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Published December 31, 2016 | Version v1
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Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species' gene coalescents and morphology (Hymenoptera, Apidae)

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Williams, Paul H., Huang, Jiaxing, Rasmont, Pierre, An, Jiandong (2016): Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species' gene coalescents and morphology (Hymenoptera, Apidae). Zootaxa 4204 (1): 1-72, DOI: 10.11646/zootaxa.4204.1.1

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https://www.checklistbank.org/dataset/37699
LSID
urn:lsid:zoobank.org:pub:C050058A-774D-49C0-93F9-7A055B51C2A0
LSID
urn:lsid:plazi.org:pub:FF971F2C7C6D330CB007A424A336FFA4
URL
http://publication.plazi.org/id/FF971F2C7C6D330CB007A424A336FFA4

Cites
Publication: 10.11646/zootaxa.3830.1.1 (DOI)
Publication: 10.1111/j.1755-0998.2009.02650.x (DOI)
Publication: 10.1046/j.1365-294X.2003.02063.x (DOI)
Publication: 10.1098/rstb.1998.0206 (DOI)
Publication: 10.1016/S0031-0182(03)00367-5 (DOI)
Publication: 10.1017/S147720000500191X (DOI)
Publication: 10.1111/j.1095-8312.2007.00784.x (DOI)
Publication: 10.1080/10635150151125851 (DOI)
Publication: 10.1371/journal.pone.0029251 (DOI)
Publication: 10.1155/1917/38198 (DOI)
Publication: 10.1007/BF00032629 (DOI)
Publication: 10.1007/s13592-011-0022-z (DOI)
Publication: 10.1080/10635150701701083 (DOI)
Publication: 10.1073/pnas.91.21.9911 (DOI)
Publication: 10.1186/1471-2148-7-214 (DOI)
Publication: 10.1016/j.ympev.2012.03.018 (DOI)
Publication: 10.2307/2412748 (DOI)
Publication: 10.1002/cjs.5550360302 (DOI)
Publication: 10.1002/mmnd.192419240508 (DOI)
Publication: 10.1093/sysbio/syt033 (DOI)
Publication: 10.1016/j.tree.2012.04.012 (DOI)
Publication: 10.1126/science.1234532 (DOI)
Publication: 10.1098/rspb.2002.2218 (DOI)
Publication: 10.1073/pnas.0406166101 (DOI)
Publication: 10.1098/rsbl.2003.0025 (DOI)
Publication: 10.2307/1934716 (DOI)
Publication: 10.1093/molbev/msp274 (DOI)
Publication: 10.1080/10635150801898912 (DOI)
Publication: 10.1111/j.1096-3642.2012.00861.x (DOI)
Publication: 10.1093/bioinformatics/17.8.754 (DOI)
Publication: 10.1038/nature11324 (DOI)
Publication: 10.1093/molbev/msg007 (DOI)
Publication: 10.1016/j.ympev.2003.12.003 (DOI)
Publication: 10.1111/ddi.12188 (DOI)
Publication: 10.1016/j.ode.2006.04.001 (DOI)
Publication: 10.1111/zsc.12107 (DOI)
Publication: 10.1186/1471-2148-13-263 (DOI)
Publication: 10.1080/09670262.2014.904524 (DOI)
Publication: 10.1093/sysbio/syp017 (DOI)
Publication: 10.1080/10635150500354928 (DOI)
Publication: 10.1006/mpev.1998.0606 (DOI)
Publication: 10.1186/1471-2164-9-214 (DOI)
Publication: 10.1098/rstb.2005.1724 (DOI)
Publication: 10.1093/sysbio/syp027 (DOI)
Has part
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Taxonomic treatment: http://treatment.plazi.org/id/03AE67547C78331AB090A6FCA6ADFC61 (URL)
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Taxonomic treatment: 10.5281/zenodo.5625289 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03AE67547C4D332DB090A088A7ADFB04 (URL)
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Figure: 10.5281/zenodo.192303 (DOI)
Dataset: http://table.plazi.org/id/DF7886CA7C6A330BB090A5B4A652FE43 (URL)
Dataset: http://table.plazi.org/id/DF7886CA7C663307B090A5B4A12BFE63 (URL)
Figure: 10.5281/zenodo.192304 (DOI)
Figure: 10.5281/zenodo.192305 (DOI)
Figure: 10.5281/zenodo.192306 (DOI)
Figure: 10.5281/zenodo.192307 (DOI)
Figure: 10.5281/zenodo.192308 (DOI)
Figure: 10.5281/zenodo.192309 (DOI)
Figure: 10.5281/zenodo.192310 (DOI)
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Figure: 10.5281/zenodo.192318 (DOI)
Figure: 10.5281/zenodo.192319 (DOI)
Figure: 10.5281/zenodo.192320 (DOI)
Is source of
Dataset: https://www.gbif.org/dataset/3f8866d2-529e-43ad-b971-29fc52a13858 (URL)

References

  • Aichhorn, A. (1976) Beitrag zur Hummelzucht zur Biologie von Bombus mendax. Berichte aus dem Haus der Natur Salzburg, 7, 13-31.
  • An, J.-D., Huang, J.-X., Shao, Y.-Q., Zhang, S.-W., Wang, B., Liu, X.-Y., Wu, J. & Williams, P.H. (2014) The bumblebees of North China (Apidae, Bombus Latreille). Zootaxa, 3830 (1), 1-89.
  • http://dx.doi.org/10.11646/zootaxa.3830.1.1
  • An, J.-D., Huang, J.-X., Williams, P.H., Wu, J. & Zhou, B.-F. (2010) Species diversity and colony characteristics of bumblebees in the Hebei region of North China. Chinese Journal of Applied Ecology, 21, 1542-1550.
  • An, J.-D., Williams, P.H., Miao, Z.-Y. & Zhou, B.-F. (2011) The bumblebees of Gansu, northwest China (Hymenoptera, Apidae). Zootaxa, 2865, 1-36.
  • Anonymous (2001) The Times comprehensive atlas of the world (10th ed.). London, Times Books.
  • Baker, A.J., Tavares, E.S. & Elbourne, R.F. (2009) Countering criticisms of single mitochondrial DNA gene barcoding in birds. Molecular Ecology Resources, 9 (supplement 1), 257-268.
  • http://dx.doi.org/10.1111/j.1755-0998.2009.02650.x
  • Ballard, J.W.O. & Whitlock, M.C. (2004) The incomplete natural history of mitochondria. Molecular Ecology, 13, 729-744. http://dx.doi.org/10.1046/j.1365-294X.2003.02063.x
  • Barraclough, T.G., Vogler, A.P. & Harvey, P.H. (1998) Revealing the factors that promote speciation. Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences, 353, 241-249. http://dx.doi.org/10.1098/rstb.1998.0206
  • Baum, D. & Smith, S. (2012) Tree thinking: an introduction to phylogenetic biology, Roberts and Company.
  • Bertsch, A. (2010) A phylogenetic framework for the bumblebee species of the subgenus Bombus sensu stricto based on mitochondrial DNA markers, with a short description of the neglected taxon B. minshanicola Bischoff, 1936 n. status. Beitrage zur Entomologie, 60, 471-487.
  • Bischoff, H. (1930) Entomologische Ergebnisse der schwedischen Kamtschatka Expedition 1920-1922. 29. Bombinae (Hymen.). Arkiv for zoologi, 21, 1-6.
  • Bischoff, H. (1931) Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II). 4. Hymenoptera VII. Zur Kenntnis einiger Hummelnester aus dem Pamir. Mitteilungen aus dem Zoologische Museum in Berlin, 16, 860- 864.
  • Bischoff, H. (1936) Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927-1930. 56. Hymenoptera. 10. Bombinae. Arkiv for zoologi, 27A, 1-27.
  • Boehme, M. (2003) The Miocene Climatic Optimum: evidence from ectothermic vertebrates of Central Europe. Palaeogeography Palaeoclimatology Palaeoecology, 195, 389-401.
  • http://dx.doi.org/10.1016/S0031-0182(03)00367-5
  • Bolton, B. (2007) How to conduct large-scale taxonomic revisions in Formicidae. In: R. R. Snelling, B. L. Fisher & P. S. Ward (Eds), Advances in ant systematics (Hymenoptera: Formicidae): homage to E. O. Wilson - 50 years of contributions. Memoirs of the American Entomological Institute, pp. 52-71.
  • Bretschneider, E. (1981) History of European botanical discoveries in China (Vol. 2). Leipzig, Zentral-Antiquariat der DDR.
  • Brower, A.V.Z. (2006) Problems with DNA barcodes for species delimitation: 'ten species' of Astraptes fulgerator reassessed (Lepidoptera: Hesperiidae). Systematics and Biodiversity, 4, 127-132.
  • http://dx.doi.org/10.1017/S147720000500191X
  • Burger, F., Creutzburg, F., Hartmann, J. & Hartmann, M. (2009) Die Hummeln von Nepal (Insecta: Hymenoptera: Apidae: Bombus). In: M. Hartmann & J. Weipert (Eds), Biodiversitat und Naturausstattung im Himalaya III. Verein der Freunde und Forderer des Naturkundemuseums Erfurt e.V., Erfurt, pp. 455-462.
  • Cameron, S.A., Derr, J.N., Austin, A.D., Wooley, J.B. & Wharton, R.A. (1992) The application of nucleotide sequence data to phylogeny of the Hymenoptera: a review. Journal of Hymenoptera Research, 1, 63-79.
  • Cameron, S.A., Hines, H.M. & Williams, P.H. (2007) A comprehensive phylogeny of the bumble bees (Bombus). Biological Journal of the Linnean Society, 91, 161-188.
  • http://dx.doi.org/10.1111/j.1095-8312.2007.00784.x
  • Cameron, S.A. & Mardulyn, P. (2001) Multiple molecular data sets suggest independent origins of highly eusocial behavior in bees (Hymenoptera: Apinae). Systematic Biology, 20, 194-214.
  • http://dx.doi.org/10.1080/10635150151125851
  • Carolan, J.C., Murray, T.E., Fitzpatrick, U., Crossley, J., Schmidt, H., Cederberg, B., McNally, L., Paxton, R.J., Williams, P.H. & Brown, M.J.F (2012) Colour patterns do not diagnose species: quantitative evaluation of a DNA barcoded cryptic bumblebee complex. PLoS ONE, 7, 662-667.
  • http://dx.doi.org/10.1371/journal.pone.0029251
  • Cederberg, B., Michez, D., Nieto, A., Radchenko, V., Rasmont, P. & Roberts, S.P.M. (2013) Bombus mendax. In: IUCN Red List of Threatened Species. IUCN. Avaliable from: http://www.iucnredlist.org (Accessed 1 Dec. 2016)
  • Cockerell, T.D.A. (1910) Some bees from high altitudes in the Himalaya mountains. Entomologist, 43, 238-239.
  • Cockerell, T.D.A. (1911) Bees from the Himalaya mountains. The Entomologist, 44, 176-177.
  • Cockerell, T.D.A. (1917) New social bees. Psyche, 24, 120-128.
  • http://dx.doi.org/10.1155/1917/38198
  • Cogbill, C.V. & White, P.S. (1991) The latitide-elevation relationship for spruce-fir forest and treeline along the Appalachian mountain chain. Vegetatio, 94, 153-175.
  • http://dx.doi.org/10.1007/BF00032629
  • de Boer, A.J. (2002) The types of Carabidae (Coleoptera) in the Zoologisch Museum Amsterdam, predominantly the collection "Oskar Vogt". Leiden, Backhuys Publishers.
  • de Boer, A.J. (2004) Oskar Vogt en zijn entomologische collecties, of: het verband tussen hommels en hersenonderzoek. Entomologische Berichten, 64, 62-68.
  • De Meulemeester, T. (2012) Approche integrative dans la systematique de taxons complexes: bourdons et abeilles fossiles. In: Laboratoire de Zoologie. Universite de Mons, Mons, pp. 316-316.
  • De Meulemeester, T., Aytekin, A.M., Cameron, S. & Rasmont, P. (2010) Nest architecture and species status of the bumble bee Bombus (Mendacibombus) shaposhnikovi (Hymenoptera: Apidae: Bombini). Apidologie, 42, 301-306. http://dx.doi.org/10.1007/s13592-011-0022-z
  • de Queiroz, K. (2007) Species concepts and species delimitation. Systematic Biology, 56, 879-886. http://dx.doi.org/10.1080/10635150701701083
  • Dowton, M. & Austin, A.D. (1994) Molecular phylogeny of the insect order Hymenoptera: apocritan relationships. Proceedings of the National Academy of Sciences of the United States of America, 91, 9911-9915. http://dx.doi.org/10.1073/pnas.91.21.9911
  • Drummond, A.J. & Rambaut, A. (2007) BEAST: Bayesian evolutionary analysis by sampling trees. BioMed Central Evolutionary Biology, 7, 214-214.
  • http://dx.doi.org/10.1186/1471-2148-7-214
  • Duennes, M.A., Lozier, J.D., Hines, H.M. & Cameron, S.A. (2012) Geographical patterns of genetic divergence in the widespread Mesoamerican bumble bee Bombus ephippiatus (Hymenoptera: Apidae). Molecular Phylogenetics and Evolution, 64, 219-231.
  • http://dx.doi.org/10.1016/j.ympev.2012.03.018
  • Ebmer, A.W. (1996) Asiatische Halictidae, 5. Daten zur Aculeata-Fauna der Ussuri-Region unter Berucksichtigung der angrenzenden Gebiete (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer biologische Beitrage, 28, 261- 304.
  • Ebmer, A.W. (2008) Die westpalaarktischen Arten der Gattung Dufourea LEPELETIER 1841 (Insecta: Hymenoptera: Apoidea: Halictidae: Rophitinae) Funfter Nachtrag. Linzer biologische Beitrage, 40, 581-625.
  • Eversmann, E. (1846) Hymenopterorum Rossicorum species novae vel parum cognitae, descriptae et ex parte depictae. Byulleten' Moskovskogo obshchestva ispytatelei prirody, 19, 436-443.
  • Farris, J.S. (1976) Expected asymmetry of phylogenetic trees. Systematic Zoology, 25, 196-198. http://dx.doi.org/10.2307/2412748
  • Ferreira, M.A.R. & Suchard, M.A. (2008) Bayesian analysis of elapsed times in continuous-time Markov chains. The Canadian Journal of Statistics, 36, 355-368.
  • http://dx.doi.org/10.1002/cjs.5550360302
  • Franklin, H.J. (1954) The evolution and distribution of American bumblebee kinds. Transactions of the American Entomological Society, 80, 43-51.
  • Friese, H. (1911b) Neue Varietaten von Bombus, III. Deutsche Entomologische Zeitschrift, 1911, 571-572.
  • Friese, H. (1913) Uber einige neue Apiden (Hym.). Archiv fur Naturgeschichte, 78, 85-89.
  • Friese, H. (1924) Uber auffallende Hummelformen. (Hym. Apid.). Deutsche Entomologische Zeitschrift, 1924, 437-439. http://dx.doi.org/10.1002/mmnd.192419240508
  • Frison, T.H. (1927) A contribution to our knowledge of the relationships of the Bremidae of America north of Mexico (Hymenoptera). Transactions of the American Entomological Society, 53, 51-78.
  • Fujisawa, T. & Barraclough, T.G. (2013) Delimiting species using single-locus data and the Generalized Mixed Yule Coalescent approach: a revised method and evaluation on simulated data sets. Systematic Biology, 62, 707-724. http://dx.doi.org/10.1093/sysbio/syt033
  • Fujita, M.K., Leache, A.D., Burbrink, F.T., McGuire, J.A. & Moritz, C. (2012) Coalescent-based species delimitation in an integrative taxonomy. Trends in Ecology and Evolution, 27, 480-488.
  • http://dx.doi.org/10.1016/j.tree.2012.04.012
  • Gardner, A.S., Moholdt, G., Cogley, J.G., Wouters, B., Arendt, A.A., Wahr, J., Berthier, E., Hock, R., Pfeffer, W.T., Kaser, G., Ligtenberg, S.R.M., Bolch, T., Sharp, M.J., Hagen, J.O., van den Broeke, M.R. & Paul, F. (2013) A reconciled estimate of glacier contributions to sea level rise: 2003 to 2009. Science, 340, 852-857.
  • http://dx.doi.org/10.1126/science.1234532
  • Gerstaecker, A. (1869) Beitrage zur naheren Kenntniss einiger Bienen-Gattungen. Stettiner entomologische Zeitung, 30, 315- 367.
  • Haas, A. (1976) Paarungsverhalten und Nestbau der alpinen Hummelart Bombus mendax (Hymenoptera: Apidae). Entomologica Germanica, 3, 248-259.
  • Hagen, E.v. & Aichhorn, A. (2003) Hummeln: Bestimmen, Ansiedeln, Vermehren, Schutzen (2 ed.). Nottuln, Fauna-Verlag.
  • Hebert, P.D.N., Cywinska, A., Ball, S.L. & DeWaard, J.R. (2003) Biological identifications through DNA barcodes. Proceedings of the Royal Society of London B, Biological Sciences, 270, 313-321. http://dx.doi.org/10.1098/rspb.2002.2218
  • Hebert, P.D.N., Penton, E.H., Burns, J.M., Janzen, D.H. & Hallwachs, W. (2004) Ten species in one: DNA barcoding reveals cryptic species in the neotropical skipper butterfly Astraptes fulgerator. Proceedings of the National Academy of Sciences, 101, 14812-14817.
  • http://dx.doi.org/10.1073/pnas.0406166101
  • Hebert, P.D.N., Ratnasingham, S. & deWaard, J.R. (2003) Barcoding animal life: cytochrome c oxidase subunit 1 divergences among closely related species. Proceedings of the Royal Society of London (B), 270, S96-S99. http://dx.doi.org/10.1098/rsbl.2003.0025
  • Heck, K.L., Belle, G.v. & Simberloff, D. (1975) Explicit calculation of the rarefaction diversity measurement and the determination of sufficient sample size. Ecology, 56, 1459-1461.
  • http://dx.doi.org/10.2307/1934716
  • Heinrich, B. (1979) Bumblebee economics. Cambridge (Massachusetts), Harvard University Press.
  • Heled, J. & Drummond, A.J. (2010) Bayesian inference of species trees from multilocus data. Molecular Biology and Evolution, 27, 570-580.
  • http://dx.doi.org/10.1093/molbev/msp274
  • Hines, H.M. (2008) Historical biogeography, divergence times, and diversification patterns of bumble bees (Hymenoptera: Apidae: Bombus). Systematic Biology, 57, 58-75.
  • http://dx.doi.org/10.1080/10635150801898912
  • Hines, H.M. & Williams, P.H. (2012) Mimetic colour pattern evolution in the highly polymorphic Bombus trifasciatus (Hymenoptera: Apidae) species complex and its comimics. Zoological Journal of the Linnean Society, 166, 805-826. http://dx.doi.org/10.1111/j.1096-3642.2012.00861.x
  • Hopkirk, P. (1982) Tresspassers on the roof of the world: the race for Lhasa. London, John Murray.
  • Huelsenbeck, J.P. & Ronquist, F. (2001) MRBAYES: Bayesian inference of phylogeny. Bioinformatics, 17, 754-755. http://dx.doi.org/10.1093/bioinformatics/17.8.754
  • ICZN (1999) International code of zoological nomenclature (4 ed.). London, International Commission on Zoological Nomenclature.
  • Ito, M. (1985) Supraspecific classification of bumblebees based on the characters of male genitalia. Contributions from the Institute of Low Temperature Science, Hokkaido University (B), 20, 143-143.
  • Ito, M. & Kuranishi, R.B. (2000) Bumble bees (Hymenoptera: Apidae) occurring in the Kamchatka Peninsula and the North Kuril Islands. Natural History Research, 7, 281-289.
  • IUCN (2001) IUCN Red List categories and criteria: version 3.1. Gland and Cambridge, IUCN Species Survival Commission.
  • Kaab, A., Berthier, E., Nuth, C., Gardelle, J. & Arnaud, Y. (2012) Contrasting patterns of early twenty-first-century glacier mass change in the Himalayas. Nature, 488, 495-498.
  • http://dx.doi.org/10.1038/nature11324
  • Kawakita, A., Sota, T., Ascher, J., Ito, M., Tanaka, H. & Kato, M. (2003) Evolution and phylogenetic utility of alignment gaps within intron sequences of three nuclear genes in bumble bees (Bombus). Molecular Biology and Evolution, 20, 87-92. http://dx.doi.org/10.1093/molbev/msg007
  • Kawakita, A., Sota, T., Ito, M., Ascher, J.S., Tanaka, H., Kato, M., et al. (2004) Phylogeny, historical biogeography, and character evolution in bumble bees (Bombus: Apidae) based on simultaneous analysis of three nuclear gene sequences. Molecular Phylogenetics and Evolution, 31, 799-804.
  • http://dx.doi.org/10.1016/j.ympev.2003.12.003
  • Kelly, S., Grenyer, R. & Scotland, R.W. (2014) Phylogenetic trees do not reliably predict feature diversity. Diversity and Distributions, 2014, 1-13.
  • http://dx.doi.org/10.1111/ddi.12188
  • Kerzhner, I.M. (1972) Historical survey of studies of the insect fauna of the Mongolian People's Republic. Nasekomye Mongolii, Leningrad, 1, 57-112.
  • Kitching, I.J., Forey, P., Humphries, C.J. & Williams, D. (1998) Cladistics: the theory and practice of parsimony analysis (2 ed.). Oxford, Oxford University Press.
  • Komarov, V.L. (1928) Les itineraires botaniques des principales expeditions russes en Asie Centrale. Itineraires de G. N. Potanine (1876-1899) Trudy Imperatorskago S.-Peterburgskago botanicheskago sada, 34, 201-404.
  • Kriechbaumer, J. (1870) Vier neue Hummelarten. Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 20, 157- 160.
  • Kruger, E. (1917) Zur Systematik der mitteleuropaischen Hummeln (Hym.). Entomologische Mitteilungen, 6, 55-66.
  • Kruseman, G. (1958) Notes sur les bourdons pyreneens du genre Bombus dans les collections neerlandaises. Beaufortia, 6, 161-170.
  • Kuhlmann, M., Else, G.R., Dawson, A. & Quicke, D.L.J. (2007) Molecular, biogeographical and phenological evidence for the existence of three western European sibling species in the Colletes succinctus group (Hymenoptera: Apidae). Organic Diversity and Evolution, 7, 155-165.
  • http://dx.doi.org/10.1016/j.ode.2006.04.001
  • Lecocq, T., Dellicour, S., Michez, D., Dehon, M., Dewulf, A., De Meulemeester, T., et al. (2015) Methods for species delimitation in bumblebees (Hymenoptera, Apidae, Bombus): towards an integrative approach. Zoologica Scripta, 44, 281-297.
  • http://dx.doi.org/10.1111/zsc.12107
  • Lecocq, T., Dellicour, S., Michez, D., Lhomme, P., Vanderplanck, M., Valterova, I., et al. (2013) Scent of a break-up: phylogeography and reproductive trait divergences in the red-tailed bumblebee (Bombus lapidarius). BMC Evolutionary Biology, 13, 263-263.
  • http://dx.doi.org/10.1186/1471-2148-13-263
  • Leliaert, F., Verbruggen, H., Vanormelingen, P., Steen, F., Lopez-Bautista, J.M., Zuccarello, G.C., et al. (2014) DNA-based species delimitation in algae. European Journal of Phycology, 49, 179-196.
  • http://dx.doi.org/10.1080/09670262.2014.904524
  • Lemmon, A.R., Brown, J.R., Stanger-Hall, K. & Lemmon, E.M. (2009) The effect of ambiguous data on phylogenetic estimates obtained by maximum likelihood and Bayesian inference. Systematic Biology, 58, 130-145. http://dx.doi.org/10.1093/sysbio/syp017
  • Lepeletier de Saint-Fargeau, A.L.M. (1832) Observations sur l'ouvrage intitule: 'Bombi scandinaviae monographice tractato, etc., a Gustav. Dahlbom.'. Annales de la Societe Entomologique de France, 1, 366-382.
  • Maddison, W.P. & Knowles, L.L. (2006) Inferring phylogeny despite incomplete lineage sorting. Systematic Biology, 55, 21- 30.
  • http://dx.doi.org/10.1080/10635150500354928
  • Mardulyn, P. & Cameron, S.A. (1999) The major opsin in bees (Insecta: Hymenoptera): a promising nuclear gene for higher level phylogenetics. Molecular Phylogenetics and Evolution, 12, 168-176.
  • http://dx.doi.org/10.1006/mpev.1998.0606
  • Meusnier, I., Singer, G.A.C., Landry, J.-F., Hickey, D.A., Hebert, P.D.N. & Hajibabaei, M. (2008) A universal DNA minibarcode for biodiversity analysis. BioMed Central Genomics, 9, 214-214. [214 pp.] http://dx.doi.org/10.1186/1471-2164-9-214
  • Michener, C.D. (2000) The bees of the world (1 ed.). Baltimore, John Hopkins University Press.
  • Monaghan, M.T., Balke, M., Gregory, T.R. & Vogler, A.P. (2005) DNA-based species delineation in tropical beetles using mitochondrial and nuclear markers. Philosophical Transactions of the Royal Society (B), 360, 1925-1933. http://dx.doi.org/10.1098/rstb.2005.1724
  • Monaghan, M.T., Wild, R., Elliot, M., Fujisawa, T., Balke, M., Inward, D.J.G., Lees, D.C., Ranaivosolo, R., Eggleton, P., Barraclough, T.G. & Vogler, A.P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298-311.
  • http://dx.doi.org/10.1093/sysbio/syp027
  • Morawitz, F.F. (1875) Bees. (Mellifera). In: Reise in Turkestan, A. P. Fedtschenko. II Zoologischer Teil. Izvestiya Imp. Obshch. Ljubit. Estestvozn., Anthrop. Ethnogr., t. 19, no. 2, Moscow, p. ii+160.
  • Morawitz, F.F. (1880) Ein Beitrag zur Bienen-Fauna mittel-Asiens. Izvestiya Imperatorskoi akademii nauk, 26, 337-379.
  • Morawitz, F.F. (1881) Die russischen Bombus -Arten in der Sammlung der Kaiserlichen Akademie der Wissenschaften. Izvestiya Imperatorskoi akademii nauk, 27, 213-265.
  • Morawitz, F.F. (1887) Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva, 20 (1886), 195-229.
  • Morawitz, F.F. (1890) Insecta a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIV. Hymenoptera Aculeata. II). III. Apidae. Trudy Russkago entomologicheskago obshchestva, 24, 349-385.
  • Newell, J. (2004) The Russian Far East, a reference guide for conservation and development (2 ed.). McKinleyville, California, Daniel & Daniel.
  • Nurk, N.M., Uribe-Convers, S., Gehrke, B., Tank, D.C. & Blattner, F.R. (2015) Oligocene niche shift, Miocene diversification - cold tolerance and accelerated speciation rates in the St John's Worts (Hypericum, Hypericaceae). BMC Evolutionary Biology, 15, 1-13.
  • Ornosa, C. & Torres, F. (2010) Mendacibombus (Mendacibombus) mendax (Gerstaecker, 1869). In: Atlas de Invertebrados Amenazados de Espana, pp. 160-164.
  • Owen, R.E. & Plowright, R.C. (1980) Abdominal pile color dimorphism in the bumble bee, Bombus melanopygus. Journal of Heredity, 71, 241-247.
  • Owen, R.E., Whidden, T.L. & Plowright, R.C. (2010) Genetic and morphometric evidence for the conspecific status of the bumble bees, Bombus melanopygus and Bombus edwardsii. Journal of Insect Science, 10, 1-18.
  • Ozbek, H. (1997) Bumblebees fauna of Turkey with distribution maps (Hymenoptera: Apidae: Bombinae) Part 1: Alpigenobombus Skorikov, Bombias Robertson and Bombus Latreille. Turkiye entomoloji dergisi, 21, 37-56.
  • Panfilov, D.V. (1957) [On the geographical distribution of bumblebees (Bombus) in China]. Acta geographica sinica, 23, 221- 239.
  • Panfilov, D.V. (1962) [On the biocoenotic structure and geographical distribution of the Issyk Kul Lake area]. In: Y. A. Isakov (Ed), [The study of geography of natural resources of animal and plant world (to the 60th anniversary of A. N. FORMOZOV's birthday)]. Academy of Sciences of the USSR, Moscow, pp. 162-198.
  • Papadopoulou, A., Bergsten, J., Fujisawa, T., Monaghan, M.T., Barraclough, T.G. & Vogler, A.P. (2008) Speciation and DNA barcodes: testing the effects of dispersal on the formation of discrete sequence clusters. Philosophical Transactions of the Royal Society (B), 363, 2987-2996.
  • Peters, G. & Panfilov, D.V. (1968) Hummeln (Bombus) und Schmarotzerhummeln (Psithyrus) Ergebnesse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. Reisbachia, 11, 117-182.
  • Phillips, J.D., Gwiazdowski, R.A., Ashlock, D. & Hanner, R. (2015) An exploration of sufficient sampling effort to describe intraspecific DNA barcode haplotype diversity: examples from the ray-finned fishes (Chordata: Actinopterygii). DNA Barcodes, 3, 66-73.
  • Pittioni, B. (1937a) Die Hummelfauna des Kalsbachtales in Ost-Tirol. Ein Beitrag zur Okologie und Systematik der Hummeln Mitteleuropas. Festschrift zum 60. Geburtstag von Professor Dr. Embrik Strand, 3, 64-122.
  • Pittioni, B. (1937b) Eine Hummelausbeute aus dem Elburs-Gebirge (Iran). Ein Beitrag zur Kenntnis der palaarktischen Hummeln und Schmarotzer-hummeln. Konowia, 16, 113-129.
  • Pittioni, B. (1939) Die Hummeln und Schmarotzerhummeln der Balkan-Halbinsel. II. Spezieller Teil. Izvestiya na Tsarskite prirodonauchni instituti v Sofiya, 12, 49-115.
  • Ploquin, E.F., Herrera, J.M. & Obeso, J.R. (2013) Bumblebee community homogenization after uphill shifts in montane areas of northern Spain. Oecologia, 173, 1649-1660.
  • Plowright, R.C. & Pallett, M.J. (1979) Worker-male conflict and inbreeding in bumble bees (Hymenoptera: Apidae). Canadian Entomologist, 111, 289-294.
  • Pons, J., Barraclough, T.G., Gomez-Zurita, J., Cardoso, A., Duran, D.P., Hazell, S., et al. (2006) Sequence-based species delimitation for the DNA taxonomy of undescribed insects. Systematic Biology, 55, 595-609.
  • Proshchalykin, M.Y. & Kupianskaya, A.N. (2005) The bees (Hymenoptera, Apoidea) of the northern part of the Russian Far East. Far Eastern Entomologist, 153, 1-39.
  • Proshchalykin, M.Y. & Kupianskaya, A.N. (2009) The bees of family Apidae (Hymenoptera, Apoidea) of Transbaikalia. Euroasian Entomological Journal, 8, 59-68.
  • Pyke, G.H., Inouye, D.W. & Thomson, J.D. (2012) Local geographic distributions of bumble bees near Crested Butte, Colorado: competition and community structure revisited. Environmental Entomology, 41, 1332-1349.
  • Qiu, J. (2008) The third pole. Nature, 454, 393-396.
  • Rasmont, P. (1983) Catalogue commente des bourdons de la region ouest-palearctique (Hymenoptera, Apoidea, Apidae). Notes Fauniques de Gembloux, 7, 1-71.
  • Rasmont, P. (1988) Monographie ecologique et zoogeographique des bourdons de France et de Belgique (Hymenoptera, Apidae, Bombinae). In: Faculte des Sciences agronomiques de l'Etat, Gembloux, p. 309+lxi.
  • Rasmont, P. & Flagothier, D. (1996) Biogeographie et choix floraux des bourdons (Hymenoptera, Apidae) de la Turquie. Rapport preliminaire 1995-1996. In: NATO-OTAN TU-Pollination Project. Universite de Mons-Hainaut, Mons, Belgium.
  • Rasmont, P., Franzen, M., Lecocq, T., Harpke, A., Roberts, S.P.M., Biesmeijer, J.C., et al. (2015) Climatic risk and distribution atlas of European bumblebees, Pensoft.
  • Rasmont, P. & Iserbyt, S. (2012) Atlas of the European bees: genus Bombus In: STEP Project, Atlas Hymenoptera, Mons, Gembloux.
  • Rasmont, P., Terzo, M., Aytekin, A.M., Hines, H.M., Urbanova, K., Cahlikova, L., et al. (2005) Cephalic secretions of the bumblebee subgenus Sibiricobombus Vogt suggest Bombus niveatus Kriechbaumer and Bombus vorticosus Gerstaecker are conspecific (Hymenoptera, Apidae, Bombus). Apidologie, 36, 571-584.
  • Ratnasingham, S. & Hebert, P.D.N. (2007) BOLD: The Barcode Of Life Data system (www.barcodinglife.org). Molecular Ecology Notes, 2007, 1-10.
  • Reid, N.M. & Carstens, B.C. (2012) Phylogenetic estimation error can decrease the accuracy of species delimitation: a Bayesian implementation of the general mixed Yule-coalescent model. BMC Evolutionary Biology, 12, 196-196.
  • Reinig, W.F. (1930) Untersuchungen zur Kenntnis der Hummelfauna des Pamir-Hochlandes. Zoologische Ergebnisse der deutsch-russischen Alai-Pamir-Expedition der Notgemeinschaft der Deutschen Wissenschaft und der Akademie der Wissenschaften der U.d.S.S.R. Zeitschrift fur Morphologie und Okologie der Tiere, 17, 68-123.
  • Reinig, W.F. (1932a) Beitrage zur Faunistik des Pamir-Gebietes. In: Wissenschaftliche Ergebnisse der Alai-Pamir Expedition 1928. Reimer and Vohsen, Berlin.
  • Reinig, W.F. (1932b) Beitrage zur Faunistik des Pamir-Gebietes. In: Wissenschaftliche Ergebnisse der Alai-Pamir Expedition 1928. Reimer and Vohsen, Berlin.
  • Reinig, W.F. (1934) Entomologische Ergebnisse der deutsch-russischen Alai-Pamir-Expedition, 1928 (III). 7. Hymenoptera VIII (Gen. Bombus Fabr.). Nachtrag. Deutsche Entomologische Zeitschrift, 1933, 163-174.
  • Reinig, W.F. (1940) Beitrage zur Kenntnis der Hummelfauna von Afghanistan. (Hym., Apid.). (Ergebnisse der Reise von H. und E. Kotzsch in den Hindukusch im Jahre 1936.). Deutsche Entomologische Zeitschrift, 1940, 224-235.
  • Richards, O.W. (1930) The humble-bees captured on the expeditions to Mt. Everest (Hymenoptera, Bombidae). Annals and Magazine of Natural History (10), 5, 633-658.
  • Richards, O.W. (1968) The subgeneric divisions of the genus Bombus Latreille (Hymenoptera: Apidae). Bulletin of the British Museum (Natural History) (Entomology), 22, 209-276.
  • Romiguier, J., Cameron, S.A., Woodward, S.H., Fischman, B.J., Keller, L. & Praz, C.J. (2015) Phylogenomics controlling for base compositional bias reveals a single origin of eusociality in corbiculate bees. Molecular Biology and Evolution, December, 1-9.
  • Sabir, A.M., Suhail, A., Ahmed, S. & Khalid, S. (2011) Diversity of bumblebees (Bombini, Apidae: Hymenoptera) in northern Pakistan. International Journal of Agriculture & Biology, 13, 159-166.
  • Sakagami, S.F. (1988) Bionomics of the halictine bees in Northern Japan. IV. Lasioglossum (Evylaeus) nupricola sp. nov., a climatic relic. Kontyu, 56, 337-353.
  • Sandhouse, G.A. (1943) The type species of the genera and subgenera of bees. Proceedings of the United States National Museum, 92, 519-619.
  • Schlick-Steiner, B.C., Steiner, F.M., Seifert, B., Stauffer, C., Christian, E. & Crozier, R.H. (2010) Integrative taxonomy: a multisource approach to exploring biodiversity. Annual Review of Entomology, 55, 421-438.
  • Simberloff, D.E., Hecht, K.L., McCoy, E.D. & Connor, E.F. (1981) There have been no statistical tests of cladistic biogeographical hypotheses. In: G. Nelson & D. E. Rosen (Eds), Vicariance biogeography: a critique. Columbia University Press, New York.
  • Skorikov, A.S. (1910a) [A contribution to the fauna and geographical distribution of bumblebees (Bombus) in northern Caucasus (a preliminary report)]. Trudy Obshch. Ispyt. Prirody Imp. Kharkov. Univ. (Kharkov), 43 (1909), 77-84.
  • Skorikov, A.S. (1910b) [Intraspecific forms of Bombus mendax Gerst.] (Hymenoptera, Bombidae). Russkoe entomologicheskoe Obozrenie, 9 (1909), 328-330.
  • Skorikov, A.S. (1912) Neue Hummelformen (Hymenoptera, Bombidae). IV. Russkoe entomologicheskoe Obozrenie, 12, 606- 610.
  • Skorikov, A.S. (1914) Les formes nouvelles des bourdons (Hymenoptera, Bombidae). VI. Russkoe entomologicheskoe Obozrenie, 14, 119-129.
  • Skorikov, A.S. (1923) [Palaearctic bumblebees. Part I. General biology (including zoogeography)]. Izvestiya Severnoi oblastnoi stantsii zashchity rastenii ot vreditelei, 4 (1922), 1-160.
  • Skorikov, A.S. (1931) Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: V. A. Lindholm (Ed.), Abhandlungen der Pamir-Expedition 1928. Academy of Sciences of the USSR, Leningrad, pp. 175-247.
  • Skorikov, A.S. (1933) Zur Fauna und Zoogeographie der Hummeln des Himalaya. Doklady Akademii nauk SSSR, 1933, 243- 248.
  • Steinbauer, M.J., Field, R., Grytnes, J.-A., Trigas, P., Ah-Peng, C., Attorre, F., et al. (2016) Topography-driven isolation, speciation and a global increase of endemism with elevation. Global Ecology and Biogeography.
  • Strange, J.P., Knoblett, J. & Griswold, T. (2009) DNA amplification from pin-mounted bumble bees (Bombus) in a museum collection: effects of fragment size and specimen age on successful PCR. Apidologie, 40, 134-139.
  • Streinzer, M. & Spaethe, J. (2014) Functional morphology of the visual system and mating strategies in bumblebees (Hymenoptera, Apidae, Bombus). Zoological Journal of the Linnean Society, 170, 735-747.
  • Suhail, A., Sabir, A.M., Asghar, M., Rafi, M.A. & Qadir, A. (2009) Geographic distributional patterns of the genus Bombus (Bombini, Apidae: Hymenoptera) in northern Pakistan. Biological Diversity and Conservation, 2, 1-9.
  • Talavera, G., Dinca, V. & Vila, R. (2013) Factors affecting species delimitations with the GMYC model: insights from a butterfly survey. Methods in Ecology and Evolution, 4, 1101-1110.
  • Thornhill, R. & Alcock, J. (1983) The evolution of insect mating systems. London, Harvard University Press.
  • Tkalcu, B. (1961) Zur Hummelfauna der Umgebung Kuku-Nors (Hymenoptera, Bombinae). Casopis Ceskoslovenske spolecnosti entomologicke, 58, 344-379.
  • Tkalcu, B. (1968) Neue Arten der Unterfamilie Bombinae der palaarktischen Region (Hymenoptera, Apoidea). Sbornik Entomologickeho oddeleni Narodniho musea v Praze, 65, 21-51.
  • Tkalcu, B. (1969) Beitrage zur Kenntnis der Fauna Afghanistans (Sammelergebnisse von O. Jakes 1963-64, D. Povolny 1965, D. Povolny & Fr. Tenora 1966, J. Simek 1965-66, D. Povolny, J. Geisler, Z. Sebek & Fr. Tenora 1967). Bombinae, Apoidea, Hym. Casopis Moravskeho musea v Brne, 53 (1968), 189-210.
  • Tkalcu, B. (1975) Beitrag zur Kenntnis der Hummelfauna der franzosischen Basses-Alpes (Hymenoptera, Apoidea, Bombinae). Sbornik Slovenskeho Narodneho Muzea, 20 (1974), 167-186.
  • Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28-84.
  • Vogt, O. (1911) Studien uber das Artproblem. 2. Mitteilung. Uber das Variieren der Hummeln. 2. Teil. (Schluss). Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1911, 31-74.
  • Wang, C.-Z., Guo, H.-D., Zhang, L., Qiu, Y.-B., Sun, Z.-C., Liao, J.-J., et al. (2013) Improved alpine grassland mapping in the Tibetan Plateau with MODIS time series: a phenology perspective. International Journal of Digital Earth.
  • Wang, S.-F. (1982) Hymenoptera: Apidae - Bombus. In: Q.-T. P. C. S. I. T. o. t. C. A. o. Sciences (Ed), Insects of Xizang. Science Press, Beijing, pp. 427-447.
  • Wang, S.-F. (1985) Apidae - Bombus. In: [Organisms of the Tumuefeng region of Tianshan], pp. 160-165.
  • Wang, S.-F. (1992) Hymenoptera: Apoidae (II) - Bombus. In: The series of the scientific expedition to the Hengduan Mountains region of Qinghai-Xizang Plateau. Chinese Academy of Sciences, pp. 1422-1430.
  • Wang, S.-F. & Yao, J. (1996) Hymenoptera: Apodae [sic] - Bombini. In: F.-S. Huang (Ed), Insects of the Karakorum-Kunlun mountains. Science Press, Beijing, pp. 303-309.
  • Wappler, T., De Meulemeester, T., Aytekin, A.M., Michez, D. & Engel, M.S. (2012) Geometric morphometric analysis of a new Miocene bumble bee from the Randeck Maar of southwestern Germany (Hymenoptera: Apidae). Systematic Entomology, 37, 784-792.
  • Williams, P.H. (1985) A preliminary cladistic investigation of relationships among the bumble bees (Hymenoptera, Apidae). Systematic Entomology, 10, 239-255.
  • Williams, P.H. (1991) The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology), 60, 1-204.
  • Williams, P.H. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology), 67, 79-152. [updated at www.nhm.ac.uk/bombus/ accessed 2015].
  • Williams, P.H. (2004) Genus Bombus Latreille. In: R. K. Gupta (Ed), An annotated catalogue of the bee species of the Indian Region. Jai Narain Vyas University, Jodhpur, p. Web page providing a list of bumblebees of India and adjacent countries with common synonyms and a summary of their distributions within the region.
  • Williams, P.H. (2007) The distribution of bumblebee colour patterns world-wide: possible significance for thermoregulation, crypsis, and warning mimicry. Biological Journal of the Linnean Society, 92, 97-118.
  • Williams, P.H. (2011) Bumblebees collected by the Kyushu University Expeditions to Central Asia (Hymenoptera, Apidae, genus Bombus). Esakia, 50, 27-36.
  • Williams, P.H., An, J.-D., Brown, M.J.F., Carolan, J.C., Goulson, D., Huang, J.-X., et al. (2012) Cryptic bumblebee species: consequences for conservation and the trade in greenhouse pollinators. PLoSONE, 7, 1-8.
  • Williams, P.H., An, J.-D. & Huang, J.-X. (2011) The bumblebees of the subgenus Subterraneobombus: integrating evidence from morphology and DNA barcodes (Hymenoptera, Apidae, Bombus). Zoological Journal of the Linnean Society, 163, 813-862.
  • Williams, P.H., Araujo, M.B. & Rasmont, P. (2007) Can vulnerability among British bumblebee (Bombus) species be explained by niche position and breadth? Biological Conservation, 138, 493-505.
  • Williams, P.H., Brown, M.J.F., Carolan, J.C., An, J.-D., Goulson, D., Aytekin, A.M., et al. (2012) Unveiling cryptic species of the bumblebee subgenus Bombus s. str. world-wide with COI barcodes (Hymenoptera: Apidae). Systematics and Biodiversity, 10, 21-56.
  • Williams, P.H., Bystriakova, N., Huang, J.-X., Miao, Z.-Y. & An, J.-D. (2015) Bumblebees, climate and glaciers across the Tibetan plateau (Apidae: Bombus Latreille). Systematics and Biodiversity, 13, 164-181.
  • Williams, P.H., Byvaltsev, A.M., Cederberg, B., Berezin, M.V., Odegaard, F., Rasmussen, C., et al. (2015) Genes suggest ancestral colour polymorphisms are shared across morphologically cryptic species in arctic bumblebees. PLoS ONE, 10, 1-26.
  • Williams, P.H., Byvaltsev, A.M., Sheffield, C.S. & Rasmont, P. (2012) Bombus cullumanus - an extinct European bumblebee species? Apidologie, 44, 121-132.
  • Williams, P.H. & Cameron, S.A. (1993) Bumble bee (Bombus Latr.) records from the Valley of Flowers, Uttar Pradesh. Bulletin of Entomology, 31, 125-127.
  • Williams, P.H., Cameron, S.A., Hines, H.M., Cederberg, B. & Rasmont, P. (2008) A simplified subgeneric classification of the bumblebees (genus Bombus). Apidologie, 39, 46-74.
  • Williams, P.H., Ito, M., Matsumura, T. & Kudo, I. (2010) The bumblebees of the Nepal Himalaya (Hymenoptera: Apidae). Insecta Matsumurana, 66, 115-151.
  • Williams, P.H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101-190.
  • Williams, S.T. & Ozawa, T. (2006) Molecular phylogeny suggests polyphyly of both the turban shells (family Turbinidae) and the superfamily Trochoidea (Mollusca: Vestigastropoda). Molecular Phylogenetics and Evolution, 39, 33-51.
  • Woodard, S.H., Fischman, B.J., Venkat, A., Hudson, M.E., Varala, K., Cameron, S.A., et al. (2011) Genes involved in convergent evolution of eusociality in bees. Proceedings of the National Academy of Sciences USA, 108, 7472-7477.
  • Wu, C.-F. (1941) Catalogus insectorum sinensium (Vol. 6). Peiping, Department of Biology, Yenching University.
  • Xu, J.-C., Grumbine, R.E., Shrestha, A., Erikson, M., Yang, X.-F., Wang, Y., et al. (2009) The melting Himalayas: cascading effects of climate change on water, biodiversity, and livelihoods. Conservation Biology, 23, 520-530.
  • Yao, J. & Wang, S.-F. (2005) Hymenoptera, Apidae, Bombini. In: X. Yang (Ed), Insect fauna of middle-west Qinling range and south mountains of Gansu province. Science Press, Beijing, pp. 890-896.
  • Yefremova, Z.A. (2001) Faunal and ecological notes on species of Bombus Latreille, 1802, and Psithyrus Lepeletier, 1833 (Hymenoptera: Apidae) from the Tarbagatai and Saur mountain ranges in eastern Kazakhstan. Entomologist's Gazette, 52, 269-275.
  • Zachos, J.C., Dickens, G.R. & Zeebe, R.E. (2008) An early Cenozoic perspective on greenhouse warming and carbon-cycle dynamics. Nature, 451, 279-283.
  • Zachos, J.C., Pagani, M., Sloan, L., Thomas, E. & Billups, K. (2001) Trends, rhythms, and aberrations in global climate 65 Ma to Present. Science, 292, 686-693.
  • Zhang, J.-J., Kapli, P., Pavlidis, P. & Stamatakis, A. (2013) A general species delimitation method with applications to phylogenetic placements. Bioinformatics, 29, 2869-2876.
  • Zink, R.M. & Barrowclough, G.F. (2008) Mitochondrial DNA under siege in avian phylogeography. Molecular Ecology, 17, 2107-2121.