brunneum brunneum (Fabricius)

Vespa brunnea Fabricius 1793: 264 —“Tranquebariae” (København); 1804: 260.

Odynerus brunneus; Latreille 1806: Tab. XIV fig. 3.

Rhynchium brunneum; Spinola 1808: 189.—de Saussure 1852: 112, p l. XIV figs. 4, 4a–d (Rygchium; expl. plate: bruneum [!]; female, male) 1855: 172.— Smith 1857: 44 (cat.).— Horne 1870: 168 (ethology).— Smith 1871a: v; 1871b: 374 (cat.).— Gribodo 1884: 357 (Rygchium; Burma).— Bingham 1888: 187 (India).— Kirby 1894: 109 (Damma I.)— Bingham 1897: 353 (key), 355, fig. 103 (distribution); 1898: 110.— Cameron 1900: 533 (compared to R. rugolatum); 1901: 243 (New Britain).— Rothney 1903: 106 (India).—de Saussure et al. 1904: 195 (Cambodia).— Schulz 1905 [1904]: 422 (probably distinct species).— Stebbing 1905: 675, fig. 70 (ethology).— Bingham 1905: 47 (Malaya).— Paiva 1907: 15 (India).— Maxwell-Lefroy & Howlett 1909: 212–213 (prey: caterpillars).— Strand 1910: 46–52 (Ceylon).— Rohwer 1911: 479 (Rhygchium; Formosa).— Dutt 1912: 240–241 (prey: caterpillars).— Fullaway 1913: 283 (Guam).— Dover 1921: 386 (India).— Dusmet 1930: 105 (India).—Hoffman 1938: 457 (Rychium; Kwangtung).— Iwata 1965: 105, pl. 9 fig. 8 (anat. ovary), pl. 10 fig. 8 (egg).—van der Vecht 1979: 28 (Nepal).— Giordani Soika 1986: 73 (India; Burma; Vietnam).— Gusenleitner 1988: 181 (Thailand).— Madl et al. 1996: 831 (Seychelles).— Gerlach 2003: 26.— Xu & Han 2008: 1129–1137 (venom).— Srinivasan & Kumar 2010: 1319, image 10 (India).— Oh et al. 2011: 55 (venom).— Srinivasan & Kumar 2013: 120 (India; Myanmar; Nepal).— Baek et al. 2013:497–502 (venom).— Pham & Li 2015: 136 (Vietnam; list).— Siddiqui et al. 2015: 509, fig. 20 (Pakistan).— Lee et al. 2016: 32 (venom).— Deshmukh 2017: 121, image 5 (India).— Rasool et al. 2017: 649 (bruneum [!]; Pakistan).— Dang 2018: 147, figs. 6–8 (nesting; Vietnam).— Mazumdar et al. 2021: 363 (India).— Guido-Patiño & Plisson 2022: 100119 (venom).

Rhynchium haemorrhoidale var. brunneum; de Saussure 1862: 190.— Maindron 1882: 277 (Rynchium; cat).— Kohl 1908: 311 (Rygchium; Neupommern).—Hoffman 1938: 457 (Rygchium; China).

Rhynchium quinquecinctum var. brunneum; Dalla Torre 1894: 45 (cat.).

Rynchium khandalense Dusmet 1930: 105, male, female—[India] “Khandala” (holotype male Madrid).— Giordani Soika 1994b: 47 (hoplomerocephaous; syn. of R. brunneum brunneum).

Odynerus haemorrhoidalis var. quinquecinctus; Bequaert & Yasmatsu 1939: 316 (distr.).— Yasumatsu 1945: 36 (distr.). Misidentification.

Rygchium haemorrhoidale var. quinquecinctum; Krombein 1949: 370 (list). Misidentification.

Rhynchium brunneum brunneum; Giordani Soika 1994b: 40 (key), 46 (India; Bangladesh; Burma; Thailand; Cambodia; Vietnam).— Madl 1995: 398 (Thailand).— Gusenleitner 2006a: 692 (India); 2008b: 769 (Indonesia: Yamdena); 2011b: 1362 (Laos).— Nugroho et al. 2012 [2011]: 120 (list).— Kumar & Sharma 2013: 105, 106 (key), figs. 1–8 (India).— Nguyen et al. 2014: 13 (Vietnam; list).— Kumar and Sharma 2014b: 564, 573, image 16 (India); 2015: 8098, 8123, image 31 (India).— Pannure et al. 2016: 38 (India).— Pham & Kumar 2016: 531, 534 (list) figs. 1–5 (Vietnam).— Nguyen 2016: 469 (Vietnam).— Jadhav et al. 2017: 142, 145 (India).— Rafi et al. 2017: 16 (Pakistan).—Kumar et al. 2018 [2017]: 16 (India).— Tan et al. 2018: 145 (list).—Nidup et al. 2018: 11688, fig. 7 (Bhutan).— Kumar & Rajmohana 2018: 338 (India).— Li et al. 2019: 157, 170 (key), figs. 110–112 (China).—Kumar & Sureshan 2019: 122 (India).— Gawas et al. 2019: 277 (India).— Dang & Nguyen 2019: 427–436 (nest, ethology; Vietnam).— Gawas et al. 2020: 38 (list).— Ahmad et al. 2021: 629–638 (Pakistan).— Kolosova et al. 2022: 1–16 (nesting ecology; Seychelles).— Luo et al. 2022: 529 (mitogenome; phylogenetic relationships).— Udayakumar et al. 2022: 794–810 (ethology).— Dal Pos et al. 2022: 60 (specs.).— Carpenter 2023: 13 (distr.).

? Rhynchium quinquecinctum; Kusigemati et al. 1996: 12 (Malakal).

Distr.: Pakistan; India; Bangladesh; Nepal; Bhutan; China; Myanmar; Thailand; Cambodia; Laos; Vietnam; Malaysia; Indonesia: Tanimbar; New Britain; Marianas;? Palau; adventive in Seychelles.