Published June 30, 2010
| Version v1
Journal article
Restricted
Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the Late Palaeocene of Itaboraí (Brazil) based on teeth and petrosal bones
Authors/Creators
Description
Ladevèze, Sandrine, Muizon, Christian De (2010): Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the Late Palaeocene of Itaboraí (Brazil) based on teeth and petrosal bones. Zoological Journal of the Linnean Society 159 (3): 746-784, DOI: 10.1111/j.1096-3642.2009.00577.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2009.00577.x
Files
Linked records
Additional details
Identifiers
- LSID
- urn:lsid:plazi.org:pub:1729AA44FFC8FFE7B62CFFA9F70A6E70
References
- Amrine-Madsen H, Scally M, Westerman M, Stanhope MJ, Krajewski C, Springer MS. 2003. Nuclear gene sequences provide evidence for the monophyly of australidelphian marsupials. Molecular Phylogenetics and Evolution 28: 186-196.
- Archer M. 1976. The basicranial region of marsupicarnivores (Marsupialia), interrelationships of carnivorous mammals, and affinities of the insectivorous marsupial peramelids. Zoological Journal of the Linnean Society 59: 217-322.
- Archibald JD. 1979. Oldest known eutherian stapes and a marsupial petrosal bone from the Late Cretaceous of North America. Nature 281: 669-670.
- Asher RJ. 1999. A morphological basis for assessing the phylogeny of the 'Tenrecoidea' (Mammalia, Lipotyphla). Cladistics 15: 231-252.
- Asher RJ, Horovitz I, Sanchez-Villagra MR. 2004. First combined cladistic analysis of marsupial mammal interrelationships. Molecular Phylogenetics and Evolution 33: 240- 250.
- Beck RMD. 2008. A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89: 175-189.
- Bremer K. 1988. The limits of amino acid sequence data in angiosperm phylogenetic reconstruction. Evolution 42: 795- 803.
- Cardillo M, Bininda-Emonds ORP, Boakes E, Purvis A. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology (London) 264: 11-31.
- Case JA, Goin FJ, Woodburne MO. 2005. 'South American' marsupials from the Late Cretaceous of North America and the origin of marsupial cohorts. Journal of Mammalian Evolution 12: 461-494.
- Cifelli RL. 1993. Early Cretaceous mammal from North America and the evolution of marsupial dental characters. Proceedings of the National Academy of Sciences of the United States of America 90: 9413-9416.
- Clemens WA. 1966. Fossil mammals of the Type Lance Formation Wyoming. Part II. Marsupialia. University of California Publications in Geological Sciences 62: 1-122.
- Dashzeveg D, Dingus L, Loope DB, Swisher CC III, Dulam T, Sweeney MR. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the upper Cretaceous Djadokhta Formation, Southern Ulan Nur Basin, Mongolia. American Museum Novitates 3498: 1-31.
- Dom R, Fisher BL, Martin GF. 1970. The venous system of the head and neck of the opossum (Didelphis virginiana). Journal of Morphology 132: 487-496.
- Fernandez C, Schmidt RS. 1963. The opossum ear and evolution of the coiled cochlea. Journal of Comparative Neurology 121: 151-159.
- Flynn JJ, Swisher CC III. 1995. Cenozoic South American land mammal ages: correlation to global geochronologies. In: Berggren WA, Kent DV, Aubry M-P, Hardenbol J, eds. Geochronology, time scales and global stratigraphic correlation. Tulsa, OK: Society of Economic Paleontologists and Mineralogists, 317-334.
- Gaudin TJ, Wible JR, Hopson JA, Turnbull WD. 1996. Reexamination of the morphological evidence for the cohort Epitheria (Mammalia, Eutheria). Journal of Mammalian Evolution 3: 31-79.
- Gaupp E. 1902. Uber die Ala temporalis des Saugetierschadels und die Regio orbitalis einiger anderer Wirbeltierschadels. Anatomische Hefte 19: 155-230.
- Gaupp E. 1905. Neue Deutungen auf dem Gebiete der Lehre vom Saugetierschadel. Anatomischer Anzeiger 27: 273-310.
- Gaupp E. 1908. Zur Entwicklungsgeschichte und vergleichenden Morphologie des Schadels von Echidna aculeata var. typica. Semon's Zoologische Forschungsreisen in Australien. Denkschriften der medicinischnaturwissenscaftliche Gesellschaft zu Jena 6: 539-788.
- Gelderen C. 1924. Die Morphologie der Sinus durae matris. Zweiter Teil. Die vergleichende Ontogenie der neurokraniellen Venen der Vogel und Saugetiere. Zeitschrift fur Anatomie und Entwickelungsgeschichte 74: 432-508.
- Godthelp H, Wroe S, Archer M. 1999. A new marsupial from the Early Eocene Tingamarra local fauna of Murgon, Southeastern Queensland: a prototypical Australian marsupial. Journal of Mammalian Evolution 6: 289-313.
- Goin FJ, Carlini AA. 1995. An early Tertiary microbiotheriid marsupial from Antartica. Journal of Vertebrate Paleontology 15: 205-207.
- Goin FJ, Oliveira EV, Candela AM. 1998. Carolocoutoia ferigoloi nov. gen. and sp. (Protodidelphidae), a new Paleocene 'opossum-like' marsupial from Brazil. Palaeovertebrata 27: 145-154.
- Goin FJ, Pascual R, Tejedor MF, Gelfo JN, Woodburne MO, Case JA, Reguero MA, Bond M, Lopez GM, Cione AL, Udrizar Sauthier D, Balarino L, Scasso RA, Medina FA, Ubaldon MC. 2006. The earliest Tertiary therian mammal from South America. Journal of Vertebrate Paleontology 26: 505-510.
- Hochstetter F. 1896. Beitraige zur Anatomie und Entwickelungsgeschichte des Blutgefaisssystems der Monotremen. Semon's Zoologische Forschungsreisen in Australien 5: 189- 243.
- Hopson JA, Rougier GW. 1993. Braincase structure in the oldest known skull of therian mammal: implications for mammalian systematics and cranial evolution. American Journal of Science 293: 268-299.
- Horovitz I, Ladeveze S, Argot C, Hooker JJ, Macrini TE, Martin T, Kurz C, de Muizon C, Sanchez-Villagra MR. 2008. The anatomy of Herpetotherium fugax Cope 1873, a metatherian from the Oligocene of North America. Palaeontographica Abt. A 284: 109-141.
- Horovitz I, Sanchez-Villagra MR. 2003. A morphological analysis of marsupial mammal higher-level phylogenetic relationships. Cladistics 19: 181-212.
- Hunt RM. 1974. The auditory bulla in Carnivora: an anatomical basis for reappraisal of carnivore evolution. Journal of Morphology 143: 21-76.
- Kearney M, Clark JM. 2003. Problems due to missing data in phylogenetic analyses including fossils: a critical review. Journal of Vertebrate Paleontology 23: 263-274.
- Kermack KA, Mussett F, Rigney HW. 1981. The skull of Morganucodon. Zoological Journal of the Linnean Society 71: 1-158.
- Kielan-Jaworowska Z, Cifelli RL, Luo Z-X. 2004. Mammals from the age of dinosaurs: origins, evolution, and structure. New York, NY: Columbia University Press.
- Kirsch JAW, Lapointe F-J, Springer MS. 1997. DNAhybridisation studies of marsupials and their implications for metatherian classification. Australian Journal of Zoology 45: 211-280.
- Kirsch JAW, Reig OA, Springer MS. 1991. DNA hybridization evidence for the Australian affinity of the American marsupial Dromiciops australis. Proceedings of the National Academy of Sciences of the United States of America 88: 10465-10469.
- Kuhn H-J, Zeller U. 1987. The cavum epiptericum in monotremes and therian mammals. In: Kuhn H-J, Zeller U, eds. Morphogenesis of the mammalian skull. Hamburg: Verlag Paul Varey, 51-70.
- Ladeveze S. 2004. Metatherian petrosals from the Late Paleocene of Itaborai (Brazil), and their phylogenetic implications. Journal of Vertebrate Paleontology 24: 202-213.
- Ladeveze S. 2007a. Isolated mammalian fossil remains of petrosals and teeth: what to do with? Morphometric and phylogenetic analyses of metatherian petrosal bones from the Paleocene of South America. 8th International Congress of Vertebrate Morphology. Journal of Morphology 268: 1097.
- Ladeveze S. 2007b. Petrosal bones of metatherian mammals from the Late Paleocene of Itaborai (Brazil), and a cladistic analysis of petrosal features in metatherians. Zoological Journal of the Linnean Society 150: 85-115.
- Ladeveze S, de Muizon C. 2007. The auditory region of early Paleocene Pucadelphydae (Mammalia, Metatheria) from Tiupampa, Bolivia, with phylogenetic implications. Palaeontology 50: 1123-1154.
- Lofgren DL. 1995. The Bug Creek problem and the Cretaceous-Tertiary transition at McGuire Creek, Montana. University of California Publications in Geological Sciences 140: 1-185.
- Luckett WP. 1994. Suprafamilial relationships within Marsupialia: resolution and discordance from multidisciplinary data. Journal of Mammalian Evolution 2: 255-283.
- Luo Z-X, Ji Q, Wible JR, Yuan C-X. 2003. An early Cretaceous tribosphenic mammal and metatherian evolution. Science 302: 1934-1940.
- Luo Z-X, Kielan-Jaworowska Z, Cifelli RL. 2002. In quest for a phylogeny of Mesozoic mammals. Acta Palaeontologica Polonica 47: 1-78.
- McKenna MC, Bell SK. 1997. Classification of mammals above the species level. New York, NY: Columbia University Press.
- MacPhee RDE. 1981. Auditory regions of primates and eutherian insectivores: morphology, ontogeny, and character analysis. Contributions to Primatology 18: 1-282.
- MacPhee RDE, Novacek MJ, Storch G. 1988. Basicranial morphology of early Tertiary Erinaceomorphs and the origin of primates. American Museum Novitates 2921: 1-42.
- Maddison DR. 1991. The discovery and importance of multiple islands of most-parsimonious trees. Systematic Zoology 40: 315-328.
- Marshall LG. 1987. Systematics of Itaboraian (Middle Paleocene) age opossum-like marsupials from the limestone quarry at Sao Jose de Itaborai, Brazil. In: Archer M, ed. Possums and opossums: studies in evolution. Sydney: Surrey Beatty & Sons and the Royal Zoological Society of New South Wales, 91-160.
- Marshall LG, Case JA, Woodburne MO. 1990. Phylogenetic relationships of the families of marsupials. In: Genoways HH, ed. Current mammalogy. New York, NY: Plenum Press, 433-505.
- Marshall LG, Kielan-Jaworowska Z. 1992. Relationships of the dog-like marsupials, deltatheroidans, and early tribosphenic mammals. Lethaia 25: 361-374.
- Marshall LG, de Muizon C. 1988. The dawn of the Age of Mammals in South America. National Geographic Research 4: 23-55.
- Marshall LG, de Muizon C. 1995. Part II. The skull. In: de Muizon C, ed. Pucadelphys andinus (Marsupialia, Mammalia) from the early Paleocene of Bolivia. Paris: Memoires du Museum national d'Histoire naturelle, 21-90.
- Marshall LG, de Muizon C, Sigogneau-Russel D. 1995. Pucadelphys andinus (Marsupialia, Mammalia) from the early Paleocene of Bolivia. Paris: Memoires du Museum national d'Histoire naturelle.
- Marshall LG, Sempere T, Butler RF. 1997. Chronostratigraphy of the mammal-bearing Paleocene of South America. Journal of South American Earth Sciences 10: 49-70.
- Meng J, Fox RC. 1995a. Osseous inner ear structures and hearing in early marsupials and placentals. Zoological Journal of the Linnean Society 115: 47-71.
- Meng J, Fox RC. 1995b. Therian petrosals from the Oldman and Milk River Formations (Late Cretaceous), Alberta. Journal of Vertebrate Paleontology 15: 122-130.
- Meredith RW, Westerman M, Case J, Springer MS. 2008. Phylogeny and timescale for marsupial evolution based on sequences for five nuclear genes. Journal of Mammalian Evolution 15: 1-36.
- de Muizon C. 1992. La fauna de mamiferos de Tiupampa (Paleoceno inferior, Formacion Santa Lucia), Bolivia. In: Suarez-Soruco R, ed. Fosiles y Facies de Bolivia, Vol. I, Vertebrados. Santa Cruz: Revista Tecnica, Yacimientos Petroliferos Fiscales Bolivianos, 575-624.
- de Muizon C. 1994. Marsupial skeletons from the Early Paleocene of Tiupampa (Bolivia) diagnostic features of fossil marsupials. Journal of Vertebrate Paleontology 14 (Suppl. 3): 39A.
- de Muizon C. 1998. Mayulestes ferox, a boryaenoid (Metatheria, Mammalia) from the early Palaeocene of Bolivia. Phylogenetic and palaeobiologic implications. Geodiversitas 20: 19-142.
- de Muizon C. 1999. Marsupial skulls from the Deseadan (Late Oligocene) of Bolivia and phylogenetic analysis of the Borhyaenoidea (Marsupialia, Mammalia). Geobios 32: 483- 509.
- de Muizon C, Brito IM. 1993. Le bassin calcaire de Sao Jose de Itaborai (Rio de Janeiro, Bresil): ses relations fauniques avec le site de Tiupampa (Cochabamba, Bolivie). Annales de Paleontologie 79: 233-268.
- de Muizon C, Cifelli RL. 2001. A new basal 'Didelphoid' (Marsupialia, Mammalia) from the Early Paleocene of Tiupampa (Bolivia). Journal of Vertebrate Paleontology 21: 87-97.
- de Muizon C, Cifelli RL, Bergqvist LP. 1998. Eutherian tarsals from the early Paleocene of Bolivia. Journal of Vertebrate Paleontology 18: 655-663.
- de Muizon C, Cifelli RL, Cespedes Paz R. 1997. The origin of the dog-like borhyaenoid marsupials of South America. Nature 389: 486-489.
- Nilsson MA, Arnason U, Spencer PBS, Janke A. 2004. Marsupial relationships and a timeline for marsupial radiation in South Gondwana. Gene 340: 189-196.
- Nilsson MA, Gullberg A, Spotorno AE, Arnason U, Janke A. 2003. Radiation of extant marsupials after the K/T boundary: evidence from complete mitochondrial genome. Journal of Molecular Evolution 57: 3-12.
- Nixon KC, Carpenter JM. 1996. On simultaneous analysis. Cladistics 12: 221-241.
- Palma RE, Spotorno AE. 1999. Molecular systematics of marsupials based on the rRNA 12S mitochondrial gene: the phylogeny of didelphimorphia and of the living fossil microbiotheriid Dromiciops gliroides Thomas. Molecular Phylogenetics and Evolution 13: 525-535.
- de Paula Couto C. 1952a. Fossil mammals from the beginning of the Cenozoic in Brazil. Marsupialia: Didelphidae. American Museum Novitates 1567: 1-26.
- de Paula Couto C. 1952b. Fossil mammals from the beginning of the Cenozoic in Brazil. Marsupialia: Polydolopidae and Borhyaenidae. American Museum Novitates 1559: 1-27.
- de Paula Couto C. 1952c. A new name for Mirandaia riberoi Paula Couto, 1952. Journal of Mammalogy 33: 503.
- de Paula Couto C. 1961. Marsupiais fosseis do Paleoceno do Brazil. Anais da Academia Brasileira de Ciencias 33: 312- 333.
- de Paula Couto C. 1962. Didelphideos fosiles del Paleoceno de Brazil. Revista del Museo Argentino de Ciencias Naturales 'Bernadino Rivadavia', Ciencias Zoologicas 112: 135- 166.
- Phillips MJ, Lin Y-H, Harrison GL, Penny D. 2001. Mitochondrial genomes of a bandicoot and a brushtail possum confirm the monophyly of australidelphian marsupials. Proceedings of the Royal Society of London Series B 268: 1533- 1538.
- Phillips MJ, MacLenachan PA, Down C, Gibb GC, Penny D. 2006. Combined mitochondrial and nuclear protein-coding DNA sequences resolve the Interrelations of the major Australasian marsupial radiations. Systematic Biology 55: 122-137.
- Reig OA, Kirsch JAW, Marshall LG. 1987. Systematic relationships of the living and Neocenozoic American 'opossum-like' marsupials (suborder Didelphimorphia), with comments on the classification of these and of the Cretaceous and Paleogene New World and European metatherians. In: Archer M, ed. Possums and opossums: studies in evolution. Sydney: Surrey Beatty & Sons and the Royal Zoological Society of New South Wales, 1-89.
- Rougier GW, Bonaparte JF. 1988. La pared lateral del craneo de Vincelestes neuquenianus (Mammalia, Eupantotheria) y su importancia en el estudo de los mamiferos mesozoicos. V Jornadas Argentina de Paleontologia de Vertebrados Resumenes: 14-15.
- Rougier GW, Wible JR. 2006. Major changes in the ear region and basicranium of early mammals. In: Carrano M, Gaudin TJ, Blob R, Wible JR, eds. Amniote paleobiology: phylogenetic and functional perspectives on the evolution of mammals, birds and reptiles. Chicago, IL: University of Chicago Press, 269-311.
- Rougier GW, Wible JR, Hopson JA. 1992. Reconstruction of the cranial vessels in the Early Cretaceous mammal Vincelestes neuquenianus: implications for the evolution of the mammalian cranial vascular system. Journal of Vertebrate Paleontology 12: 188-216.
- Rougier GW, Wible JR, Hopson JA. 1996. Basicranial anatomy of Priacodon fruitaensis (Triconodontidae, Mammalia) from the Late Jurassic of Colorado, and a reappraisal of mammaliaform interrelationships. American Museum Novitates 3183: 1-38.
- Rougier GW, Wible JR, Novacek MJ. 1996. Middle-ear ossicles of the multituberculate Kryptobaatar from the Mongolian Late Cretaceous: implications for mammaliamorph relationships and the evolution of the auditory apparatus. American Museum Novitates 3187: 1-43.
- Rougier GW, Wible JR, Novacek MJ. 1998. Implications of Deltatheridium specimens for early marsupial history. Nature 396: 459-463.
- Rougier GW, Wible JR, Novacek MJ. 2004. New specimen of Deltatheroides cretacicus (Metatheria, Deltatheroidea) from the Late Cretaceous of Mongolia. Bulletin of Carnegie Museum of Natural History 36: 245-266.
- Sanchez-Villagra MR. 2001a. Ontogenetic and phylogenetic transformations of the vomeronasal complex and nasal floor elements in marsupial mammals. Zoological Journal of the Linnean Society 131: 459-479.
- Sanchez-Villagra MR. 2001b. The phylogenetic relationships of argyrolagid marsupials. Zoological Journal of the Linnean Society 131: 481-496.
- Sanchez-Villagra MR, Ladeveze S, Horovitz I, Argot C, Hooker JJ, Macrini T, Martin T, Moore-Fay S, de Muizon C, Schmelzle T, Asher RJ. 2007. Exceptionally preserved North American Paleogene metatherians: adaptations and discovery of a major gap in the opossum fossil record. Proceedings of the Royal Society, Biology Letters 3: 318-322.
- Sanchez-Villagra MR, Schmelzle T. 2007. Anatomy and development of the bony inner ear in the woolly opossum, Caluromys philander (Didelphimorphia, Marsupialia). Mastozoologia Neotropical 14: 53-60.
- Sanchez-Villagra MR, Wible JR. 2002. Patterns of evolutionary transformation in the petrosal bone and some basicranial features in marsupial mammals, with special reference to didelphids. Journal of Zoological Systematics and Evolutionary Research 40: 26-45.
- Segall W. 1970. Morphological parallelisms of the bulla and auditory ossicles in some insectivores and marsupials. Fieldiana: Zoology, 51: 169-205.
- Sorenson MD, Franzosa EA. 2007. TreeRot, Version 3. Boston, MA: Boston University.
- Springer MS. 1997. Molecular clocks and the timing of the placental and marsupial radiations in relation to the Cretaceous-Tertiary boundary. Journal of Mammalian Evolution 4: 285-302.
- Springer MS, Kirsch JAW, Case JA. 1997. The chronicle of marsupial evolution. In: Givnish TJ, Sytsma KJ, eds. Molecular evolution and adaptative radiation. New York, NY: Cambridge University Press, 129-161.
- Springer MS, Westerman M, Kavanagh JR, Burk A, Woodburne MO, Kao DJ, Krajewski C. 1998. The origin of the Australasian marsupial fauna and the phylogenetic affinities of the enigmatic monito del monte and marsupial mole. Proceedings of the Royal Society of London. Series B. Biological Sciences 265: 2381-2386.
- Swisher CC III, Wang Y-Q, Wang X-L, Xu X, Wang Y. 1999. Cretaceous age for the feathered dinosaurs of Liaoning, China. Nature 400: 58-61.
- Swofford DL. 2002. PAUP*. Phylogenetic analysis using parsimony (*and other methods), Version 4. Sunderland, MA: Sinauer Associates.
- Szalay FS. 1982a. A new appraisal of marsupials phylogeny and classification. In: Archer M, ed. Carnivorous marsupials. Sydney: Royal Zoological Society of New South Wales, 621-640.
- Szalay FS. 1982b. Phylogenetic relationships of the marsupials. Geobios memoire special 6: 177-190.
- Szalay FS. 1994. Evolutionary history of the marsupials and an analysis of osteological characters. Cambridge; New York, NY: Cambridge University Press.
- Szalay FS, Sargis EJ. 2001. Model-based analysis of postcranial osteology of marsupials from the Paleocene of Itaborai (Brazil) and the phylogenetics and biogeography of Metatheria. Geodiversitas 23: 139-302.
- Van der Klaauw CJ. 1931. The auditory bulla in some fossil mammals, with a general introduction to this region of the skull. Bulletin of the American Museum of Natural History LXII: 1-352.
- Voit M. 1909. Das primordialcranium des kaninchens unter berucksichtigung der deckknochen. Ein beitrag zur morphologie des saugetierschadels. Anatomische Hefte 38: 425- 616.
- Voss RS, Jansa SA. 2003. Phylogenetic studies on didelphid marsupials II. Nonmolecular data and new IRBP sequences: separate and combined analyses of didelphine relationships with denser taxon sampling. Bulletin of the American Museum of Natural History 276: 1-82.
- Wible JR. 1986. Transformations in the extracranial course of the internal carotid artery in mammalian phylogeny. Journal of Vertebrate Paleontology 6: 313-325.
- Wible JR. 1987. The eutherian stapedial artery: character analysis and implications for superordinal relationships. Zoological Journal of the Linnean Society 91: 107-135.
- Wible JR. 1990. Petrosals of Late Cretaceous marsupials from North America, and a cladistic analysis of the petrosal in therian mammals. Journal of Vertebrate Paleontology 10: 183-205.
- Wible JR. 2003. On the cranial osteology of the short-tailed opossum Monodelphis brevicaudata (Didelphidae, Marsupialia). Annals of Carnegie Museum 72: 137-202.
- Wible JR, Hopson JA. 1993. Basicranial evidence for early mammal phylogeny. In: Szalay FS, Novacek MJ, McKenna MC, eds. Mammal phylogeny: Mesozoic differentiation, multituberculates, monotremes, early therians, and marsupials. New York, NY: Springer-Verlag, 45-62.
- Wible JR, Hopson JA. 1995. Homologies of the prootic canal in mammals and non-mammalian cynodonts. Journal of Vertebrate Paleontology 15: 331-356.
- Wible JR, Rougier GW, Novacek MJ, McKenna MC. 2001. Earliest eutherian ear region: a petrosal referred to Prokennalestes from the Early Cretaceous of Mongolia. American Museum Novitates 3322: 1-44.
- Woodburne MO, Case JA. 1996. Dispersal, vicariance, and the Late Cretaceous to Early Tertiary land mammal biogeography from South America to Australia. Journal of Mammalian Evolution 3: 121-161.
- Wroe S. 1997. A reexamination of proposed morphology-based synapomorphies for the families of Dasyuromorphia (Marsupialia): Part I, Dasyuridae. Journal of Mammalian Evolution 4: 19-52.
- Wroe S. 1999. The geologically oldest dasyurid, from the Miocene of Riversleigh, north-west Queensland. Palaeontology 42: 501-527.
- Wroe S, Ebach M, Ahyong S, de Muizon C, Muirhead J. 2000. Cladistic analysis of dasyuromorphian (Marsupialia) phylogeny using cranial and dental characters. Journal of Mammalogy 81: 1008-1024.