Records associated with mangroves: Pinnacle Rock, Isla Bartolomé, Galápagos, moist mangrove leaf litter (Laguncularia racemosa); Punta Cormorán, Flamingo Lagoon, Isla Floreana, Galápagos, decayed mangrove leaf litter (Laguncularia racemosa); Darwin Bay and Lake Arcturus, Isla Genovesa, Galápagos, moist decayed leaf litter and black humus under Rhizophora mangle; Flamingo Bay, Isla Rábida, Galápagos, moist decayed leaf litter under Rhizophora mangle; South of Wreck Bay, Isla San Cristobal, Galápagos, moist leaf litter, humus and rotten wood under Conocarpus erecta, Laguncularia racemosa and Avicennia germinans; Puerto Ayora, Divine’s Bay and west of Canal de Itabaca, Isla Santa Cruz, Galápagos, moist decayed leaf litter under Avicennia germinans and Rhizophora mangle mangroves (Schatz 2003).
Distribution: Galápagos.
Remarks: Although this species occurs predominantly in the littoral mangrove habitat, it can also occur under other deciduous trees and in deeper humus layers. There are also a few remarkable records from a saltwater Crater Lake, old giant tortoise droppings and epiphytic lichens and moss in a highland cloud forest (Schatz 2003).