Hebrus ruficeps Thomson, 1871

(Figs. 44–45, 73)

Hebrus ruficeps Thomson, 1871: 395. Syntype(s): sex not indicated, Sweden, Skåne, Ringsjön (MZLU). Hebrus ruficeps var. transversalis Rey, 1893: 109. Syntype(s): Carpathian Mts. (probably lost).

Material examined. BULGARIA: Sofia Province: Bajana [= Bayana], 17. vii.1956, 1 ♀ (micropterous), L. Hoberlandt lgt., P. Kment det. (NMPC). CZECH REPUBLIC: Bohemia: Filipovka (5056), peatbog, 26. iv.2002, 1 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Česká Kamenice, Noldenteich NM (5152; 50°48′43.3″N 14°26′15.0″E), peatbog, 2. viii.2004, 3 Ƌ 3 ♀ (micropterous), 23. viii.2004, 2 ♀ (micropterous), 3. x.2004, 2 Ƌ 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Heřmanice (5156), flooded basalt quarry, 26. iv.2002, 2 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Mařeničky, Rašeliniště Mařeničky NM (5254; 50°47′41.2″N 14°40′27.8″E), peatbog, 29. vi.2004, 2 ♀ (micropterous), 1. x.2004, 1 Ƌ 3 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Provodín, sand-pit NW of the village (5353), 8. v.2002, 1 ♀ (micropterous), M. Fikáček & H. Poláček lgt., P. Kment det. (NMPC); Karlsbad, Zinnbach [= Lužní potok near Karlovy Vary (56–5739)], 9. ix.1937, 1 ♀ (micropterous), W. Reinhard lgt., P. Kment det. (NMPC); Lomnice (5742), experimental reservoirs on recultivated spoil tips 2 km NE of the village, 20. iv.2002, 1 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Karlsbad, b. Flugplatz, Waldrand bei Espenthor [= Karlovy Vary, near airport, forest margin near Olšová Vrata] (5743), 1. xi.1938, 1 Ƌ (micropterous), L. Hoberlandt det. (NMPC); Neudau [= Nejda] (5743), 24. x.1937, 1 Ƌ 2 ♀ (micropterous), P. Kment det. (NMPC); Otovice u Karlových Varů (5743), 5 ♀ (micropterous), P. Kment det. (NMPC); Račice—Předonín (5552), 22. iv.1976, 2 ♀ (micropterous), I. Kovář & J. Ježek lgt., P. Kment det. (NMPC); Rožďalovice (5656–57), 19. v.1930, 1 Ƌ (micropterous), Dr. Klička lgt., L. Hoberlandt det. (NMPC); Kateřina, Soos NNR (5840), 27. v.1987, 3 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Čelákovice (5854), ix.1946, 1 ♀ (micropterous), Mat lgt., J. L. Stehlík det. (MMBC); Sedlčánky, oxbow-lakes on left bank of Labe river (5854; 50°10′05″N 14°47′53.1″E), 27. v.2003, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Přerov [nad Labem] (5854–55), 5. v.1930, 1 ♀ (micropterous), Dr. Klička lgt., P. Kment det. (NMPC); Prameny, Mokřady pod Vlčkem NR (5942), vegetated pool in spring fen, 750–760 m a.s.l., 13. ix.2011, 1 Ƌ 1 ♀ (micropterous), J. Škodová lgt., P. Kment det. (NMPC); [Praha], Krč (5952), 1 Ƌ 1 ♀ (micropterous), Zeman lgt., L. Hoberlandt det. (NMPC); Horní Ředice (5961b; 50°05.508′N 15°57.064′E), NR Žernov, marsh at Mordýř pond, 20. iv.2007, 1 ♀ (micropterous), M. Boukal lgt., P. Kment det. (NMPC); Benešov (6254), 31. vii.1973, 1 ♀ (micropterous), Z. Jindra lgt., P. Kment det. (VURV); Lnáře (6548), Malý Pálenec [pond?], iv.1942, Mařan lgt., L. Hoberlandt det. (NMPC); Jindřichův Hradec (6855–56), 18. vii.1885, 1 ♀ (micropterous), coll. L. Duda, L. Hoberlandt det. (NMPC); Horusice, NPR Ruda (6854; 49°09′03.9″N 14°41′27.7″E), peatbog, 416 m a.s.l., 18. vii.1970, 2 Ƌ 1 ♀ (micropterous), 18. vii.1971, 1 ♀ (micropterous), L. Škapec lgt., P. Kment det.; 29. iv.1975, 7 Ƌ 4 ♀ (micropterous), P. Štys lgt., J. Vilímová det.; 18. vii.1976, 1 Ƌ 1 ♀ (micropterous), L. Škapec lgt., P. Kment det. (all NMPC); 13. ix.2010, 2 Ƌ 5 ♀ (micropterous), extracted from wet Sphagnum, P. Kment lgt. & det. (NMPC); viii.2011, 1 Ƌ 1 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); 29. viii.2014, 5 Ƌ 3 ♀ (micropterous), extracted from wet Sphagnum, J. Vilímová lgt., P. Kment det. (NMPC); Ponědrážka env., W bank of Švarcenberk pond (6854; 49°08.72′N 14°42.03″E), 27. vii.2014, 1 Ƌ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Lomnice nad Lužnicí (6954), 16. vi.1992, 1 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Stožec, Mrtvý Luh (7048; 48°51′34″N 13°53′35″E), peatbog, 740 m a.s.l., extracted from wet Sphagnum, 18. viii.2015, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); České Budějovice, Čtyři Dvory, small peatbog near U hada street (7052; 48°58′39.1″N 14°25′33.3″E), 11. vii.2000, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Nová Pec, Pěkná env. (7149; 48°50′49″N 13°55′27″E), Vltavský luh, oxbow lake of Vltava river, 18. viii.2015, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC). Moravia: Rychlebské hory Mts., Javorník env., Uhelná (5668), 15. vii.1981, 1 Ƌ 2 ♀ (micropterous), Z. Jindra lgt. & det. (ZJPC); Rejvíz, Rejvíz NNR (5769; 50°13′15.5″N 17°17′58.5″E), alluvial peaty meadow, 736 m a.s.l., in wet Sphagnum, 15. viii.2011, 3 Ƌ 3 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Ostrava, Martinov, Štěpán pond (6175), shore, sifting, 22. ix.1998, 1 Ƌ (micropterous), 12. ii.2000, 1 Ƌ (micropterous), M. Mantič lgt., P. Kment det. (MMHC); Křižánky, Kyšperský rybník pond (6362; 49°41′10.1″N 16°03′32.8″E), waterlogged shore, 24. vi.2000, 1 Ƌ (micropterous), P. Kment lgt. & det. (NMPC); Rudice, Rudická jezírka (6666, 49°20′09.7″N 16°43′24.4″E), flooded pits after iron ore mining, 1. v.1999, 2 Ƌ 1 ♀ (micropterous), 6. vi.1999, 2 Ƌ 2 ♀ (micropterous), 6. vii.1999, 2 ♀ (micropterous), P. Kment lgt. & det. (NMPC, MHBC, MUBC); Velká Bíteš, Dolní Klečanský rybník pond (6763; 49°16′39″N 16°14′28″E), waterlogged grassy shore, 1. v.2000, 3 Ƌ 10 ♀ (micropterous), P. Kment lgt. & det. (NMPC, MUBC); Střelice (6865), ix.1943, 1 ♀ (micropterous), Rauscher lgt., L. Hoberlandt det. (MMBC); Pouzdřany (7065), 7. vi.1967, 1 ♀ (macropterous), Volák lgt., P. Kment det. (SMOC); Šafov env., Janský rybník pond (7160), litter on banks (sifting) + in water (high water level), 10. iv.2007, 5 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Hrabětice, Travní dvůr SAC, Za drahou pool (7164; 48°48′55.9″N 16°27′8.8″E), 15. v.2011, 1 Ƌ (micropterous), J. Sychra lgt., P. Kment det. (NMPC); Lednice, Nejdek (7166), Nejdecké louky meadows, Bažina u Azontu reed marsh, 18. ix.1998, 2 ♀ (micropterous), P. Kment & M. Horsák lgt., P. Kment det. (NMPC); Podivín, pools by road to Lednice (7167; 48°49′24″N 16°50′41″E), 26. iv.2003, 2 Ƌ 1 ♀ (micropterous), M. Horsák lgt., P. Kment det. (NMPC); Valtice, Alach IV (7266), small silting pond in forest, 9. v.1998, 1 ♀ (macropterous), P. Kment lgt. & det. (NMPC). POLAND: Lesser Poland Voivodeship: Chochołów, sepage on south margin of village, 14. vi.2002, 1 ♀ (micropterous), M. Horsák lgt., P. Kment det. (NMPC). ROMANIA: Caraș-Severin county: Caransebeș, litter on pond banks (45.4188°N 22.1875°E), 217 m a.s.l., 24. xii.2014, 1 ♀ (micropterous), G. M. Berchi lgt. & det. (GBCR). Cluj county: Turda, litter on pond banks (46.47388°N 23.77694°E), 317 m a.s.l., 22. xii.2014, 1 ♀ (macropterous) 2 Ƌ (micropterous), G. M. Berchi lgt. & det (GBCR). Timiș county: Dumbrăvița, litter on pond banks (45.80638°N 21.26416°E), 96 m a.s.l., 19. xii.2014, 1 Ƌ 1 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR); Jimbolia, litter on pond banks (45.8099°N 20.7177°E), 75 m a.s.l., 26. xii.2014, 2 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR); Satchinez, collected by pitfall trap in farmland (45.9622°N 21.0466°E), 92 m a.s.l., 10. vii.2012, 1 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR). RUSSIA: North European Territory: Leningrad Province: Karelian Isthmus, St. Petersburg reg., Svetloe ozero Lake (ca. 60°18′N 29°50′E), surrounding peat bog, 21. vii.2002, 2 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Karelian Isthmus, 4 km S of Dibuny railway station, river bank, 13. vi.2004, 1 ♀ (macropterous) 9 Ƌ 5 ♀ (micropterous), P. Kment lgt. & det. (NMPC). SLOVAKIA: Svätý Jur, Šúr NNR (7768), 4. vi.1941, 1 Ƌ 3 ♀ (micropterous), L. Korbel lgt., P. Kment det. (MMBC); Palárikovo (7974), salty clay-pit, in water, 1. v.1999, 1 ♀ (micropterous), M. Mantič lgt., P. Kment det. (MMHC); Tvrdošovce (7974), meadow behind railway, canal, 1. v.1999, 1 ♀ (micropterous), M. Mantič lgt., P. Kment det. (MMHC).

Distribution (Fig. 73). Europe: Albania (Josifov 1970), Austria (Benedek 1970, Rabitsch 2005), Belgium (Stoffelen et al. 2013), Bulgaria (Josifov 1964), Byelorussia (Lukashuk 1997, Kanyukova 2006), Croatia (Protić 1998, Kment & Beran 2011), Czech Republic (Hoberlandt 1977), Denmark (Andersen & Gaun 1974), Estonia (Coulianos 2005), Finland (Rintala & Rinne 2010), France (Poisson 1957a), Germany (Hoffmann & Melber 2003, Wachmann et al. 2006, Strauss & Niedringhaus 2014), Great Britain (Savage 1989), Hungary (Horváth 1897, Benedek 1970, Boda et al. 2015), Ireland (Savage 1989), Italy (Servadei 1967, Tamanini 1979, Cianferoni & Mazza 2012), Latvia (Lukashuk 1997, Spuņģis 2008), Luxembourg (Reichling & Gerend 1994), Macedonia (Protić 1998), Moldavia (Derzhansky 1984, 1997; Kanyukova 2006), the Netherlands (Aukema et al. 2002), Norway (Coulianos 1998, Coulianos et al. 2008), Poland (Jaczewski & Wróblewski 1976, Wróblewski 1980), Romania (Horváth 1897, Benedek 1970), Russia: North, Central and South European Territory of Russia (Kanyukova 1997, 2006), Serbia (Horváth 1897, Benedek 1970, Protić 1998), Slovakia (Horváth 1897, Benedek 1970, Kment et al. 2003, Reduciendo Klementová et al. 2015), Slovenia (Gogala 2003), Sweden (Coulianos & Ossiannilsson 1976), Switzerland (Dethier & Matthey 1977, Matthey & Fiora 1979), Ukraine (Putshkov & Putshkov 1996; Kanyukova 1997, 2006; Grandova 2013). Asia: Iran: Mazandaran (Ghahari et al. 2013), Kazakhstan: Asian part (Andersen 1995; Kanyukova 1997, 2006; Esenbekova et al. 2015), Russia: East Siberia, Far East (Kanyukova 1997, 2006; Vinokurov et al. 2010), Turkey: Asian part (Fent et al. 2011), Uzbekistan (Kanyukova 1997, 2006).

The species was listed from Bosnia Herzegovina by Andersen (1995), probably based on Lindberg’s (1922) record from the locality Ilidze, which probably is a misidentification of H. fulvinervis recorded from the same locality by Benedek (1970).