Figs 1, 2, 5, 7, 11, 100 – 110.
flavipes Fallén, 1819: 9 (Cordylura). Type-locality: “ad lacum Gyllebo Scaniae” (Sweden). frontalis Macquart, 1835: 387 (Cleigastra). Type-locality: “Du nord de la France ”. trilineata Meigen, 1838: 341 (Cordylura). Type-locality: not given.
minuta Becker, 1894: 116 (Amaurosoma). Type-locality: “Livland” (Latvia or Estonia). multisetosum Hackman, 1956: 16 (Amaurosoma). Type-locality: “N: Helsinge” (Nyland, Finland).
Remarks. Noted by Gorodkov (1970: 449; 1986: 18) for European part of Russia without indicating specific locality, by Šifner (2008: 153) from Kamchatka Kray and by Ozerov & Krivosheina (2014 b: 214) from Russian Far East.
Material examined. Altay: Lake Manzherok (51.825278 N 85.812778 E), 10.V. 2008, O. Kosterin (1 ♂, ZMUM); Arkhangelsk Oblast: Nenetskiy Rezerve, Сordon “Bol'shoy Gusinets” (68.1759 N 53.6453 E), 11.VII. 2008, A.L. Ozerov (1 ♀, ZMUM); Island Kashin (68.242042 N 53.856746 E), 10.VII. 2008, A.L. Ozerov (1 ♂, ZMUM); Chelyabinsk Oblast: 63 km NNW of Chelyabinsk (55.8000N 61.0333 E), 17–19.V. 1992, M. Krivosheina (1 ♀, ZMUM); Irkutsk Oblast: Novochunka (56.1144 N 99.2735 E), 5.VI. 1957, Gorodkov (1 ♂, ZISP); Os'kino (60.8116 N 107.9858 E), 12.VI. 1875, Chekanovsky (1 ♂, ZISP); Yurty (56.05 N 97.63 E), 21.V. 1912, Mishin & Verchov (1 ♂, 1 ♀, ZISP); Kamchatka Kray: Kluchi (56.3226 N 160.8295 E), 5.VI. 1909, A. Derzhavin (1 ♂, ZISP); Karelia: Reserve Kivach (ca. 62.2954 N 33.9214 E), 18–21.V.1989, 14– 21.VI. 1990, A. Polevoy (2 ♂♂, ZISP); Kizhi (62.0663 N 35.2377 E), 19.VI. 1979, Gorodkov (1 ♀, ZISP); Krasnodar Kray: Sochi, Estosadok env., Mt. Psekhako (43.6911 N 40.3666 E), 14–18.VI. 2008, K. Tomkovich (2 ♀♀, ZMUM); range Azish- Tau, Biological Station “Kamyshanova Polyana” env. (44.168784 N, 40.044638 E), 1240 m, 13.VI.– 6.VII. 2010, Kustov, (1 ♂, ZMUM); Chechtshir (48.2911 N 135.095 E), 18.V. 1973, Gorodkov (1 ♂, ZISP); Krasnoyarsk Kray: Turuchansk (65.7972 N 87.9586 E), 27.VI. 1967, Gorodkov (4 ♂♂, ZISP); Kursk Oblast: Streletskaya Steppe (51.5795 N 36.0870 E), 5–15.V. 2008, K. Tomkovich (2 ♂♂, 11 ♀♀, ZMUM); Borisovka (51.6705 N 35.4515 E), 18–25.IV. 1915, Sokolov (1 ♂, 1 ♀, ZISP); Leningrad Oblast and St.-Petersburg: Gatshina (59.56 N 30.13 E), 24.V. and 10.VI. 1940, A. Stackelberg (2 ♀♀, ZISP); Jukki (ca. 60.11 N 30.29 E), 5.VI. 1931, A. Stackelberg (1 ♂, ZISP); Luga (ca. 58.73 N 29.84 E), 27.V. 1956, A. Stackelberg (1 ♂, 1 ♀, ZISP); Peri (ca. 60.24 N 30.46 E), 15.VI. 1952, A. Stackelberg (2 ♀♀, ZISP); Pesochnaya (60.1175 N 30.1518 E), 30.V.1963, 22.V. 1966, K. Gorodkov (14 ♂♂, 7 ♀♀, ZISP); Island Bol'shoy Berezovyy (60.3 N 28.62 E), 29 and 31.V. 1981, Kandybina (2 ♂♂, 1 ♀, ZISP); Yashchera (58.8945 N 29.8206 E), 13 and 26.V.1957, 19.VI.1957, 23.VI.1964, 25.V.1965, 26.VI.1966, 24 and 26.V.1968, 1.VI. 1968, A. Stackelberg (3 ♂♂, 8 ♀♀, ZISP); Komarovo (60.18 N 29.81 E), 2.V. 1953, A. Stackelberg (1 ♂, ZISP); N. Bronnaya (ca. 60.19 N 29.64 E), 8.VI. 1918, A. Stackelberg (1 ♀, ZISP); Rozhdestveno (59.3236 N 29.9472 E), 20.VI. 1956, A. Stackelberg (2 ♀♀, ZISP); Moscow and Moscow Oblast: Izmaylovo (55.8028 N 37.8408 E, 55.7870 N 37.8345 E, 55.7871 N 37.8358 E, 55.7872 N 37.8365 E, 55.7923 N 37.8385 E, 55.7923 N 37.8385 E, 55.7867 N 37.8350 E, 55.7882 N 37.8384 E), 11.V.2007, 26.IV., 15–18.V.2008, 18– 31.V., 12. VI. 2009, A.L. Ozerov (17 ♂♂, 18 ♀♀, ZMUM); Petrovsko-Razumovskoe (55.8399 N 37.5694 E), 6.V. 1931, N. Violovich (1 ♂, ZMUM); Andreevskoe (55.9783 N 35.5885 E, 55.9815 N 35.5972 E), 13 and 14.V. 2007, A.L. Ozerov (2 ♂♂, 2 ♀♀, ZMUM); Burtsevo (55.9817 N 35.5982 E, 55.9799 N 35.5898 E, 55.9752 N 35.5862 E), 18 i 25.V., 2–3.VI. 2006, A.L. Ozerov (3 ♂♂, 4 ♀♀, ZMUM); Golitsyno (55.6496 N 37.0117 E), 21 and 29.V., 11.VI.1977, 8 and 20.V.1978, 9.V.1979, 29.V.1982, 25.V. 1990, A. Shatalkin (6 ♂♂, 13 ♀♀, ZMUM); Dmitrov env. (56.3163 N 37.7258 E), 19.V., 10.VI.2006, 7– 9.V., 15 and 19. V.2007, 30. IV., 9 and 10. V.2008, 27. V., 9. VI.2009, 7.V.2010, 15.V. 2011, D. Gavryushin, N. Vikhrev (17 ♂♂, 29 ♀♀, ZMUM); Ivanovskoe (55.9262 N 35.6198 E, 55.9338 N 35.6275 E), 14.V., 15.VI. 2006, A.L. Ozerov (5 ♂♂, 4 ♀♀, ZMUM); Naro-Fominsk (55.3643 N 36.7410 E, 55.3955 N 36.7823 E, 55.3729 N 36.7578 E, 55.3574 N 36.7361 E, 55.3930 N 36.77790 E, 55.3678 N 36.7381 E), 8–13.V.2006, 1.V.2008, 14 and 17.V., 26. VI.2009, 14.V., 22.VI. 2010, D. Gavryushin (7 ♂♂, 6 ♀♀, ZMUM); Ozhigovo (55.448 N 36.8709 E), 2.V. 2008, D. Gavryushin (1 ♂, 1 ♀, ZMUM); Zvenigorod env. (55.7003 N 36.7223 E), 24.V. 1981, A.L. Ozerov (1 ♀, ZMUM); River Ruza near Rozhdestveno (55.9322 N 35.6327 E), 5.V. 2011, A.L. Ozerov (1 ♂, ZMUM); Desna (55.5230 N 37.3720 E), 9.V. 1982, A.L. Ozerov (2 ♂♂, ZMUM); Stepan'kovo (55.0045 N 35.6247 E), 3.VI. 2007, A.L. Ozerov (4 ♀♀, ZMUM); Elektrogorsk env. (55.9124 N 38.8411 E), 15.V. 2010, A.L. Ozerov (1 ♂, ZMUM); Yakshino (55.9198 N 35.5833 E), 2.V. 2008, A.L. Ozerov (1 ♂, 2 ♀♀, ZMUM); Novgorod Oblast: Vereb'e (58.6782 N 32.6965 E), 10.V. 1975, Gorodkov (2 ♂♂, ZISP); Novosibirsk Oblast: (55.52 N 83.24 E), 22.V. 2011, O. Kosterin (1 ♂, 1 ♀, ZMUM); Zherebtsovo env. (55.125833 N 83.256111 E), 31.V. 2008, O. Kosterin (15 ♀♀, ZMUM); 4 km S of Novososedovo (54.6169 N 83.9850 E), 7.VII. 2008, O. Kosterin (1 ♀, ZMUM); Sakhalin Oblast: Yuzho-Sakhalinsk (46.95 N 142.74 E), 27.V. 1954, Violovich (2 ♂♂, ZISP); Stavropol' Kray: Teberda Reserve (43.4472 N 41.744 E), 12.V. 1964, Gorodkov (1 ♂, ZISP); Tatarstan: Reserve Volzhsko-Kamskiy, Lake Raifa (55.8973 N 48.733 E), 5–15.V. 2005, Basov (6 ♂♂, 21 ♀♀, ZMUM); Tula Oblast: near Novomoskovsk (55.0352 N 38.1547 E), 13. VI. 2009, N. Vikhrev (1 ♀, ZMUM); Tyumen' Oblast: (63.818 N 59.562 E), 5–11.VII. 2010, K. Tomkovich (1 ♂, 2 ♀♀, ZMUM); Neroyka (ca. 64.57 N 59.67 E), 700 m, 24–27.VI.1990, 6– 9.VII. 1990, Malozemov (4 ♂♂, ZISP); same place, 6– 12.VII. 1990, Malozemov (2 ♂♂, ZISP); Yakutia: River mouth of Viluy (63.0406 N 112.3146 E), 8–16.VI. 1875, Chekanovsky (1 ♀, ZISP); Yaroslavl' Oblast: Berditsino (57.45 N 40.10 E), 8, 9 and 29.V. 1907, A.I. Yakovlev (2 ♂♂, 2 ♀♀, ZISP).
Additional material examined. China: prov. Sichuan, Tatsienlu (=Kangting) (29.9995 N 101.955 E), 21.V 1893, Potanin (1 ♂, ZISP).
Diagnostic description. Body-length 3.8 –6.0 mm. Head. Frons from completely yellow to black, only lower quarter always yellow. Female frons usually yellow completely, while male frons black in upper part. 3 orbital setae present. Antenna black. Postpedicel with acutely angled upper apical corner, approximately 2 times as long as wide, in male wider than in female. Thorax black, grey dusted. Proepisternum with hairs in central part. Katepisternum with 3 strong setae, from black to yellow. Legs yellow, only mid and hind coxae and tarsi darkened. Fore femur usually with dark stripe posterodorsally (Fig. 102), with 7–17 anteroventral setae in 2–3 irregular rows (Figs 100, 101). Fore tibia with 1 anterodorsal, 1 posterodorsal and 1 posterior setae at middle. Hind tibia with apical posteroventral and anteroventral setae, posteroventral one is equal to anteroventral or 1 / 2 as long as anteroventral. Wing tinged with brownish, sometimes strongly darkened along anterior margin. Abdomen black, greyish dusted, covered with pale hairs in male and black short hairs in female. Male sternite 4 rectangular, approximately 1.5 times as long as wide (Fig. 109). Male sternite 5 with black lobes, as in Fig. 106. Epandrium, cerci and surstyli as in Figs 107, 108. Female sternite 8 from single plate (Fig. 103, 105) to two separate sclerites (Fig. 104).
Distribution. Russia (Fig. 110): Altay, Amur Oblast, Arkhangelsk Oblast, Chelyabinsk Oblast, Irkutsk Oblast, Kamchatka Kray, Karelia, Krasnodar Kray, Krasnoyarsk Kray, Kursk Oblast, Leningrad Oblast and St.- Petersburg, Magadan Oblast, Moscow and Moscow Oblast, Novgorod Oblast, Novosibirsk Oblast, Primorskiy Kray, Sakhalin Oblast, Stavropol' Kray, Tatarstan, Tula Oblast, Tyumen' Oblast, Yakutia, Yaroslavl' Oblast.— Europe (widespread), China (first record), Mongolia.