Year,Site,Mean,SE,Sample 1859,Den,0.50525,0.12175,2 1860,Den,0.45525,0.04075,2 1861,Den,0.49525,0.15625,2 1862,Den,0.50075,0.15125,2 1863,Den,0.336,0.0105,2 1864,Den,0.60925,0.15025,2 1865,Den,0.47625,0.10275,2 1866,Den,0.397,0.0835,2 1867,Den,0.2825,0.0225,2 1868,Den,0.383666667,0.056847994,3 1869,Den,0.319,0.046981379,3 1870,Den,0.452,0.007522189,3 1871,Den,0.3765,0.059378026,3 1872,Den,0.5585,0.209509049,4 1873,Den,0.65975,0.213344938,4 1874,Den,0.558125,0.201098481,4 1875,Den,0.603375,0.1935304,4 1876,Den,0.52875,0.183285126,4 1877,Den,0.611375,0.171407577,4 1878,Den,0.868916667,0.268915644,6 1879,Den,0.622833333,0.164697079,6 1880,Den,1.023142857,0.252250813,7 1881,Den,1.1195625,0.241326439,8 1882,Den,0.990166667,0.172731728,9 1883,Den,1.043611111,0.156893603,9 1884,Den,1.13345,0.161952607,10 1885,Den,1.074136364,0.167072209,11 1886,Den,1.097818182,0.172816314,11 1887,Den,1.1875,0.201430701,11 1888,Den,0.953958333,0.154741799,12 1889,Den,0.822166667,0.138546887,12 1890,Den,0.940653846,0.123572041,13 1891,Den,0.864,0.103609273,13 1892,Den,0.930230769,0.123564833,13 1893,Den,0.968807692,0.127933471,13 1894,Den,1.044615385,0.142080446,13 1895,Den,1.151115385,0.161128733,13 1896,Den,0.938357143,0.128268428,14 1897,Den,0.919678571,0.143251149,14 1898,Den,0.870928571,0.121668369,14 1899,Den,0.824,0.117205828,14 1900,Den,1.026366667,0.151639912,15 1901,Den,0.855366667,0.121001776,15 1902,Den,0.900766667,0.141948917,15 1903,Den,0.936933333,0.140017931,15 1904,Den,0.868766667,0.132746015,15 1905,Den,0.9302,0.136487491,15 1906,Den,0.8117,0.107737134,15 1907,Den,0.796533333,0.108372137,15 1908,Den,0.824,0.104652452,15 1909,Den,0.733266667,0.088640873,15 1910,Den,0.754533333,0.088777296,15 1911,Den,0.8464,0.113821071,15 1912,Den,0.815033333,0.095366043,15 1913,Den,0.969633333,0.112971704,15 1914,Den,0.767466667,0.092802228,15 1915,Den,0.7536,0.074075206,15 1916,Den,0.876,0.091936352,15 1917,Den,0.801133333,0.076880604,15 1918,Den,0.831966667,0.068195229,15 1919,Den,0.912066667,0.076203126,15 1920,Den,0.796133333,0.068951388,15 1921,Den,0.8611,0.079155531,15 1922,Den,0.770566667,0.079623771,15 1923,Den,0.690766667,0.082116741,15 1924,Den,0.70309375,0.096173067,16 1925,Den,0.7371875,0.103246266,16 1926,Den,0.866294118,0.095859167,17 1927,Den,0.761,0.076656606,17 1928,Den,0.789611111,0.080882061,18 1929,Den,0.790473684,0.097766606,19 1930,Den,0.940026316,0.128742793,19 1931,Den,0.7233,0.120729822,20 1932,Den,0.9415,0.134096696,21 1933,Den,0.89,0.102612081,22 1934,Den,0.927590909,0.102739708,22 1935,Den,1.038568182,0.096072474,22 1936,Den,0.981159091,0.095488805,22 1937,Den,0.847409091,0.102617595,22 1938,Den,0.933318182,0.11122375,22 1939,Den,0.933113636,0.121133968,22 1940,Den,0.870795455,0.101586584,22 1941,Den,1.037869565,0.142391333,23 1942,Den,0.938673913,0.122198607,23 1943,Den,1.130434783,0.129221718,23 1944,Den,1.229583333,0.118899462,24 1945,Den,0.97476,0.098226363,25 1946,Den,1.22242,0.120444841,25 1947,Den,0.93146,0.101665231,25 1948,Den,1.0637,0.095738302,25 1949,Den,0.87546,0.077024877,25 1950,Den,0.913596154,0.068975185,26 1951,Den,0.941980769,0.067665253,26 1952,Den,0.768557692,0.056420102,26 1953,Den,1.019740741,0.078593654,27 1954,Den,1.11862963,0.080778811,27 1955,Den,0.943111111,0.080423498,27 1956,Den,0.883648148,0.072523274,27 1957,Den,1.108925926,0.104035458,27 1958,Den,1.122035714,0.234459685,28 1959,Den,1.035051724,0.144030704,29 1960,Den,0.897413793,0.097416852,29 1961,Den,1.097465517,0.121483533,29 1962,Den,1.147466667,0.123690018,30 1963,Den,1.214283333,0.112317056,30 1964,Den,1.057316667,0.099422844,30 1965,Den,1.227032258,0.121304232,31 1966,Den,1.371935484,0.124482256,31 1967,Den,0.975546875,0.112552878,32 1968,Den,0.915140625,0.110219382,32 1969,Den,1.060818182,0.106993767,33 1970,Den,1.123848485,0.101346878,33 1971,Den,0.925515152,0.10156595,33 1972,Den,1.052681818,0.099272243,33 1973,Den,0.905318182,0.073163681,33 1974,Den,1.022220588,0.148645915,34 1975,Den,0.920472222,0.120367381,36 1976,Den,1.090256757,0.122080074,37 1977,Den,1.116171053,0.131128617,38 1978,Den,0.998039474,0.113406442,38 1979,Den,1.028315789,0.119976385,38 1980,Den,0.96881579,0.115383901,38 1981,Den,1.233184211,0.142672487,38 1982,Den,0.970710526,0.1085105,38 1983,Den,1.237631579,0.128998709,38 1984,Den,1.131907895,0.108448388,38 1985,Den,1.085384615,0.109585987,39 1986,Den,1.230333333,0.118446295,39 1987,Den,1.1332625,0.117477484,40 1988,Den,1.103025,0.118973341,40 1989,Den,0.9789875,0.099084229,40 1990,Den,1.1205125,0.118387557,40 1991,Den,1.1621125,0.12445057,40 1992,Den,0.820325,0.085446759,40 1993,Den,1.0808875,0.097789568,40 1994,Den,1.0201,0.095219243,40 1995,Den,1.0716875,0.114546451,40 1996,Den,1.084875,0.119562549,40 1997,Den,0.982175,0.097509514,40 1998,Den,1.0053625,0.09582192,40 1999,Den,1.084175,0.097057144,40 2000,Den,1.0201375,0.102614792,40 2001,Den,1.171525,0.122895223,40 2002,Den,1.312166667,0.137563897,39 2003,Den,1.455435897,0.147227115,39 2004,Den,1.201884615,0.130535623,39 2005,Den,1.397820513,0.145768904,39 2006,Den,1.188730769,0.109569645,39 2007,Den,1.069576923,0.093921198,39 2008,Den,1.028320513,0.082036175,39 2009,Den,0.799641026,0.07145937,39 2010,Den,1.111564103,0.093169151,39 2011,Den,0.931179487,0.08773953,39 2012,Den,0.915641026,0.101179111,39 2013,Den,1.022092105,0.107300881,38 2014,Den,1.015302632,0.105938127,38 2015,Den,0.952157895,0.099958494,38 2016,Den,0.941552632,0.104368757,38 2017,Den,1.025315789,0.109567497,38 1827,Control,0.44425,0.16425,2 1828,Control,0.51575,0.20125,2 1829,Control,0.451,0.202,2 1830,Control,0.4145,0.2625,2 1831,Control,0.37975,0.20625,2 1832,Control,0.4155,0.2605,2 1833,Control,0.51275,0.32975,2 1834,Control,0.41975,0.22325,2 1835,Control,0.3255,0.1265,2 1836,Control,0.39025,0.19825,2 1837,Control,0.3745,0.1475,2 1838,Control,0.4705,0.162,2 1839,Control,0.28425,0.07525,2 1840,Control,0.31725,0.07275,2 1841,Control,0.4195,0.095,2 1842,Control,0.3775,0.1325,2 1843,Control,0.359,0.121,2 1844,Control,0.29025,0.09225,2 1845,Control,0.3235,0.0765,2 1846,Control,0.32525,0.09625,2 1847,Control,0.33375,0.13125,2 1848,Control,0.30975,0.12125,2 1849,Control,0.48125,0.27025,2 1850,Control,0.495,0.217,2 1851,Control,0.4545,0.1925,2 1852,Control,0.44575,0.17875,2 1853,Control,0.43875,0.11425,2 1854,Control,0.352,0.058,2 1855,Control,0.4035,0.1465,2 1856,Control,0.452,0.1405,2 1857,Control,0.31825,0.06525,2 1858,Control,0.38625,0.11825,2 1859,Control,0.3595,0.1225,2 1860,Control,0.39175,0.07425,2 1861,Control,0.409166667,0.052979031,3 1862,Control,0.358166667,0.05543189,3 1863,Control,0.347666667,0.041134265,3 1864,Control,0.429833333,0.088883976,3 1865,Control,0.344333333,0.065437205,3 1866,Control,0.356833333,0.073683068,3 1867,Control,0.250333333,0.048761608,3 1868,Control,0.311375,0.038435755,4 1869,Control,0.267,0.018984643,4 1870,Control,0.33875,0.069789117,4 1871,Control,0.35075,0.099048073,4 1872,Control,0.3913,0.097142627,5 1873,Control,0.3764,0.099638145,5 1874,Control,0.3204,0.072664709,5 1875,Control,0.3237,0.083405725,5 1876,Control,0.2999,0.077077461,5 1877,Control,0.3499,0.106984041,5 1878,Control,0.363666667,0.088247726,6 1879,Control,0.3695,0.086896682,6 1880,Control,0.436416667,0.06720211,6 1881,Control,0.452166667,0.081479411,6 1882,Control,0.453666667,0.099968217,6 1883,Control,0.438583333,0.084706593,6 1884,Control,0.453583333,0.098654567,6 1885,Control,0.463583333,0.114918407,6 1886,Control,0.514083333,0.123397692,6 1887,Control,0.462357143,0.078122364,7 1888,Control,0.4195,0.094282125,7 1889,Control,0.606,0.224088929,8 1890,Control,0.5986875,0.20080194,8 1891,Control,0.50675,0.118931478,8 1892,Control,0.480125,0.121997173,8 1893,Control,0.486375,0.118597675,8 1894,Control,0.451666667,0.087761277,9 1895,Control,0.558333333,0.091600089,9 1896,Control,0.527666667,0.085482901,9 1897,Control,0.591454545,0.096520593,11 1898,Control,0.511545455,0.07922346,11 1899,Control,0.519636364,0.071790766,11 1900,Control,0.636545455,0.095647612,11 1901,Control,0.557772727,0.081203123,11 1902,Control,0.550590909,0.087271776,11 1903,Control,0.579333333,0.069092202,12 1904,Control,0.552083333,0.082926374,12 1905,Control,0.574416667,0.083322048,12 1906,Control,0.578333333,0.095175423,12 1907,Control,0.516708333,0.074525988,12 1908,Control,0.5035,0.070121567,12 1909,Control,0.447541667,0.057926627,12 1910,Control,0.533807692,0.0842504,13 1911,Control,0.528423077,0.085636902,13 1912,Control,0.51021875,0.059790101,16 1913,Control,0.520205882,0.057200757,17 1914,Control,0.423166667,0.051422669,18 1915,Control,0.442277778,0.057084406,18 1916,Control,0.490075,0.052682131,20 1917,Control,0.507190476,0.055285563,21 1918,Control,0.46847619,0.048146991,21 1919,Control,0.490809524,0.050345112,21 1920,Control,0.470071429,0.050413909,21 1921,Control,0.563404762,0.063488981,21 1922,Control,0.495452381,0.057776137,21 1923,Control,0.471977273,0.044733972,22 1924,Control,0.462195652,0.049870472,23 1925,Control,0.516152174,0.056901833,23 1926,Control,0.622583333,0.082155515,24 1927,Control,0.6103125,0.061730611,24 1928,Control,0.6759,0.062480737,25 1929,Control,0.6574,0.066215387,25 1930,Control,0.70594,0.061990425,25 1931,Control,0.56694,0.060308197,25 1932,Control,0.70292,0.069143532,25 1933,Control,0.63464,0.060899648,25 1934,Control,0.71022,0.067145893,25 1935,Control,0.79652,0.07447953,25 1936,Control,0.77128,0.082306131,25 1937,Control,0.699519231,0.070733462,26 1938,Control,0.716,0.066575493,27 1939,Control,0.644481481,0.065282072,27 1940,Control,0.639388889,0.054623461,27 1941,Control,0.620446429,0.061184237,28 1942,Control,0.658017241,0.059156621,29 1943,Control,0.783310345,0.067959934,29 1944,Control,0.860806452,0.094885995,31 1945,Control,0.641322581,0.071954363,31 1946,Control,0.84484375,0.088895174,32 1947,Control,0.77121875,0.081378951,32 1948,Control,0.794924242,0.078894643,33 1949,Control,0.632954545,0.061611881,33 1950,Control,0.688515152,0.0626001,33 1951,Control,0.685560606,0.061181056,33 1952,Control,0.5585,0.044707123,34 1953,Control,0.594814286,0.048438751,35 1954,Control,0.731114286,0.062547797,35 1955,Control,0.688157143,0.063818215,35 1956,Control,0.652114286,0.057003789,35 1957,Control,0.784457143,0.064978986,35 1958,Control,0.645071429,0.058263835,35 1959,Control,0.688742857,0.063251417,35 1960,Control,0.610842857,0.049469674,35 1961,Control,0.693657143,0.051684075,35 1962,Control,0.787621622,0.059482593,37 1963,Control,0.881135135,0.06584795,37 1964,Control,0.748539474,0.052060018,38 1965,Control,0.869381579,0.061584341,38 1966,Control,1.001294872,0.077905648,39 1967,Control,0.771589744,0.069425984,39 1968,Control,0.717410256,0.05979841,39 1969,Control,0.786038462,0.068309169,39 1970,Control,0.802153846,0.061959831,39 1971,Control,0.700538462,0.061129352,39 1972,Control,0.791125,0.063883748,40 1973,Control,0.6347125,0.050349413,40 1974,Control,0.6388875,0.060051688,40 1975,Control,0.621775,0.056284946,40 1976,Control,0.7281625,0.06508708,40 1977,Control,0.7777375,0.066557354,40 1978,Control,0.6811875,0.057890751,40 1979,Control,0.686425,0.062126831,40 1980,Control,0.6118625,0.056944673,40 1981,Control,0.8128625,0.07401768,40 1982,Control,0.622375,0.055821627,40 1983,Control,0.79635,0.063171529,40 1984,Control,0.6738,0.055510258,40 1985,Control,0.6281125,0.05166425,40 1986,Control,0.728925,0.055960925,40 1987,Control,0.689475,0.057209976,40 1988,Control,0.683375,0.059746389,40 1989,Control,0.6571,0.055057287,40 1990,Control,0.7233375,0.06417642,40 1991,Control,0.6988,0.069237481,40 1992,Control,0.4639125,0.045829428,40 1993,Control,0.607,0.055128039,40 1994,Control,0.6107875,0.062783745,40 1995,Control,0.6287,0.06907752,40 1996,Control,0.60675,0.058018308,40 1997,Control,0.63025,0.058390353,40 1998,Control,0.5500625,0.050714113,40 1999,Control,0.5772625,0.04967525,40 2000,Control,0.6242,0.054383744,40 2001,Control,0.69,0.060544956,40 2002,Control,0.7380625,0.070619757,40 2003,Control,0.7304875,0.06770306,40 2004,Control,0.6916625,0.061815615,40 2005,Control,0.834575,0.071430302,40 2006,Control,0.68645,0.061552346,40 2007,Control,0.5992875,0.046088873,40 2008,Control,0.6044,0.045945296,40 2009,Control,0.4529375,0.038155036,40 2010,Control,0.6488875,0.052451123,40 2011,Control,0.555,0.049295989,40 2012,Control,0.5297125,0.044263097,40 2013,Control,0.6058125,0.052105329,40 2014,Control,0.5829125,0.047571124,40 2015,Control,0.5655125,0.045096931,40 2016,Control,0.5502375,0.046043044,40 2017,Control,0.6235,0.052720784,40