Akrav israchanani Levy, 2007 (Scorpiones: Akravidae) from Levana Cave, Israel. Euscorpius, 247: 1-12.
  • FOMICHEV, A. A. 2011. A new locality of Mesobuthus eupeus thersites (C. L. Koch, 1839) (Scorpiones: Buthidae) in East Kazakhstan. Euscorpius, 136: 1-3.
  • GANTENBEIN, B., V. FET, I. A. GANTENBEIN-RITTER & F. BALLOUX. 2005. Evidence for recombination in scorpion mitochondrial DNA (Scorpiones: Buthidae). Proceedings of the Royal Society: Biological Sciences, 272(1564): 697-704.
  • GANTENBEIN, B., V. FET & A. V. GROMOV. 2003. The first DNA phylogeny of four species of Mesobuthus Vachon, 1950 (Scorpiones: Buthidae) from Eurasia. Journal of Arachnology, 31: 412-420.
  • GHARAKHLOO, M. M., S. A. HEYDARABAD & E. A. YAGMUR. 2018. The scorpion fauna of West Azerbaijan Province in Iran (Arachnida: Scorpiones). Biharean Biologist, 12(2): 84-87.
  • GRAHAM, M. R., E. A. MYERS, R. KAISER & V. FET. 2019. Cryptic species and co-diversification in sand scorpions from the Karakum and Kyzylkum deserts of Central Asia. Zoologica Scripta, 48(6): 801-812.
  • GRAHAM, M. R., V. OLAH-HEMMINGS & V. FET. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a longstanding component of Central Asian biogeography (Scorpiones: Buthidae). Zoology in the Middle East, 55: 95-110.
  • GRAHAM, M. R., D. A. WOOD, J. A. HENAULT, Z. J. VALOIS & P. E. CUSHING. 2017. Ancient lakes, Pleistocene climates and river avulsions structure the phylogeography of a large but little-known rock scorpion from the Mojave and Sonoran deserts. Biological Journal of the Linnean Society, 122(1): 133-146.
  • GROMOV, A. V. 2001. The northern boundary of scorpions in Central Asia. Pp. 301-306 in Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society.
  • GROMOV, A. V. & Ye. Ye. KOPDYKBAEV. 1994. [The fauna of the scorpions and solpugids (Arachnida: Scorpiones, Solifugae) in Kazakhstan]. Selevinia, 2(2): 19-23 (in Russian).
  • HEDDERGOTT, M., M. STUBBE, W. STUBBE, P. STEINBACH & A. STUBBE. 2016. Geographical distribution of the genus Mesobuthus (Scorpiones: Buthidae) in Mongolia. Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, 13: 147-164.
  • HUSSEN, F. S. & S. T. AHMED. 2020. New data of scorpion fauna, include two new records with identification key of scorpion species (Arachnida: Scorpiones) in Iraq. Plant Archives, 20 (2): 6711-6725.
  • KACHEL, H. S, A. M. AL-KHAZALI, F. S. HUSSEN & E. A.YAGMUR. 2020. Checklist and review of the scorpion fauna of Iraq (Arachnida: Scorpiones). Arachnologische Mitteilungen, 61(1): 1-10.
  • KALTSAS, D., I. STATHI & V. FET. 2008. Scorpions of the Eastern Mediterranean. Pp. 209-246 in Makarov, S.E. & R.N. Dimitrijevic (eds.). Advances in Arachnology and Developmental Biology. Papers dedicated to Prof. Dr. Bozidar Curcic. Vienna-Belgrade-Sofia, 517 pp.
  • KARATAS, AY. & M. COLAK. 2005. Scorpions of Gaziantep Province, Turkey (Arachnida: Scorpiones). Euscorpius, 30: 1-7.
  • KARATAS, A., V. FET & M. M. GHARKHELOO. 2012. On the type locality of Mesobuthus vesiculatus (Pocock, 1899) (Scorpiones: Buthidae). Euscorpius, 147: 1-3.
  • KARATAS,A. & M. M. GHARKHELOO.2012. Redescription of Mesobuthus vesiculatus (Pocock, 1899) (Scorpiones: Buthidae) based on specimens from Iran. Turkish Journal of Zoology, 36(5): 576-584.
  • KARATAS, A., M. M. GHARKHELOO & M. UCAK. 2012. Contribution to the distribution of the scorpions of Iran. Zoology in the Middle East, 55: 111-120.
  • KARATAS, A. & A. KARATAS. 2001. First record of Mesobuthus eupeus (C. L. Koch, 1839) from central Anatolia (Scorpiones: Buthidae). Pp. 297-299 in Fet, V. & P. A. Selden (eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK.
  • KARATAS, A. & A. KARATAS. 2003. Mesobuthus eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) in Anatolia. Euscorpius, 7: 1-7.
  • KARATAS, A., M. UCAK, A. KARATAS & O. EROGLU. 2019. Records of chromosomal data of some scorpions (Arachnida: Scorpiones) from Turkey. Hacettepe Journal of Biology and Chemistry, 47: 1-6.
  • KESSLER, K. F. 1874. O russkikh skorpionakh [On Russian scorpions]. Trudy Russkago Entomologicheskago Obshchestva v S.-Peterburge [Transactions of the Russian Entomological Society in St. Petersburg], (1): 3-27 (in Russian) [date on the volume cover, 1876; separatum first available in 1874].
  • KOCH, C. L. 1839a. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung, 5(6): 125-158, figs. 418-419.
  • KOCH, C. L. 1839b. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung, 6(3): 51-53, figs. 466-467.
  • KOCH, L. 1878. Kaukasische Arachnoideen. Pp. 36-71 in Schneider, O. (ed.) Naturwissenschaftliche Beitrage zur Kenntnis der Kaukasuslander,auf Grund seiner Sammelbeute herausgegeben. Dresden: Gesellschaft "Isis", 3.
  • KOVARIK, F. 2002. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket, 8(1): 1-23.
  • KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.
  • KOVARIK, F. 2019. Taxonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae). Euscorpius, 288: 1-27.
  • KOVARIK, F. & P. BERON. 2015. A checklist of the scorpions (Arachnida, Scorpiones) in the collections of the National Museum of Natural History (Sofia). Historia naturalis bulgarica, 22: 37-44.
  • KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.
  • KOVARIK, F., F. STAHLAVSKY, T. KORINKOVA, J. KRAL & T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1-12.
  • KOVARIK, F. & S. WHITMAN. 2005. Cataloghi del Museo di Storia Naturale dell'Universita di Firenze - sezione di zoologia "La Specola" XXII. Arachnida Scorpiones. Tipi. Addenda (1998-2004) e checklist della collezione (Euscorpiinae esclusi). Atti della Societa Toscana di Scienze Naturali, Memorie, serie B, 111 (2004): 103-119.
  • KOVARIK, F., E. A. YAGMUR, V. FET & S. NAVIDPOUR. 2011. On two subspecies of Mesobuthus eupeus (C. L. Koch, 1839) in Turkey (Scorpiones: Buthidae). Euscorpius, 109: 1-15.
  • KRYZHANOVSKY, O. L. 1965. Sostav i proiskhozhdenie nazemnoi fauny Srednei Azii [The Composition and Origin of the Terrestrial Fauna of Middle Asia]. Moscow & Leningrad: Nauka, 419 pp. (in Russian; English translation: New Delhi, 1980).
  • KUMAR, S., G. STECHER, M. LI, C. KNYAZ & K. TAMURA. 2018. MEGA X: molecular evolutionary genetics analysis across computing platforms. Molecular Biology and Evolution, 35(6): 1547.
  • LARSSON, A. 2014. AliView: a fast and lightweight alignment viewer and editor for large datasets. Bioinformatics, 30(22): 3276-3278 (Nov 15). doi: 10.1093/bioinformatics/btu531.
  • LIECHTENSTEIN, H. 1823. Naturhistorische Anhang. Pp. 113-150 in: Eversmann, E. Reise von Orenburg nach Buchara. Berlin: E. H. G. Christiani.
  • LIPSKY, V. I. 1902. Gornaya Bukhara: rezul'taty tryokhletnikh puteshestvii v Srednyuyu Aziyu v 1896, 1897 i 1899 godu. Chast' 1. Gornaya Bukhara: Gissarskaya ekspeditsiya, 1896 g. [The Mountainous Buchara: the results of three years of travels to Central Asia in 1896, 1897 and 1899. Part 1. The Mountainous Buchara: the Gissar Expedition, 1896]. St. Petersburg (in Russian).
  • MAKINO, S. 1956. A review of the chromosome numbers in animals. Tokyo: Hokurykan.
  • MAYR, E. 1969. Principles of Systematic Zoology. New York: McGraw-Hill Book Co., xi + 428 pp.
  • MAYR, E. 1971. Prinstipy zoologicheckoi sistematiki [Principles of Systematic Zoology]. Moscow: Mir, 454 pp. (in Russian).
  • MIRSHAMSI, O. 2012. Ecological niche modeling of two scorpion species Mesobuthus eupeus (C. L. Koch, 1839) and M. phillipsii (Pocock, 1889) from the Iranian Plateau and Zagros region (Arachnida: Scorpiones). Euscorpius, 154: 1-10.
  • MIRSHAMSI, O., A. SARI, E. ELAHI & S. HOSSEINIE. 2010. Phylogenetic relationships of Mesobuthus eupeus (C. L. Koch, 1839) inferred from COI sequences (Scorpiones: Buthidae). Journal of Natural History, 44(47): 2851-2872.
  • MIRSHAMSI, O., A. SARI, E. ELAHI & S. HOSSEINIE. 2011a. Mesobuthus eupeus (Scorpiones: Buthidae) from Iran: A polytypic species complex. Zootaxa, 2929: 1-21.
  • MIRSHAMSI, O., A. SARI & S. HOSSEINIE. 2011b. History of study and checklist of the scorpion fauna (Arachnida: Scorpiones) of Iran. Progress in Biological Sciences, 1(2):16-28.
  • MLIKOVSKY, J. & S. FRAHNERT. 2009. Type specimens and type localities of birds (Aves) collected by Eduard Eversmann and Christian Pander during the Negri Expedition to Bukhara in 1820-1821. Zootaxa, 2297: 15-26.
  • MORADI, M., E. A. YAGMUR, P. M. GHARAKHLOO & F. AHMADI. 2015. Scorpion fauna of Zanjan Province, Iran (Arachnida: Scorpiones). Journal of Applied Biological Sciences, 9(1): 11-14.
  • MORITZ, M. & S.-CH. FISCHER. 1980. Die Typen der Arachniden-Sammlung des Zoologischen Museums Berlin. III. Scorpiones. Mitteilungen aus dem Zoologischen Museum in Berlin, 2(36): 309-326.
  • NAVIDPOUR, S., M. EZATKHAH, F. KOVARIK, M. E. SOLEGLAD & V. FET. 2011. Scorpions of Iran (Arachnida, Scorpiones). Part VII. Kerman Province. Euscorpius, 131: 1-32.
  • NAVIDPOUR, S., V. FET, F. KOVARIK & M. E. SOLEGLAD. 2008a. Scorpions of Iran (Arachnida, Scorpiones). Part III. Ilam Province. Euscorpius, 69: 1-29.
  • NAVIDPOUR, S., V.FET, F. KOVARIK& M. E. SOLEGLAD. 2012. Scorpions of Iran (Arachnida, Scorpiones). Part VIII. Fars Province. Euscorpius, 139: 1-32.
  • NAVIDPOUR, S., F. KOVARIK, M. E. SOLEGLAD & V. FET. 2008b. Scorpions of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province. Euscorpius, 65: 1-41.
  • NAVIDPOUR, S., F. KOVARIK, M. E. SOLEGLAD & V. FET. 2008c. Scorpions of Iran (Arachnida, Scorpiones). Part IV. Kohgilouyeh & Boyer Ahmad Province. Euscorpius, 74: 1-24.
  • NAVIDPOUR, S., F. KOVARIK, M. E. SOLEGLAD & V. FET. 2019. Scorpions of Iran (Arachnida, Scorpiones). Part X. Alborz, Markazi and Tehran Provinces with descriptions of Orthochirus carinatus sp. n. (Buthidae). Euscorpius, 276: 1-20.
  • NAVIDPOUR, S., H. H. NAYEBZADEH, M. E. SOLEGLAD, V. FET, F. KOVARIK & M. H. KAYEDI. 2010. Scorpions of Iran (Arachnida, Scorpiones). Part VI. Lorestan Province. Euscorpius, 99: 1-23.
  • NAVIDPOUR, S., M. E. SOLEGLAD, V. FET & F. KOVARIK. 2013. Scorpions of Iran (Arachnida, Scorpiones). Part IX. Hormozgan Province, with description of Odontobuthus tavighiae sp. n. (Buthidae). Euscorpius, 170: 1-29.
  • NAVIDPOUR, S., M. E. SOLEGLAD, F. KOVARIK & V. FET. 2008d. Scorpions of Iran (Arachnida, Scorpiones). Part II. Bushehr Province. Euscorpius, 67: 1-33.
  • NORDMANN, A. 1840. Notice sur les Scorpions de la faune pontique. In: Voyage dans la Russie meridionale et la Crimee, par la Hongrie, la Valachie et la Moldavie, execute an 1837, sous la direction de M. Anatole de Demidoff, par Mm. de- Sainson, Le-Play, Huot, Leleveille, Raflet, Rousseau, de Nordmann et du Ponceau; dedie a S. M. Nic h olas 1-er, Empereur de toutes les Russies. Paris, 3: 751-752.
  • NOVRUZOV, N. E., F. KOVARIK & V. FET. 2022. Mesobuthus zarudnyi sp. n. from Azerbaijan (Scorpiones: Buthidae). Euscorpius, 347: 1-9.
  • OVCHINNIKOVA, O. G., M. G. VOLKOVICH & V. FET. 2021. Remembering Victor Anatolyevich Krivokhatsky (January 20, 1954 - August 18, 2021). Caucasian Entomological Bulletin, 17(2): 499-509.
  • PALCU, D. V., I. S. PATINA, I. SANDRIC, S. LAZAREV, I. VASILIEV, M. STOICA & W. KRIJGSMAN. 2021. Late Miocene megalake regressions in Eurasia. Scientific Reports, 11: 11471. https://doi.org/10.1038/s41598-021- 91001-z
  • PALLAS, P. S. 1776. Reise durch verschiedene Provinzen des Russischen Reichs. Dritter Theil vom Jahr 1772. und 1773. Reise aus Sibirien zuruck an die Wolga im 1773sten Jahr. Des Dritten Theil. Zweytes Buch. St. Petersburg: Kayserliche Academie der Wissenschaften, S. 455-760 + [1-26].
  • PAVLOVSKY, E. N. 1916a. [Dzhulek, of Perovsky District, Syr-Darya Province, and some biological observations in its environs]. Trudy Imperatorskago Petrogradskago Obshchestva Yestestvoispytatelei [Travaux de la Societe imperiale des Naturalistes de Petrograd], 47(1): 27-68 (in Russian).
  • PAVLOVSKY, E. N. 1916b. Quelques observations biologiques sur les scorpions de la famille des Buthidae. Comptes rendus de la Societe biologique [Petrograd], 81: 243-246.
  • PAVLOVSKY, E. N. 1934. [Some data on the poisonous animals of Turkmenia]. Pp. 191-204 in: Trudy Kara- Kalinskoi i Kzyl-Atrekskoi parazitologicheskoi ekspeditsii 1931 goda i materialy po faune Turkmenii [Transactions of the 1931 Kara-Kala and Kzyl-Atrek Parasitological Expedition, with contributions to the fauna of Turkmenia]. Leningrad: Ed. Acad. Sci. USSR and Narkomzdrav of Turkmenia (in Russian).
  • PENTHER, A. 1912. Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Scorpiones. Annalen des Kaiserlich-Koniglichen Naturhistorischen Hofmuseums in Wien, 26(1/2): 109-115.
  • PEREZ MINNOCCI, S. 1974. Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identificacion de los generos de la region Paleartica Occidental. Madrid: Universidad Complutense de Madrid, Facultad de Ciencias, Departamento de Zoologia, Catedra de Artropodos, 7: 1-45.
  • PIRALI-KHEIRABADI, K., S. NAVIDPOUR, V. FET, F. KOVARIK & M. E. SOLEGLAD. 2009. Scorpions of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province. Euscorpius, 78: 1-23.
  • POCOCK, R. I. 1889a. Arachnida, Chilopoda and Crustacea. On the zoology of the Afghan Commission on Delimitation. Transactions of Linnean Society of London, Zoology, ser. 2, 5(3): 110-122.
  • POCOCK, R. I. 1889b. Notes on some Buthidae, new and old. Annals and Magazine of Natural History, 3: 334- 351.
  • POCOCK, R. I. 1899. Chilopoda and Arachnida. In: R. T. Gunther. Contributions to the natural history of Lake Urmi, N. W. Persia, and its neighbourhood. Journal of the Linnean Society of London, Zoology, 27: 399-406.
  • POCOCK, R. I. 1900. Arachnida. The Fauna of British India, including Ceylon and Burma. Published under the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp.
  • POVERENNYI, N. M. 2015. [The scorpion Mesobut[h] us eupeus (Scorpiones) in various habitats of the Bogdo-Baskunchak Reserve]. Entomologicheskie i parazitologicheskie issledovaniia v Povolzhye [Entomological and Parasitological Research in Povolzhye], 12: 153-155 (in Russian).
  • POVERENNYI, N. M. 2018. [The scorpion Mesobut[h] us eupeus (C. L. Koch, 1839) (Scorpiones) in the Ryn Sands of the Western Kazakhstan]. Entomologicheskie i parazitologicheskie issledovaniia v Povolzhye [Entomological and Parasitological Research in Povolzhye], 15: 109-111 (in Russian).
  • POVERENNYI, N. M. & V. V. ANIKIN. 2017. [Study of phylogenetic relationships of the representatives of the genus Mesobuthus (Arachnida: Scorpiones) based on the molecular DNA analysis]. Ivzestiia Saratovskogo universiteta. Nov. ser. Khimiia. Biologiia. Ekologiia [Proceedings of the Saratov University. New Ser. Chemistry, Biology, Ecology], 17(3): 333-334 (in Russian).
  • POVERENNYI, N. M. & V. V. ANIKIN. 2019. [Phylogenetic relationships of the scorpion Mesobut[h]us eupeus (С. L. Koch, 1839) (Scorpiones) from the Lower Povolzhye and Southern Kazakhstan. Part 1. Molecular genetic similarity of the CO1 gene sequences]. Entomologicheskie i parazitologicheskie issledovaniia v Povolzhye [Entomological and Parasitological Research in Povolzhye], 16: 9-14 (in Russian).
  • POVERENNYI, N. M. & V. V. ANIKIN. 2020a. [Studies of phylogenetic relationships of the scorpion Mesobuthus eupeus volgensis (Birula, 1925) from the Lower Povolzhye with the closely related species from Central Asia based on coxI DNA analysis]. Pp. 98-101 in: Issledovaniia molodykh uchenykh v biologii i ekologii [Studies by Young Researchers in Biology and Ecology]. Saratov: Amirit (in Russian).
  • POVERENNYI, N. M. & V. V. ANIKIN. 2020b. [Phylogenetic relationships of the scorpion Mesobuthus eupeus volgensis (Birula, 1925) from the Lower Povolzhye with the closely relatedspeciesfromCentralAsiabasedoncoxIDNAanalysis]. Entomologicheskie i parazitologicheskie issledovaniia v Povolzhye [Entomological and Parasitological Research in Povolzhye], 17: 12-17 (in Russian).
  • PUILLANDRE, N., S. BROUILLET & G. ACHAZ. 2021. ASAP: assemble species by automatic partitioning. Molecular Ecology Resources, 21: 609-620.
  • RAMBAUT, A., A.J. DRUMMOND, D. XIE, G. BAELE & M. A. SUCHARD. 2018. Posterior summarisation in Bayesian phylogenetics using Tracer 1.7. Systematic Biology. syy032. doi:10.1093/sysbio/syy032
  • ROSELAAR, C. S. & M. ALIABADIAN. 2007. A century of breeding bird assessment by western travellers in Iran, 1876-1977. Appendix 1. Podoces, 2(2): 77-96.
  • SADILEK, D., P. NGUYEN, H. KOC, F. KOVARIK, E. A. YAGMUR & F. STAHLAVSKY. 2015. Molecular cytogenetics of Androctonus scorpions: an oasis of calm in the turbulent karyotype evolution of the diverse family Buthidae. Biological Journal of the Linnean Society, 115: 69-76.
  • SCHNEIDER, M. C., V. F. MATTOS & D. M. CELLA. 2021. The scorpion cytogenetic database. Available at: http:// www.arthropodacytogenetics.bio.br/scorpiondatabase/ index.html. Accessed 1 September 2021.
  • SHARMA, G. P., R. PARSHAD & M. G. JONEJA. 1959. Chromosome mechanism in the males of three species of scorpions (Scorpiones - Buthidae). Research Bulletin of the Panjab University, 10: 197-207.
  • SHEDD, M. L. 1922. The Measure of a Man: The Life of William Ambrose Shedd, Missionary to Persia. New York: George H. Doran.
  • SHI, C. M., Z. S. HUANG, L. WANG, L. J. HE, Y. P. HUA, L. LENG, & D. X. ZHANG. 2007. Geographical distribution of two species of Mesobuthus (Scorpiones, Buthidae) in China: insights from systematic field surveys and predictive models. Journal of Arachnology, 35: 215-226.
  • SHI, C. M., Y. J. JI, L. LIU, L. WANG & D. X. ZHANG. 2013. Impact of climate changes from Middle Miocene onwards on evolutionary diversification in Eurasia: Insights from the mesobuthid scorpions. Molecular Ecology, 22: 1700-1716.
  • SHI, C.-M., H.-B. LIANG, D. ALTANCHIMEG, NONNAIZAB, C. CHULUUNJAV & D.-X. ZHANG. 2015. Climatic niche defines geographical distribution of Mesobuthus eupeus mongolicus (Scorpiones: Buthidae) in Gobi desert. Zoological Systematics, 40(3): 339-348.
  • SHI, C.-M., X.-S. ZHANG, L. LIU, Y.-J. JI & D.-X. ZHANG. 2021. Phylogeography of the desert scorpion Mesobuthus mongolicus illuminates a route out of Central Asia and Junggar Basin as a corridor for faunal exchange between Central and East Asia. bioRxiv preprint, doi: https://doi.org/10.1101/2021.10.31.466646 (posted November 2, 2021): 1-25.
  • SOLEGLAD, M. E. & V. FET. 2003. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1-34.
  • STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.