Echinocardium cordatum (Pennant, 1777)

Fig. 69 A–E.

Amphidotus cordatus: Forbes, 1841: 190 –193.

Echinocardium cordatum: A. Agassiz, 1872: 109; 349. Pl. XIX, Figs 10–17. Pl. XX, Figs 5–7; Döderlein, 1910: 248; H.L. Clark, 1923 *: 405. Pl. XIII; H.L. Clark, 1925a: 232 –233; Mortensen, 1951: 152 –157. Pl. XVIII, Figs 6, 7. (1–3, 8); Clark & Courtman-Stock, 1976: 251 –252; Schultz, 2010: 411 –412, Figs 772–774; Branch et al., 2010: 236, Fig. 106.5. Echinocardium australe. A. Agassiz, 1872 –74: 109; 580. Pl. XXXVII, Fig. 15; Bell, 1904: 174.

Material examined. MBC-A077941; MBC-A 022287; MBC-A 022288; MBC-A 022289; MBC-A 022293; MBC- A023309; MBC-A023402; MBC-A027879; MBC-A077955; MBC-A077956; MBC-A077959; MBC-A077963; MBC-A077961; MBC-A077962; MBC-A077958; MBC-A077957; MBC-A077960; MBC-A077954; A27739; A6457; MBC-A077965; MBC-A077966; MBC-A077967; MBC-A077968; MBC-A077969; MBC-A077970; MBC-A077971; MBC-A078006; A 22354; A 22355; MBC-A 022287; MBC-A 022288; MBC-A 022289; MBC- A 022293.

Identification. Test moderately high, with deep anterior notch. Apical disc posteriorly positioned. Anterior ambulacrum distinctively sunken, two-series of irregularly arranged, large pores, associated with tube feet; paired petals sunken, wide at internal fasciole, tapering distally. Periproct opening at truncated end, variable in shape. Internal fasciole conspicuously long, subanal fasciole shield-shaped, with four pore-pairs, pointed at end; anal fasciole extending along periproct, onto upper side, separated from subanal fasciole. Spines uniform. Live specimen brownish, denuded test white.

Global maximum size. Maximum test length 90 mm.

Global distribution. From northern Norway along European coasts into Mediterranean and Morocco, South Africa, Japan, New Zealand and southern Australia; from littoral to 500 m (Döderlein 1910; Schultz 2010).

Remarks. Samples identified through current study extend previously known distribution northwards towards Sodwana Bay. Differs from E. capense in large shield shape internal fasciole and frontal notch.