Emended diagnosis. CuA with four posterior branches; most basal posterior branch of CuA basal to fork of M into its two main branches; no arculus (oblique crossvein in area between M and CuA); the veinlets in costal area long and oblique (angles between veinlets and main stem of ScP ranging between 20°–30°).
Locality and horizon. Holotype PE 2020/5002- LS a, b (part and counterpart): south of the village of Sitters, Saar-Nahe Basin, Germany; Niedermoschel black shale, Jeckenbach Subformation, Meisenheim Formation, Lower Rotliegend, probably Asselian–?Sakmarian, early Permian. Probably specimen NM 790 (basal fourth of a forewing, private collection of H. Stapf / Nierstein); also from the Niedermoschel black shale, east of the village of Niedermoschel, stored at the Paläontologisches Museum Nierstein, Germany.