(Figs. 34, 169–170, 371, Map 33)
Diagnosis.— Hypatopa manus possesses the following unique combination of features; an anteriorly directed ventroposterior margin of the gnathos; an apical process of the ventral part of the valva that is acutely curved inwardly from base; a digitate process of the dorsal part of the valva that is straight from a slightly angled base; a proximal flange that is elongate, narrowly angular basally, gradually widening dorsolaterally to a broadly rounded apical part; and a phallus and sclerite of phallus that is longer than the valva. This combination of character states differentiates it from all other congeners.
Description.—Head: Scales on vertex and frontoclypeus grayish brown tipped with pale grayish brown. Outer surface of labial palpus brown intermixed with pale-brown scales along apical margins of all segments, inner surface pale brown. Antennal scape with grayish-brown scales tipped with white, pecten grayish brown, flagellum gray. Proboscis pale brown.
Thorax: Tegula brown; mesonotum with basal 1 / 2 brown, apical 1 / 2 pale brown. Legs brown intermixed with pale-brown scales near midsegments and along apical margins of all segments and tarsomeres. Forewing (Fig. 371): Length 4.0–6.0 mm (n = 73), pale brown intermixed with brown scales; base dark gradually brightening to 2 / 3 [some specimens with a dark costa]; cell with three spots present or absent, if present, one near middle, one near middle on apical end along crossvein; marginal spots faint. Undersurface brown. Venation (Fig. 34) with M 3 and CuA 1 arising from a common point on distoposterior part of cell; cubital veins divergent from bases with M 3 straight and CuA 2 broadly curved. Hindwing: Translucent pale brown. Venation (Fig. 34) with cubitus 4 -branched; M 2 arising from point beyond distoposterior part of cell and branching point of M 3 and CuA 1 near 1 / 3.
Abdomen: Male genitalia (Figs. 169–170): Uncus narrowed from broadly rounded base, acutely downcurved and laterally flattened apically, sparsely setose, about equal in length to width of anal opening. Gnathos, narrow, anteriorly directed band, confluent with tegumen, ventroposterior margin widely emarginate mesially. Sockets of tergal setae extending beyond midlength of tegumen. Valva divided, ventral part basally protracted inwardly, angular at 2 / 3, narrowing to inwardly curved, spinelike apical process; process acutely curved inwardly from base, setose on outer surface, planate on inner surface; ventral margin entire, sparsely setose, with setose lobe at base of apical process; dorsal part with apical portion of costa extending dorsally from an angled base, forming setose digitate process; process straight from base; basal ridge of digitate process extending ventrally fusing with dorsolateral ridge of proximal flange; flange elongate, narrowly angular basally, gradually widening dorsolaterally to broadly rounded apical margin; flange densely setose, each seta pointed toward middle; margin entire. Juxta bandlike. Vinculum semicircular. Phallus and sclerite of phallus longer than valva; phallus slightly curved from 1 / 3, sclerite of phallus slightly curved at 2 / 3; anellus about 1 / 5 length of phallus, parallelsided along basal 1 / 2, gradually narrowing to rounded apex, setose on basal 1 / 2. Female Genitalia: Unknown.
Holotype, 3, “Est[ación] La Casona, 1520 m, Res[erva] Biol[ógica] Monteverde, Prov[incia] Puntarenas, COSTA RICA, Ago[sto] 1992, N. Obando, L-N- 253250, 449700, “ INBio: COSTA RICA: CRI 000, 947143 [barcode label], “ INBio, 3 Genitalia Slide by D. Adamski, No. 2566 [yellow label].
Paratypes (73 3): 10 3, same data as for holotype except, “ CRI 000, 947047, “Slide No. 2569; “ CRI 000, 947175, “Slide No. 2571, “ USNM 83936; “ CRI 000, 947068, “Slide No. 2570, “ USNM 83937; “ CRI 000, 947053, “Slide No. 2577, “ USNM 83938; “ CRI 000, 970311, “Slide No. 2578; “ CRI 000, 970252, “Slide No. 2580, “ USNM 83939; “ CRI 000, 947186, “Slide No. 2581; “ CRI 000, 947081, “Slide No. 2584; “ CRI 000, 947121, “Slide No. 2585, “ USNM 83940; “ CRI 000, 970249, “Slide No. 2587, “ USNM 83941; 1 3,”Set., “ CRI 000, 946857, “Slide No. 2573, “ USNM 83942; 1 3, “Feb., “ CRI 000, 801536, “Slide No. 2609; 1 3, “ Nov. 1991, “ CRI 000, 487012, “Slide No. 2607; 4 3, “ Ene. 1993, “ CRI 001, 368980, “Slide No. 2594, “ USNM 83943; “ CRI 001, 368990, “Slide No. 2596, “ USNM 83944; “ CRI 001, 369094, “Slide No. 2599; “ CRI 001, 369111, “Slide No. 2604, “ USNM 83945; 1 3, “C. Chaves, Abr. 1991, “ CRI 000, 434741, “Slide No. 2293; 6 3, “ Jun. 1991, “ CRI 000, 586070, “Slide No. 2256, “ USNM 83946; “ CRI 000, 623497, “Slide No. 2285; “ CRI 000, 623446, “Slide No. 2286, “ USNM 83947; “ CRI 000, 586021, “Slide No. 2236, “ USNM 83948; “ CRI 000, 585971, “Slide No. 2252; “ CRI 000, 613329, “Slide No. 2294, “ USNM 83949; 2 3, 23 Oct.- 9 Nov. 1990, “ CRI 000, 316243, “Slide No. 2195; “ CRI 000, 576662, “Slide No. 2198; 2 3, 25 Set.- 11 Oct. 1990, “ CRI 000, 577687, “Slide No. 2163; “ CRI 000, 577738, “Slide No. 2185, “ USNM 83950; 4 3, “ Set. 1991, “ CRI 000, 357222, “Slide No. 2223, “ USNM 83951; “ CRI 000, 357383, “Slide No. 2246; “ CRI 000, 357240, “Slide No. 2247, “ USNM 83952; “ CRI 000, 356652, “Slide No. 2218; 4 3, “C. Chaves & R. Espinoza, Nov.-Dic., 1990, “ CRI 000, 321920, “Slide No. 2158, “ USNM 83953; “ CRI 000, 321536, “Slide No. 2192, “ USNM 83954; “ CRI 000, 321953, “Slide No. 2203, “ USNM 83955; “ CRI 000, 321672, “Slide No. 2159, “ USNM 83956; 3 3, “F.A. Quesada, Jun., 1991, “ CRI 000, 338540, “Slide No. 2385; “ CRI 000, 613333, “Slide No. 2389, “ USNM 83957; “ CRI 000, 613355, “Slide No. 2390; 1 3, “M. Segura, 23 Mar.- 21 Apr., 1992, “ CRI 000, 416797, “Slide No. 2471; 1 3, “A. Martin, 21–29 May, 1992, “ CRI 000, 684384, “Slide No. 2311; 1 3, “D. Brenes, 21–29 May, “ CRI 000, 487953, “Slide No. 2321, “ USNM 83958; 1 3, “M. Ramirez, May, 1991, “ CRI 000, 355373, “Slide No. 2610; 2 3, “ Abr. 1994, N. Obando, # 2820, “ CRI 001, 781891, “Slide No. 2548; “ CRI 001, 781864, “Slide No. 2550, “ USNM 83959; 2 3, “Mar., 1994, # 2819, “ CRI 001, 764782, “Slide No. 2555, “ USNM 83960; “ CRI 001, 764636, “Slide No. 2562, “ USNM 83961; 3 3, “ Jul. 1993, # 2287, “ CRI 001, 130666, “Slide No. 2603; “ CRI 001, 730703, “Slide No. 2590, “ USNM 83962; “ CRI 001, 130649, “Slide No. 2592, “ USNM 83963; 3 3, “Agos. 1992, “ CRI 001, 910053, “Slide No. 2593, “ USNM 83964; “ CRI 001, 910170, “Slide No. 2600; “ CRI 001, 910010, “Slide No. 2602; 4 3, “Oct., 1993, “ CRI 001, 162892, “Slide No. 2630; “ CRI 001, 162899, “Slide No. 2626; “ CRI 001, 162842, “Slide No. 2629; “ CRI 001, 162856, “Slide No. 2631, “ USNM 83965; 1 3, “ May 1993, N.G. Obando, # 2184, “ CRI 001, 810549, “Slide No. 2591; 8 3, “Derrumbe, III Curso Parataxon., May, 1992, “ CRI 000, 416812, “Slide No. 2264, “ USNM 83966; “ CRI 000, 416849, “Slide No. 2266, “ USNM 83967; “ CRI 000, 426529, “Slide No. 2340; “ CRI 000, 417040, “Slide No. 2337; “ CRI 000, 417085, “Slide No. 2326; “ CRI 000, 416847, “Slide No. 2317, “ USNM 83968; “ CRI 000, 416799, “Slide No. 2274, “ USNM 83969; “ CRI 000, 416984, “Slide No. 2273, “Wing Slide No. 7010; 1 3, “II Curso Parataxon., Jun. 1990, “ CRI 000, 609183, “Slide No. 2169; 1 3, “San Luis, Monteverde, Prov. Punta., COSTA RICA, 1000–1350 m, Jul. 1994, Z. Fuentes, L-N- 250850, 449250, # 3074, “ CRI 002, 0 25719, “Slide No. 2634, “ USNM 83970; 1 3, “Estac. Mengo, 100 m, SW side Volcon Cacao, Prov. Guan., COSTA RICA, Feb. 1989, GNP Biodiversity Survey, 85 ° 28 ’ 10 ”W, 10 ° 55 ’ 43 ”N, “ CRI 001, 0 55023, “Slide No. 2522; 1 3, COSTA RICA, Prov. Puntarenas, El Ripario, 3 km NE de Progresso, 1300 m, 5–7 Jun. 1997, A. Picado, de Luz, L-S- 319000, 597000, # 47405, “ CRI 002, 569827, “Slide No. 2671; 3 3, “ COSTA RICA, Puntarenas, Monteverde, Eatación Biológica Monteverde, 1500 m, 11 -XI- 2001, Col. Kenji Nishida, luz en la noche, “Slide No. 2700, “ USNM 83971; “Slide No. 2708, “ USNM 83972; “Slide No. 2713, “ USNM 83973 [35 in INBio, 38 in USNM].
MAP 33. Distribution of Hypatopa manus (●) and H. caepae (˔).
Distribution (Map 33). Hypatopa manus is known from nine collecting sites; five in western Costa Rica along the Cordillera de Guanacaste, two on the Cordillera de Tilarán, and two closely adjacent sites on the eastern most part of the Cordillera de Talamanca near the border of Panama.
Etymology. The specific epithet manus is derived from the Latin meaning, the hand.