Tryphosa nana (Krøyer, 1846a)

(Fig. 23)

Anonyx nanus Krøyer, 1846a: 30.― Krøyer, 1846b: pl. 17 fig. 2a–t.― Bruzelius, 1859: 42.― Bate, 1862: 78, pl. 12 fig. 2. ― Lilljeborg, 1865a: 18.― Lilljeborg, 1865b: 28.― Heller, 1866: 24.― Norman, 1869: 274.― Stalio, 1877: 243.― Graeffe, 1880: 48.― Stossich, 1880: 71.― Nebeski, 1881: 34.― Krapp-Schickel, 1974: 321, 338.

Tryphosa nanus.― Boeck, 1871: 118.― Boeck, 1876: 181.

Tryphosa ciliata Sars, 1883: 81, pl. 3 fig. 4.― Chevreux, 1887: 295, 331.― Walker, 1888: 172, pl. 13 figs 1–4.― Walker, 1889: 77.― Walker, 1892: 241.

Tryphosa nana.― Chevreux, 1888: 3.― Meinert, 1890: 155.― Bonnier, 1893: 191, 196, pl.7.― Norman, 1900: 203.― Scott, 1901.― Christodoulou, Paraskevopoulou, Syranidou & Koukouras, 2013: 1312, table 2.

Tryphosa höringii.― Walker, 1889: 77.

Orchomenella ciliata.― Sars, 1891: 69, pl. 25 fig. 2.― Robertson, 1892: 204.― Sars, 1895: 683.― Walker, 1895a: 420. ― Walker, 1895b: 292.

Anonyx pinguis.― Della Valle, 1893: 821, pl. 28 figs 22–35.

Orchomenella nana.― Chevreux, 1898: 476.― Walker, 1898: 166.― Walker, 1904: 244.― Stephensen, 1923: 12.―Chevreux, 1925: 288.― Chevreux & Fage, 1925: 71, fig. 62.― Stephensen, 1929: 60.―Marine Biological Association of the UK, 1931: 188.― Stephensen, 1932: 357.― Stephensen, 1935: 104 (key).― Crawford, 1937: 644.― Moore, 1937: 117. ― Schellenberg, 1942: 108.― Stephensen, 1942: 474.― Gorbunov, 1946: 43.― Jones, 1948: 404.― Enequist, 1949: 387, 400.― Reid, 1951: 195.―J.L. Barnard, 1958: 97.― Oldevig, 1959: 22.― Duhig, 1960: 64.― Agrawal, 1963: 632, pl. 3. ― Gurjanova, 1964: 279.―Toulmond, 1964: 321.― Toulmond & Truchot, 1964: 7.― Agrawal, 1966: 459, pl. 5.― Nayar, 1966: 135.― O'Riordan, 1969: 32.― Bellan-Santini & Ledoyer, 1973: 920.― Laverack & Blackler, 1974: 81.― Drago & Albertelli, 1978: 212.― Myers & Southgate, 1980: 975.― Walker & Rees, 1980: 56.― Macquart-Moulin, 1984: 185. ― Diviacco & Ruffo, 1989: 525, fig. 359 (in part).— Dauvin & Sorbe, 1995: 443, 447, 449, 456, tables 4, 5.— Stefanidou & Voultsiadou-Koukoura, 1995: 602, table 1.— Cartes & Sorbe, 1999: 1138, table 1, 1134, table 4.— Bakir & Çevirgen, 2010: 83, table 1.

Orchomenella nanus.― Stebbing, 1906: 81.― Tattersall, 1913: 4.― Schellenberg, 1925: 118.

Orchomene nana.―J.L. Barnard, 1964: 89 (key).― Karaman, 1973: 133, figs 14–18 (in part).― Krapp-Schickel, 1974: 321, 338.― Kitching et al, 1976: 756.― Lincoln, 1979: 70, figs 25a–g, 26e–h.― O'Riordan, 1982: 551.― Moore, 1984: 35. ― Parker, 1984: 122.― Bellan-Santini, 1985: 275.― Marques & Bellan-Santini, 1985: 323, 349.― Holmes, 1986: 37. ― Costello et al, 1989: 34.― Kaartvedt, 1989: 189, table 3, 191, table 5.― Dauvin & Gentil, 1990: 126, 128.― Buhl-Jensen & Fosså, 1991: 248.― Hamerlynck & Mees, 1991: 208. table 2.― Mees & Hamerlynck, 1992: 360, table 2.― Cattrijsse, Mees & Hamerlynck, 1993: 191, table 1.― Marques & Bellan-Santini, 1993: 557, table 2, 559, table 3, 560, table 4, 561, table 5.― Dauvin et al, 1994: 550, table 3.― Dauvin & Zouhiri, 1996: 914, table 3, 916, table 5, 921, table 6.― Conradi & Lopez-Gónzález, 1999: 6, table 1.

Orchomene nanus.― Barnard & Karaman, 1991: 509.― Conlan, 1994: 520

not Tryphosa nana.― Sars, 1891: 76, pl. 27 fig. 1 (= Tryphosella sarsi).― Sars, 1895: 684 (= Tryphosella sarsi).―Walker, 1895a: 422 (= Tryphosella sarsi).― Walker, 1895b: 294 (= Tryphosella sarsi).

not Tryphosa nanus.― Robertson, 1892: 205 (?= Tryphosella sarsi).

not Orchomene nana.― Karaman, 1973: 133, figs 14–18 (in part = Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).

not Orchomenella nana.― Ledoyer, 1977: 384 (= Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).― Diviacco & Ruffo, 1989: 525, fig. 359 (in part = Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).

Types. According to Dr Jørgen Olesen (ZMUC) there are two sets of material (ZMUC CRU-7476?type and ZMUC CRU-7487 syntype) which might be considered as original material.

Material examined. 23 specimens & 1 slide (female, 5.5 mm), SMNH N. Amph. 6904, from Gåsörännan, Sweden, 30–40 m; 8 specimens, AM P.11222, from Plymouth, Devon, United Kingdom (50°24'N 4°7'W), coll. A.A. Cameron.

Type locality. The Kattegat.

Habitat. Sand, mud and shelly debris.

Feeding strategies. Often found ‘stripping bait in lobster pots’ (O’Riordan 1982) on carcasses of dead crabs (Norman 1900, Moore 1984), also in Echinocardium tests (Laverack & Blackler 1974).

Depth range. 0–250 m (Lincoln 1979).

Distribution. Tryphosa nana is widely recorded and locally very common along the Atlantic coasts of Europe and the North Sea, from Sweden and southern Norway to Spain (Chevreux & Fage 1925); also reported from Senegal (Schellenberg 1925) and Guinea (Reid 1951). It is also recorded from the Adriatic Sea (Heller 1866, Karaman 1973), Aegean Sea (Bellan-Santini 1985, Christodoulou et al. 2013) and Mediterranean Sea (Bellan- Santini & Ledoyer 1973).