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<title>Fusarium houttuyniae Y. K. He, Yong Wang</title>
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<h2>
<i>Fusarium houttuyniae</i>
Y. K. He, Yong Wang bis,
<i>sp. nov.</i>
Fig. 2.</h2>

<p>Mycobank: MB846236; FOF number: 14109</p>
<p>
Etymology:—
<i>houttuyniae</i>
refers to the host genus
<i>Houttuynia</i>, which the holotype was isolated.</p>

<p>
Holotype:— CHINA, Guangxi Province, Nanning City, Guang Xi Medicinal Botanical Garden, 108°19’ E, 22°51’ N, leaf of
<i>Houttuynia cordata</i>, November 2017, collected by Qian Zhang (Collection number: HGUP 10007, culture number: GUCC 21004).</p>

<p>
Additional specimens examined:— CHINA, Guangxi Province, Nanning City, Guang Xi Medicinal Botanical Garden, 108°19’ E, 22°51’ N, leaf of
<i>Houttuynia cordata</i>, November 2017, collected by Qian Zhang (Collection number: HGUP 10007, culture number: GUCC 21008).</p>

<p>
Description:
<b>Colonies</b>
on PDA media with an average radial growth rate of 2.1‒3.0 mm/d at 24℃. The colony was white with abundant and flocculent aerial hyphae, but later turned light purple, odour absent. Colony margins regularly.
<b>Conidiophores</b>
unbranched or sparingly branched, aerial phialides subulate to subcylindrical, straight, thin- and smooth- walled.
<b>Microconidia</b>
hyaline, ellipsoidal to falcate, smooth- and thin-walled, 0‒1 septate; 0-septate, 4‒10 × 1‒5 (av. 6.4 ± 1.53 × 3.2 ± 1.08) µm or 1-septate, 9‒17 × 2‒6 (av. 13.9 ± 2.11 × 4.3 ± 1.21) µm.
<b>Macroconidia</b>
formed on aerial mycelium, falcate, curved dorsiventrally with almost parallel sides tapering slightly towards both ends, with a blunt to papillate, curved apical cell and a blunt to foot-like basal cell, hyaline, smooth- and thin-walled, 1‒3 septate; 1-septate conidia: 12‒20 × 2‒5 (av. 16.5 ± 2.1 × 3.5 ± 0.65) µm; 2-septate conidia: 15‒22 × 2‒4 (av. 18.15 ± 1.92 × 3 ± 0.44) µm; 3-septate conidia: 28‒45 × 3‒6 (av. 38.5 ± 3.18 × 4.71 ± 0.7) µm.
<b>Chlamydospores</b>
globose to sub globose, 4.5‒9 (av. 6.92 ± 1.06) µm diam.</p>

<p>
Notes: Phylogenetically,
<i>F</i>.
<i>houttuyniae</i>
(GUCC 21004) formed a well-supported, distinct clade sister to
<i>F. liriopes</i>
(GUCC 21005), and
<i>F. nirenbergiae</i>
(CBS 129.24) (Fig. 1). Between
<i>F. houttuyniae</i>
and
<i>F. liriopes</i>, there are 6/606 character differences in
<i>cmdA</i>, 1/876 character in
<i>rpb2</i>, 1/621 character in
<i>tef1</i>, and 10/534 characters in
<i>tub2</i>. Between
<i>F. houttuyniae</i>
and
<i>F. nirenbergiae</i>, there are 4/606 characters in
<i>cmdA</i>, 2/876 characters in
<i>rpb2</i>, 7/621 characters in
<i>tef1</i>
and 12/534 characters in
<i>tub2</i>
(Table 2). This species (conidia 1‒3-septate) can be distinguished from
<i>F. nirenbergiae</i>
(0‒5-septate; (Zhao
<i>et al.</i>
2020)) based on the septation of the macroconidia.
<i>Fusarium houttuyniae</i>
differs from
<i>F. liriopes</i>
by having bigger conidia (<i>F. houttuyniae</i>: 1-septate conidia: 12‒20 × 2‒5 µm; 2-septate conidia: 15‒22 × 2‒4 µm; 3-septate conidia: 28‒45 × 3‒6 µm.
<i>vs. F. liriopes</i>: 1-septate conidia: 11‒15 × 2‒3 µm; 2-septate conidia: 14‒22 × 2‒4 µm; 3-septate conidia: 18‒28 × 3‒5 µm.). Thus, we propose
<i>F. houttuyniae</i>
as a novel species.</p>

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