Manerebia pauperata Pyrcz & Willmott, 2006, n. stat.

Manerebia satura pauperata Pyrcz & Willmott, 2006: 47. Type locality: Km 40, road Loja – Zamora, Zamora-Chinchipe, Ecuador. Holotype male: CEPUJ [examined].

Remarks. In both BI (Fig.12) and ML (Fig. 13) trees M. pauperata n. stat. and M. satura situate in distant branches of the tree and cluster in highly resolved clades with other congeners. Both M. lamasi n. stat., and M. pauperata n. stat. have also been collected in sympatry in the Cordillera del Cóndor in southern Ecuador. Additionally, the two taxa also consistently differ in wing colour patterns (Fig. 5C, D) and male genitalia. We therefore formally recognize M. lamasi n. stat. as a distinct species. Despite an overall similar colour pattern in comparison with M. satura from central and southern Peru, with the exception of the absence of the HWV yellow median band, M. pauperata n. stat. shows some genitalic differences. Moreover, the species-delimitation methods (Figs. 14, 15) and genetic distances (Supplementary material 2) clearly support treating the two taxa as separate species.