(Fig. 12)
Previously reported from northwestern Mexico and Puerto Rico to Costa Rica (Guanacaste, Alajuela, and San José provinces) (Champion 1898; Torre-Bueno 1906; Wolcott 1924, 1936; Drake & Harris 1928c, 1934, 1938; Kuitert 1942; Drake & Maldonado-Capriles 1954; Maier 1977; Maes 1989; Bass 1993; Morales & Molano 2009; Damgaard et al. 2014), being herein recorded for the first time from Puntarenas Province. In Costa Rica, T. quadrilineatus is found in the same type of small forest puddles along the Atlantic and Pacific slopes as is T. opacus, also up to 1,200 m a.s.l.
Material examined. Alajuela— El Calabazo Stream, Fundación Madreverde, Palmares, 1200 m a.s.l., (B. Pacheco): 1 female (MZUCR). Bosquecito Stream, Sede de Occidente, UCR, X 483,539, Y Y 230,010. 4, 1074 m a.s.l, 15.XI.2008, (B. Pacheco): 1 female (MZUCR). Guanacaste — Papagayo Substation, stream, 25.XI.2005: 1 female, 2 nymphs (MZUCR). 3 km NW from Nacaome, 100 m a.s.l, Barra Honda National Park, X 239,000, Y 386,000, III.1993, (M. Reyes): 4 males, 2 females (InBio). 3 km NW from Nacaome, Barra Honda National Park, X 239,000, Y 386,000, 100 m a.s.l., VIII.1993, (M. Reyes): 5 males, 2 females (InBio Cat. # 2317). Las Pailas Station, Rincon de la Vieja National Park, A. C. Guanacaste, X 306,300, Y 388,600, 800 m a.s.l., 18.XII.1993, (F. A. Quesada): 2 males, 1 female (InBio Cat. # 2541). Murciélago Station, 8 km SW from Cuajiniquil, X 320,300, Y 347,200, 100 m a.s.l, VIII.1993, (F. Quesada): 3 males, 2 females (InBio). Murciélago Station, 8 km SW from Cuajiniquil, X 320,300, Y 347,200, 100 m a.s.l, IV.1994, (F. Quesada): 1 male, 1 female (InBio Cat. #2809). Murciélago Station, 8 km SW from Cuajiniquil, Guanacaste National Park, X 320,300, Y 347,200, 100 m a.s.l, I.1994, (F. A. Quesada): 1 male, 2 females (InBio Cat. #2556). Murciélago Station, Guanacaste National Park, X 320,300, Y 347,200, 100 m.a.s.l. X–XI.1994, (F. A. Quesada): 1 female (InBio Cat. #3328). Santa Rosa Station, 300 m a.s.l, X 313,000, Y 359,800, I.1990 (GNP Biodiver. Survey): 2 females (InBio). Santa Rosa National Park, Naranjo Beach, X 309,300, Y 353,300, IV.1991, (E. Alcazar): 1 male, 1 female (InBio). Naranjo Beach, Santa Rosa, Guanacaste National Park, X 309,300, Y 353,300, II.1991, (E. Alcazar): 8 males, 9 females (InBio). Naranjo Beach, Santa. Rosa, Guanacaste National Park, X 309,300, Y 353,300, III.1991, (E. Alcazar): 2 males, 2 females (InBio). Santa Rosa National Park, 300 m a.s.l., XII. 1981, (D.H. Janzen & W. Hallwachs): 2 females (InBio). Santa María Sector, 6 km S from Santa María Volcano, X 304,8000, Y 393,700, 750– 790 m a.s.l III.1996, (F. A. Quesada, D. Briceño & A. Masis): 1 male, 1 female (InBio Cat., #7335). Puntarenas— Puntarenas, San Lucas Island, Tumbabote Beach, X 213,800, Y 435,700, 0–10 m a.s.l, 11. III.2005, (Y. Cardenas): 5 males, 5 females (InBio Cat. #79797). Montezuma, IV.1995, (C. Nolthing): 1 male, 1 female (MZUCR). Barranquilla Stream, Jabonal, Esparza, Site 6, 19.VIII.2008, (B. Pacheco): 1 male (MZUCR). San José— Acosta, Palmichal, Tabarcia and Negro Rivers, N 0 9.8333333, W -84.2, 1200 m a.s.l, 25.II.2008, (B. Pacheco): 2 males, 1 female, 1 nymph (MZUCR). Puriscal, Mercedes Sur, 1175 m a.s.l, 22.XI.1998, (C. Obando): 1 male (MZUCR).