A new species-group of the genus Anthaxia ( Haplanthaxia ) from south-eastern Asia , with descriptions of two new species ( Coleoptera : Buprestidae : Anthaxiini )

Two new species of the genus Anthaxia Eschscholtz, 1829 are described: A. (Haplanthaxia) phobos sp. nov. (Thailand) and A. (H.) deimos sp. nov. (China, Laos, Thailand). Anthaxia (H.) phobos species-group is defi ned and a review of all the currently defi ned species-groups in the subgenus Haplanthaxia Reitter, 1911 is provided. New species are illustrated and the newly defi ned species-group is differentiated from previously defi ned groups.


Introduction
The large genus Anthaxia Eschscholtz, 1829 (about 800 species according to BELLAMY (2008)) of worldwide distribution (except for the Australian Region) is currently divided into six subgenera (BELLAMY, 2008).There is no doubt that the subgenus Haplanthaxia Reitter, 1911 is the largest subgenus, comprising about 60 % of species of the genus.So far 11 species-groups have been defi ned in this subgenus according to BELLAMY (2008) and 5 species-groups were defi ned subsequently by BÍLÝ (2010BÍLÝ ( , 2015)), BÍLÝ & KUBÁŇ (2010, 2012) and BÍLÝ & SAKALIAN (2014).Moreover two species-groups were omitted by BELLAMY (2008) in his catalogue and one more was transferred from Anthaxia s. str.into Haplanthaxia by BÍLÝ & KUBÁŇ (2012), so at present there are 19 species-groups defi ned in the subgenus Haplanthaxia and many more are awaiting defi nition; one of them (20 th ) is defi ned in this paper.
The currently defi ned species-groups of the subgenus Haplanthaxia are: A. aeneocuprea species-group (BÍLÝ 2015) A. atomaria species-group (BÍLÝ 2000) A. cichorii species-group (OBENBERGER 1917 Having studied the rich material of Anthaxia collected in the course of the last decades in south-eastern Asia I found two undescribed species from China, Laos and Thailand belonging to a very characteristic, so far undefi ned species-group, which is defi ned in this contribution together with the descriptions of the two new species.

Material and methods
A Canon D-550 digital camera with attached Canon MP-E65mm f/2.8-5× macro lens was used to capture the colour images; multiple photographs taken were combined with Helicon Focus image stacking software.
The body length was measured in the middle of the body following the elytral suture (the same for the pronotal and elytral length); width of body means the maximum body width (usually the maximum span between lateral pronotal margins or span between outer margin of humeral callosities).The double slash (//) is used for the separating data on the different labels, the square brackets ([]) for clarifi cation of the text of the locality labels.The morphological terminology follows that of BÍLÝ (2010) and BÍLÝ & KUBÁŇ (2010).
The species-group can be simply attributed to the subgenus Haplanthaxia due to the general body-shape, morphology of the elytral margins and the form of the elytral epipleura at subhumeral part.The species-group is defi ned by the complete set of the above mentioned characters since each of them separately can be found also in other species-groups and not only in the subgenus Haplanthaxia.Anthaxia (Haplanthaxia) phobos sp.nov.Diagnosis.Medium-sized (length 4.2-6.0mm), stout, convex, completely asetose, lustrous (Figs 1-2); dorsal surface black-bronze with fi ne green lustre (more intense at basal portion of elytra and along suture); pronotum with red-orange lateroposterior depressions and narrow, medial stripe; frons dark golden-orange in both sexes, vertex black; antennae and legs black with green lustre; ventral surface black, proepisterna and lateral portions of abdominal ventrites golden-orange.Description of the male holotype (Fig. 1).Head large as wide as width of anterior pronotal margin; anterior margin of frontoclypeus almost straight; frons regularly, rather strongly convex, vertex weakly convex, 0.9 times as wide as width of eye; eyes large, widely reniform, not projecting beyond outline of head; sculpture of head homogeneous consisting of small, dense, regular, oval cells with well developed central grains; antennae short reaching posterior third of lateral pronotal margins when laid alongside; scape 5 times as long as wide, slightly curved, claviform, pedicel elliptical, 1.5 times as long as wide; third antennomere obtusely triangular, as wide as long, antennomeres 4-10 trapezoidal, somewhat wider than long; terminal antennomere rhomboid, 1.5 times as long as wide.
Pronotum rather strongly convex, 1.8 times as wide as long, with weak, shallow lateroposterior depressions; anterior margin deeply bisinuate with wide, projecting medial lobe, posterior margin very weakly bisinuate, almost straight; lateral margins weakly arcuate, posterior angles obtuse-angled, maximum pronotal width just anterior to mid-length; sculpture consisting of small, dense, regular, polygonal cells with tiny central grains; sculpture rougher and more distinct in lateroposterior depressions.Scutellum large, triangular, fl at, microsculptured, slightly longer than wide.
Elytra almost regularly convex, 1.9 times as long as wide, weakly wedge-shaped, very slightly uneven at posterior fourth; each elytron regularly rounded apically, humeral callosities small, not projecting beyond outline of elytra; basal, transverse depressions deep, wide, almost reaching scutellum; elytral epipleura narrow, parallel-sided, not reaching elytral apex; only very tips of elytra with very fi ne, lateral serrations; sculpture very fi ne, homogeneous, almost scale-shaped, rougher, weakly granulate in basal, transverse depressions.
Ventral surface lustrous, with rather rough eye-like sculpture with large central grains, abdominal ventrites with very fi ne, eye-like sculpture; prosternal process weakly convex, strongly widening behind procoxae, pointed apically.Anal ventrite narrowly truncate with very fi ne, lateral serrations.Legs thin, relatively short, tibiae almost straight, without inner serrations; tarsi slender, distinctly shorter than corresponding tibiae, fourth tarsomere yellow-brown with wide adhesive pads.Tarsal claws fi ne, weakly hook-shaped, yellow-brown, only slightly enlarged at base.Aedeagus (Fig. 3) wide, short spindle-shaped, moderately sclerotised, parameres conspicuously narrowed in posterior fi fth, obliquely truncate apically; median lobe broad, widely pointed apically with fi ne dorsolateral serrations.Measurements.Length: 4.2-6.0mm (holotype 4.3 mm); width: 1.5-2.2mm (holotype 1.5 mm).Sexual dimorphism.Only very slightly expressed; the female differs from the male only by the stouter body, slightly different colouration (Fig. 2) and by the wider vertex (vertex as wide as width of eye in female).Variability.No variability was observed except for the size and sexual dichromatism.Bionomy.Unknown.Differential diagnosis.Anthaxia (Haplanthaxia) phobos sp.nov. is similar and most probably related to the following species, A. (H.) deimos sp.nov., from which it differs by the characters given in the differential diagnosis of A. (H.) deimos sp.nov.(see below).Etymology.This species is named after the larger moon of Mars -"Phobos"; noun in apposition.Distribution.Thailand.Note.One female labelled: ʻLaos NE, Hua Phan Province, Ban Saleui, Phou Pan (Mt), 20°12′N 104°01′E, 1300-1900m, 1.-31.v.2011, Leg. C. Holzschuhʼ (NMPC) was not included among paratypes since (apart from the locality) it differs by the completely golden-orange ventral surface and almost unicolorous, dark, black-bronze pronotum; other characters completely fi t to Anthaxia (H.) phobos sp.nov.but it could belong to a different species and without a male it is impossible to evaluate this specimen.Diagnosis.Relatively small (length 3.2-4.3mm), fl attened, subparallel, completely asetose, very lustrous (Figs 4-5); dorsal surface bronze-violet, frons, anterior pronotal angles, antennae and legs golden green, vertex bronze (male) or anterior pronotal angles coppery red, antennae and legs green, frons and vertex dark bronze (female); ventral surface red-bronze with green lustre, meso-and metafemora bright green (male) or ventrally entirely red-bronze (female).Description of the male holotype (Fig. 4).Head large as wide as width of anterior pronotal margin; anterior margin of frontoclypeus almost straight; frons regularly, rather strongly convex, vertex weakly convex, 1.1 times as wide as width of eye; eyes large, widely reniform, only very weakly projecting beyond outline of head; sculpture of head homogeneous consisting of small, dense, regular, oval cells with tiny central grains; antennae relatively short almost reaching posterior pronotal angles when laid alongside; scape 4.5 times as long as wide, slightly curved, claviform, pedicel elliptical, twice as long as wide; third antennomere obtusely triangular, as wide as long, fourth antennomere triangular, slightly wider than long, antennomeres 5-10 trapezoidal, somewhat wider than long; terminal antennomere rhomboid, 1.5 times as long as wide.
Pronotum fl attened, 1.7 times as wide as long, with deep, wide lateroposterior depressions almost occupying posterior half of pronotum; anterior margin deeply bisinuate with wide, projecting medial lobe, posterior margin very weakly bisinuate, almost straight; lateral margins weakly arcuate, posterior angles obtuse-angled, maximum pronotal width at anterior third; sculpture consisting of small, dense, rather poorly defi ned, polygonal cells with tiny central grains.Scutellum large, subcordiform, fl at, microsculptured, 1.4 times as long as wide.