Southern seas between 35°S and 66°S, including the surroundings of Marion Island and cold waters south of Australia (Spence Bate 1888; Illig 1914; Hale 1941; Iwasaki & Nemoto 1987a; Wasmer 1993; Tiefenbacher 1991, 1994; Vereshchaka & Lunina 2015); bathypelagic.