Anacochylidia Brown 2019, new genus

Anacochylidia Brown, new genus

Type species: Anacochylidia maderana Brown, new species, by monotypy.

Diagnosis. The short broad valvae and the long, slender secondary process of the phallus of Anacochylidia (Fig. 19) are most similar to those of Cochylidia Obraztsov, 1956, but other features of the male genitalia deviate from those of all known Cochylidia. In Anacochylidia the top of the tegumen is broad with a shallow, rounded concavity; the costa of the valva is short, unmodified, and upcurved, without a basal process; and the phallus has an elongate secondary process ca. 0.5 times as long as the phallus itself. In contrast, the male genitalia of Cochylidia are characterized by a dorsally attenuate tegumen, often with a narrowly bifurcate process at the top; the base of the costa of the valva has a long, erect, somewhat digitate, setose process; and the phallus has a short, curved digitate process apically. The latter is probably homologous with the extraordinarily long process of the phallus of Anacochylidia.

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna about 0.9 times flagellomere diameter in male, shorter, sparser in female; labial palpus porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus inconspicuous. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing (Fig. 1) length 4.5–4.8 mm, length ca. 2.8 times width; costa slightly and evenly arched throughout, without costal fold in male, apex rounded; termen obliquely angled toward hind margin; Sc ca. 0.45 times wing length; R 1 originating from about middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa well before apex; M 3 and CuA 1 separate; CuA 2 originating about 0.75 times length of discal cell; CuP absent; A1+2 stalked in distal 0.65. Hindwing length ca. 2.5 times width; costa weakly undulate, male with narrow costal roll in basal 0.5 of wing; apex rounded; termen slightly concave near middle; Sc+R 1 ca. 0.25 times wing length, Rs and M 1 stalked ca. 0.3 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.6 length of discal cell; frenulum with one acanthus in male, two acanthi in female. Abdomen: Unmodified. Male genitalia (Fig. 19) with tegumen short, broad, with shallow, rounded excavation at top bordered laterally by short lobelike processes; subscaphium subrectangular, semi-sclerotized; socius inconspicuous; transtilla well developed, broad, weakly bilobed with small rounded median lobe; valva short, broad, subquadrate, pointed apically, outer margin undulate, length ca. 0.4 times width; costa sclerotized, curved dorsally; sacculus represented by narrow, curved, sclerotized edge, lacking free apical process; vinculum arms weakly joined by membrane distally. Phallus elongate, ca. 4.0 times length of valva, strongly curved in basal 0.4, with slender curved spine originating dorsally just beyond curve, extending ca. 0.5 length of phallus, vesica with dense bundle of 7–8 small, non-deciduous cornuti. Female genitalia (Fig. 33) with ovipositor long, slender, telescopic; papillae anales slender, slightly pointed posteriorly; apophyses posteriores ca. 2.0 times as long as apophyses anteriores; sterigma mostly membranous, lamella postvaginalis a subquadrate plate, attenuate and distinctly emarginate posteriorly, with a pair of narrow, submedial sclerites extending anteriorly from plate; colliculum sclerotized with longitudinal wrinkles, occupying entire ductus bursae; corpus bursae rounded, differentiated from ductus bursae by absence of sclerotization, signum lacking; origin of accessory bursa uncertain.

Etymology. The name comes from the similar genus Cochylidia and the Greek “ana” (= against) and is feminine in gender.