(Figs 9, 21, 33)
Description. Holotype male. Size, 10.2 mm x 4.3 mm. Sculpture irregular, rough dorsally, shining bronzyblack with elytral foveae dull brassy; more shining ventrally than above. Head with frons greenish and transverse rusty area near middle of eyes and larger transverse goldish-rusty area just above antennal socket, both areas interrupted at middle of frons; occiput with broad smooth, grooved longitudinal carina forming indistinct Y anteriorly; frons nearly flat, its surface coarsely, evenly punctate and clothed with long white setae; clypeus greenish-gold, semicircularly rounded on each side of median groove; antennae greenish at base, apical four antennomeres magenta, slightly narrowed toward apex, intermediate antennomeres compact- about as long as wide; third antennomere shorter than combined length of next two antennomeres. Pronotum less than twice as wide as long, widest at anterior ¼ with lateral margin slightly undulating to posterior ¼ then narrowed sharply; anterior margin sinuate; posterior margin strongly sinuate; disc convex, uneven with longitudinal median depression deeper in middle and raised area lateral to it behind middle of disc; surface irregularly punctate with larger punctations laterally; disc with raised callosities with some parallel transverse ridges on either side of middle, and with long white setae laterally. Elytra distinctly wider than pronotum, sides rounded basally then subparallel to widest at posterior 1 / 3; lateral margin serrulate from basal 1 / 5 to apex with serrulations largest on apical half; three raised dark indistinct transverse bands- one near basal 1 / 5, one just behind middle, and last at apical 1 / 3; depressed brassy irregular indistinct foveae at basal 2 / 5 and 4 / 5 not reaching sutural elytral margin; four longitudinal costae: 1 st, indistinct, laterally parallel with apical half of elytra, 2 nd, about ¼ from lateral margin, strong near anterior fovea, weaker apically, 3 rd, indistinct except near anterior fovea, and 4 th, strongest costa, about a ¼ from and parallel with sutural margin, from near base to apex. Underside: bronze, with long white prosternal setae; laterally with dark bluish raised callosities on abdominal ventrites 2, 3, and 4. Last ventrite semicircularly emarginate; profemur, pro- and mesotibiae greenish-bronze. Inner margin of protibia with row of 6 small teeth; metatibia straight and brownish; tarsi greenish-blue. Male genitalia (Fig. 21) strongly constricted before middle and tapering toward base, spines of paramere noticeable and pointing toward base of male genitalia.
Allotype female. Size, 9.5 mm x 4.0 mm. Frons brownish-magenta with moderately sized irregular raised areas on either side of midline, two near antennal bases and two below vertex; antennae dark bronze with sensory area on apical eight antennomeres reddish-brown; clypeus maroon, margin greenish-black. pronotal raised areas bluish-black with interconnecting transverse ridges interrupted by median depression. Scutellum black. Elytral discal foveae rather indistinct, more brassy than darker elytra. Underside: with legs more magenta than males, tarsi bluish-black. There is some color variation in the female's clypeus, usually bronzegreen. Female pygidium (Fig. 33) with a shallow depression on either side of the median carina.
Type specimens. Described from 144 males and 164 females. Holotype male labeled USA, CALIFOR- NIA: Shasta Co., 4 mi W Redding, 19 -VI- 1965. F. M. & V. S. Beer. Beating Quercus garryana Dougl. ex Hook. FRANK M. BEER COLLECTION, Univ. of Idaho. Acc. Dec. 1986. Female allotype labeled: USA, CALIFORNIA: Shasta Co., 6 S. Lakehead, 20 -VI- 1965, F. M. & V. S. Beer. From Quercus garryana Dougl logs. FRANK M. BEER COLLECTION, Univ. of Idaho. Acc. Dec. 1986. Holotype (red label) and allotype (blue label) specimens deposited in USNM. Paratypes: ARIZONA: 1 ɗ Huachuca Mts., Copper Canyon S. end, 15 -V- 85, G. H. Nelson; 1 ɗ Yuma Co., Yuma vic. Moralis Dam, VI-VIII- 82 (emg.), C. L. Bellamy. CAL- IFORNIA: 1 Ψ Autioch, 26 -V- 33 - M. Cazier; 1 ɗ, 1 Ψ Bass Lake, 28 -IV- 34, R. P. Allen; Boulevard 1 ɗ 9 -VII- 52, D. J. & J. N. Knull; Calaveras Co., 1 Ψ Mokell Hill, 22 -VI- 21, F. E. Blaisdell. 2 ɗɗ, 4 ΨΨ Castella, VII- 12, Van Dyke; Colusa Co., 1 ɗ, 4 ΨΨ 15 mi SW Williams on Hwy 20 @ mile marker 6 80, 25 -IV- 92, T. C. MacRae; Contra Costa Co., 4 ΨΨ Alamo, 24 -VI- 47, D. Giuliani; 1 ɗ E. side Mt. Diablo, 12 -V- 62, R. L. Westcott; 1 Ψ Alhambra Val., 20 -V- 19, F. E. Blaisdell, 1 ɗ Vine Hill, 7 -VI- 31 Blaisdell; Davis, 1 Ψ 8 -VI- 38, B. E. White, 1 Ψ 9 -VI- 62, R. L. Westcott; El Dorado Co., 1 Ψ Georgetown, 29 -VIII- 48, D. Giuliani, 1 ɗ 3 mi SW Somerset, 1- 20 -V- 78, B. Warton; 1 ɗ Emigrant Gap, 26 -VI- 37, M. Cazier; Fresno Co., Clovis, 1 ɗ 3 -V-, 1 ɗ 28 -V-, 1 Ψ 3 -VI- 79, all A. J. & M. E. Gilbert; 2 ɗɗ Firebaugh, 6,7 - VI- 75 M. Robey; 1 Ψ Idlewild, 8 -VII- 28, Van Dyke; Imperial Co., 1 Ψ Laguna Dam, 21 -V 1 -79, J. Cope; Kern Co., 3 ɗ Arvin, 29 -VI- 67, L; Frudden; 1 Ψ 81 / 2 mi. SW Bodefish, 1 -VIII- 68. F.M. & V.S. Beer; 1 Ψ 4 mi NE Buttonwillow, 28 -VI- 61, C. W. O'Brien; 1 ɗ 5.3 mi N. Caliente, em. 20-24 -VII- 99, R. Turnbow; 1 ɗ, 2 ΨΨ 2.3 mi E Lost Hills, 28 -VI- 91 T. C. MacRae; Lake Co., 1 ɗ Middletown, 2 -VI- 56, D. Giuliani; 1 ɗ Mt. St. Helena, 22-23 -V- 54, D. Giuliani; La Grange: 1 ɗ 17 -VII- 49, R. P. All e n; 1 Ψ V- 30, 1 Ψ 28 -VII- 61, R. P. Allen; Los Angeles Co., Placerita Canyon Parks, 1 Ψ 10 -VI- 73, 1ɗ 16 - VII- 74, F. T. Hovore; 2 ɗɗ, 2 ΨΨ S. Gabriel Mts., 1 / 2 mi SE Crystal Lake, 11 -VII- 68, F. M. & V. S. Beer; 1 Ψ Crystal Lake, 29 -VI- 50, F. X. Williams; Madera Co., 2 ΨΨ New Columbia Ranch, 11 -VI- 73, M. Robey; Marin Co., 1 ɗ, 1 Ψ Fairfax Canyon. 3 -V- 53, 1Ψ Fairfax, 28 -V- 55, D. Giuliani; 1 ɗ, 1 Ψ Larkspur, 20 -VI- 54, D. Giulani; 1 Ψ Mill Valley, 18 -V- 23, E. P. Van Duzee; 1 Ψ Woodacre, 16 -VI- 57, D. Giuliani; Mendocino Co., 1 ɗ 30 - V- 22, E. R. Leach; 1 ɗ Dos Rios, 27 -V- 39, W. F. Barr; 1 ɗ, 2 ΨΨ Twin Rocks, 10 -VII- 29, E. C. Van Dyke; 2 ɗɗ 28 -V- 22, 20-VI- 20, E. R. Leach; 1 Ψ Midway, 24 -V- 36, E. S. Ross; 1 Ψ 13mi W. Mineral, 9 -VII- 54, D. J. & J. N. Knull; 3 ɗɗ, 4 ΨΨ Mt. St. Helena, 9 -VI- 18, E. P. Van Duzee; 1 ɗ 27 -IV- 50 R. P. Allen, 1 Ψ 11 -V- 2001, Sk. E. Haskins; Napa Co., N. side Howell Mt., 2 mi NNE Angwin, 1300 ', 1 Ψ 26 -VII- 81; 1 Ψ 7 -VII-81. 1Ψ 19 -VII- 81, H. B. Leech; Nevada Co., 2 ΨΨ 10 mi S. Grass Valley, 2 -V- 70 F.M. Beer; 4 ɗɗ, 1 Ψ Northfork, 11 -VI- 33, R. P. Allen; 2 ΨΨ Newton, 14 -VII- 49, D. J. & J. N. Knull; Orange Co., 1 Ψ San Juan Canyon, 13 -VI- 64, E. Weidert; Harding Canyon, 8 ɗɗ, 2 ΨΨ 29 -V- 78, R. Velten; 1 ɗ Parasio Hot Springs, 15 -VII- 54, B. Lot; 1 ɗ Paso Robles, 9 -VI- 27, Blaisdell; Placer Co., 1 Ψ Auburn State Recreation Area, 1-4mi SE Auburn, 1 -VI- 91, T. C. MacRae, 2 ɗɗ, 1 Ψ Placerville, 4 -III- 15, F. B. Herbert; Placerville, 3 ɗɗ 9 -III- 13, 1ɗ - 14, J. J. Sullivan; Placerville, 1 ɗ VI- 15, 1 Ψ 6 -VI- 16, 1ɗ 3 -VII- 16, 1Ψ 26 -VII- 16 (bred) H. E. Burke; 1 ɗ, 1 Ψ Palo Alto, 5 -VI- 18, Burke & Hartman; Pumas Co., 1 ɗ Meadow Valley, 3500-4000 ', 2 -VII- 24, E. C. Van Dyke; Riverside Co., 1 ɗ Anza, 28 -VI- 60; 1 ɗ Pinyon Flats, 13 -V- 64; 1 ɗ Pinyon Flats 4000 ', 10 -VI- 59; 1 Ψ 2.5 mi. N. Pinyon Flats, 21 -VI- 62, all G. H. Nelson; 1 Ψ San Bernadino Mts., Cajon Pass, 29 -V- 62, G. H. Nelson; San Bernardino Co., 1 Ψ Victorville, 6 -VII- 63, R. L. Westcott, 1 Ψ Mt. Baldy, 25 -VIII- 63, Weidert. 1 Ψ Lake Arrowhead, 22 -VI- 64, R. D. Ward; San Diego Co., 2 ΨΨ 5 mi E. Oak Grove, 5 -VII- 59, R. L. Westcott; 6 ɗɗ, 3 ΨΨ 3 mi. E. of Banner, 24 -V- 62 & 1 ɗ 24 -V- 62, G. H. Nelson; 1 ɗ, 1 Ψ San Gabriel Mts., 1 mi above Mt. Baldy Village, Glendora Ridge Rd., 27 -VII- 93, G. H. Nelson; San Joaquin Co., 1 ɗ Stockton, 10 -VIII- 74; 1 Ψ 4 mi S. Ft. Stockton, 2 -I- 76 (reared ex. Salix), K. W. Brown; Santa Clara Co., 1 Ψ Almaden, 9 -VI- 35, E. P. Van Duzee; 1 ɗ Morgan Hill, 27 -VIII- 51, V; Harris; 1 Ψ Guadalupe Res., 4 -IV- 74, B. Carscens; 11 ɗɗ, 10 ΨΨ Guadalupe Cr., IV- 73, C. Y. Kitayama; 4 ɗɗ, 6 ΨΨ Hicks Road, 3 -III- 73 & VI- 74, L. Bezark; 7 ɗɗ, 13 ΨΨ San Jose-Hicks Rds., 20 -VI- 73, L. Bezark; 1 ɗ, 2 ΨΨ 18 -IV- 74, M. J. Robey; 2 ɗɗ, 1 Ψ Los Gatos (IBM), 29 -lV- 73, C. Y. Kitayama; 1 ɗ, 2 ΨΨ San Jacinto Mts., Idyllwild, 9,17,19- VI- 40, F.M. & V.S. Beer; 3 ɗɗ, 2 ΨΨ 11 -IV- 73, M. Ebertz; 1 Ψ San L. Obispo, 30 -V- 32, V. L. Clemence; Santa Cruz Co., 1 Ψ Soquel, 14 -VIII- 67, A. & A. Gillogly; 1 Ψ Santa Cruz Mts., Boulder Crk., 26 -IV- 68, A. & A. Gillogly; 1 ɗ Santa Rosa, 8 -VI- 40, J. R. Helfer. 1 ɗ 21 -IV- 72, G. M. Nishida; Shasta Co., 3 ΨΨ 6 mi. S. LakeHead, 20 -VI- 65, F. M. Beer; 3 ɗɗ, 1 Ψ 4 mi W. Redding 19 -VI- 65, F. M & V.S. Beer; Siskiyou Co., 1 ɗ Shasta Retreat, 3 -VII-05; Solano Co., 4 ɗɗ Putah Canyon 27 -VII- 38, B. E. White; Sonoma Co., Sobre Vista: 2 ɗɗ 24 -IV- 10, 2ΨΨ 6 -V-10. 1ɗ 10 -IV- 11, all Van Dyke; 1 ɗ, 1 Ψ Springville, VII- 33, F. T. Scott; 1 ɗ Stanford Univ., 6 -V- 54, P. S. Bartholomew; Stanislaus Co., 1 ɗ Highway I- 5 -J 18 exit-Newman overpass, 14 -V- 80, F.M. Beer; 1 Ψ La Grange, 20 -VII- 69, R. P. Allen; Tehama Co., 2 ΨΨ Rancho Tehama, 6 - VI- 73, B. Carstens; Trinity Co., 2 ΨΨ 27 -IX- 17, E. R. Leach; 1 ɗ, 1 Ψ Coffee Creek, 7 -VII- 50, D. Giuliani; 1 Ψ 28 -V- 34, T. Aiken; Tulare Co., 1 Ψ Camp Creek, 8 -VI- 17, R. L. Beardsley; 1 Ψ Carrville, 29 -VI- 13, Van Dyke; 1 Ψ 19 -VII- 32, T. S. Scott, 2 ΨΨ Carrville, 2400-2500 ', 8 -VI- 34, G. E. Bohart; 1 Ψ Hot Springs, 2 -VI- 39, Van Dyke; 1 ɗ VI- 30, F. T. Scott; 2 ΨΨ Visalia, VI- 30, F. T. Scott; 10 ɗɗ, 11 Ψ 21 -VIII- 2005, R. D. Haines; 1 ɗ Ash Mtn Kwh Pwr Sta # 3, 8 -VI- 85, 2 ΨΨ 15 -VI- 85, R. D. Haines, 1 Ψ 17 -V- 86, 1Ψ 31 -V- 86, 1Ψ 9 -VI- 86, all R. D. Haines; 1 Ψ Yermo, 5 -VII- 39, W. M. Pearce, 1 ɗ Yosemite, 3800-4000 ', 1 -VI- 31, G. L. Smith; 2 ɗɗ Yosemite, 20, 23 -V- 21, Van Dyke; 1 ɗ, 1 Ψ 3880-4000 ' 14-15 -VI- 38, C. T. Sierra; 1 Ψ Yosemite Valley, 10 -VI-, 1 Ψ 25 -VI- 21, Van Dyke; Yuba Co., Sierra Foothill Field Stat., 3 mi N. Smartville, 7 -V- 80, 1ɗ J. Chemsak, 1 Ψ J. DeBenedictis. OREGON: 1 ɗ Ashland, 16 -IV- 10, P. D. Sergent; Benton Co., 1 Ψ 1 mi N. Suver, 1 -VIII- 66, F. M. & V. S. Beer; 1 ɗ, 1 Ψ Douglas Co., Cabin Creek Rest Stop, 1-5 -IX- 99, R. L. Westcott; Jackson Co., 1 Ψ 4 mi S. Ruch, 13 -VII- 68, R. L. Westcott; Josephine Co., Grants Pass, 2 ɗɗ, 2 ΨΨ 10 -V- 39, F. M. Beer, 1 Ψ 4 -VII- 69, R. L. Westcott; Polk Co., 1 Ψ 2 mi N. Suver, 1 -VIII- 49, F. M. Beer, 2 mi NW Suver, 1 ɗ 8 -VIII- 69, 1ɗ 16 - VI- 71, 1ɗ 8 -IX- 72, R. L. Westcott; 2 ɗɗ, 2 ΨΨ Rogue River, 16 -VI- 59, K. Goeden; Yamhill Co., 1 ɗ 6.7mi W. Carlton, 18 -VII- 68, R. L. Westcott. WA S H I N GTO N: 1 Ψ Asotin Co., Anatone, 3 -III- 47, F. Beer. MEXICO: BAJA CALIFORNIA: 1 Ψ N. Par Nac. Const. 1857, 1615m, 26 -V- 89, R. L. Westcott on Quercus. Paratypes deposited in following collections: AMNH, BKDC, BMNH, CASC, CLBC, DSVC, FMNH, FTHC, GHNC, GVMC, HAHC, LACM, LGBC, LBC, MCZC, MSUE, RDHC, RHTC, RLWE, RMBC, SGWC, TAMU, TCMC, UICM, USNM and UICM.
Hosts. Adult hosts are usually oaks, and specimens have been collected on: Quercus agrifolia Nee, Q, chrysolepis Liebm., Q. gravesii Sudw., Q. garryana Douglas, Arctostaphylos viscida Parry, Pyrus communis Linnaeus, Salix scouleriana Barratt ex Hook., and Betula spp. Larval host records from California based on adult emergence include: 1 ɗ Platanus racemosa Nutt., in Stanislaus Co., 14 -V- 80, F. M. Beer; 1 ɗ Quercus spp. 1974, L. Bezark; 1 Ψ Q. kelloggii Newb. (Q. california) by H. E. Burke from Placerville also same host; 3 ɗɗ reared from Q wislizenii A. DC, Placerville by J. J. Sullivan; 1 Ψ from Napa Co., 2 mi NNE Angwin by H. B. Leech; 1 ɗ Q. douglasii Hook. & Arn., at Guadalupe Creek, Santa Clara Co. on 10 -IV- 73 by C. Y. Kitayama; 1 Ψ Q. berberidifolia Liebm. {dumosa Nutt.}, from 2.5 mi N Pinyon Flats, Riverside Co., emerged 21 -VI- 62 (host collected 21 -VI- 61) G. H. Nelson. From Placerville, 1 ɗ reared from Prunus domestica Linnaeus by J. J. Sullivan and 2 ɗɗ were reared from Salix lasiolepis Benth., by H. E. Burke. One male was reared from black willow, Salix nigra Marsh. at Yuma, AZ. by C. L. Bellamy.
Comments. Males: 9.7 – 11.2 mm long, 4.3 – 5.0 mm wide. Females: 9.5 – 13.1 mm long, 3.9 – 5.8 mm wide.