Speocera amber Li & Li sp. nov. (Figs 1–2, 55A)

Material examined. Holotype ♂ (IZCAS), Madagascar, Antsiranana Province, Antsiranana District, Amber Mountain (12º31.796′S, 49º10.460′E), elev. 1084 m, 14.XII.2016, leg. H. Zhao. Paratype 1♀ (IZCAS), same data as holotype.

Etymology. The specific name is derived from the type locality; noun.

Diagnosis. The new species can be distinguished from all other congeners by the very long, ribbon-like embolus of the males (Figs 1A–B) and the spiral spermathecae of the females (Fig. 2A).

Description. Male. Total length 0.95; carapace length 0.45; carapace width 0.43; abdomen length 0.47; abdomen width 0.34. Prosoma: carapace pyriform, narrowing gradually anteriorly; light yellow, suffused with purplish pattern (Fig. 2C); pars cephalica flat; fovea not visible. Sternum light yellow, suffused with purplish color, longer than wide. Endites light yellow, suffused with purplish tinge; with serrula; labium light yellow, suffused with purplish tinge. Clypeus nearly vertical; short. Chelicerae light yellow; anterolateral margin with a cluster of short setae; promargin with a series of six teeth reaching the lamina; retromargin with two small denticles (Fig. 55A). Eyes: six contiguous eyes surrounded by black pigmentation, PME smallest, PLE largest (Fig. 2C). Opisthosoma oval; purplish. Legs light yellow; leg lengths I 2.45 (1.06, 0.16, 0.58, 0.39, 0.27); II 1.67 (0.46, 0.11, 0.50, 0.34, 0.26); III 1.47 (0.41, 0.13, 0.38, 0.33, 0.23); IV missing. Genitalia (Fig. 1). Palpal femur long; palpal tibia about 1.3 times as long as cymbium (Figs 1A–B). Cymbium conical, without prolateral extension (Fig. 1A). Bulb spherical. Embolus very long, ribbon-like, nearly transparent, associated sclerite absent; sperm duct initially medium size, gradually tapering distally (Fig. 1D).

Female. Total length 1.20; carapace length 0.51; carapace width 0.43; abdomen length 0.64; abdomen width 0.50. Similar to male in color and general features (Figs 2D–E) but larger. Chelicerae unmodified. Leg lengths I missing; II 1.92 (0.53, 0.15, 0.57, 0.39, 0.28); III 1.50 (0.41, 0.14, 0.38, 0.33, 0.25); IV 1.98 (0.55, 0.14, 0.56, 0.43, 0.30). Genitalia. Internal genitalia with afferent spermathecae short, straight, not extending past the sides of the abdomen (Fig. 2B); spermathecae spiral (Fig. 2A).

Distribution. Known only from the type locality.