Domiporta pulchra Harzhauser & Landau 2021, nov. sp.

Domiporta pulchra nov. sp.

Figs 7 1 E 1–E 2, F 1–F 2, G 1–G 2, H 1–H 2, 8G 1–G 2, H 1–H 2, I 1–I 2

? Mitra (Mitra) adsita Bell. — Boettger 1906: 7 [non Bellardi, 1887a].

Type material. Holotype. NHMW 2020 /0129/0001, SL: 24.7 mm, MD: 7.3 mm, Lăpugiu de Sus (Romania), figs 7G 1 –G 2, 8I 1 –I 2. Paratypes: NHMW 2020 /0129/0002, SL: 20.9 mm, MD: 6.0 mm, Lăpugiu de Sus (Romania), figs 7E 1 –E 2; NHMW 2020 /0129/0003, SL: 24.7 mm, MD: 7.3 mm, Lăpugiu de Sus (Romania), figs 7F 1 –F 2, 8G 1 –G 2; NHMW 2020 /0129/0004, SL: 21.0 mm, MD: 5.9 mm, Lăpugiu de Sus (Romania), figs 7H 1 –H 2, 8H 1 –H 2.

Additional material. NHMW 1807/0019/0037, Coşteiu de Sus.

Type locality. Lăpugiu de Sus (Romania), Făget Basin.

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, early/middle Badenian (Langhian).

Etymology. Latin pulcher (= beautiful).

Diagnosis. Domiporta species of small to medium size, slender to moderately fusiform, shell with well-developed, regular spiral sculpture of prominent spiral cords separated by deep spiral grooves with delicate growth lines. Colour pattern in UV light of regularly spaced spiral stripes (figs 7E–G).

Description. Shell small to medium sized, slender to moderately slender fusiform with weakly incised suture. Protoconch high conical, mammillate of about three moderately convex, smooth whorls. Teleoconch of seven whorls. Early teleoconch whorls straight sided, almost subcylindrical. Later spire whorls faintly convex with periphery at abapical suture. Sculpture of first teleoconch whorls of weakly opisthocline axial ribs crossed by four convex spiral cords. Axial ribs weakening on third teleoconch whorl. Spiral cords increasing to five on later teleoconch whorls by bifurcation of adapical primary spiral cord. Spiral grooves deep, narrower than cords, with delicate, densely spaced growth lines. Spiral cords slightly widening on last whorl with flattish profile; about 13–15 spiral cords of more or less equal width on last whorl. Last whorl elongate, weakly convex with periphery slightly below adapical tip of aperture; weakly constricted with shallow basal concavity. Aperture narrow, elongate with indistinct anal notch. Columellar callus narrow, thin, sharply delimited. Columella with four oblique columellar folds. Outer lip thin. Siphonal canal long, narrow, straight with deeply incised siphonal notch. Colour pattern in UV light intense, consisting of dark stripes coinciding with spiral grooves.

Shell measurements and ratios. SL = 20.9–24.7 mm, MD: 5.9–7.3 mm; AA = 29–31°, SL/MD: 3.5–3.6, AL/ AW: 6.1–7.1, AH/S: 2.0–2.1.

Discussion. At first sight, specimens of Domiporta pulchra nov. sp. could be mistaken as juveniles of Cancilla planicostata (Bellardi, 1887), but the colour pattern allows a distinct separation. Moreover, the spiral cords are flatter and closer spaced in Cancilla planicostata. Domiporta turpis nov. sp. is even more slender, its spire is higher and its spiral cords are much coarser. The prominent and regularly spaced spiral cords allow a separation from the slightly larger Domiporta amoena nov. sp. Mitra praecedens Bellardi, 1887, from the late Miocene of Italy, is superficially similar, but differs in its lower spire, more incised suture and lower and less constricted last whorl, which excludes it from the genus Domiporta (see Ferrero-Mortara et al. 1981, pl. 48, figs 13a–b). The Pliocene Mediterranean Mitra interposita Bellardi, 1887, differs in its weaker spiral sculpture, higher spire and subcylindrical last whorl (see Ferrero-Mortara et al. 1981, pl. 49, figs 3a–b). Boettger (1906) seemed to have this species at hand when referring to Mitra adsita (Bellardi, 1887), which was described from the Langhian of the Monte dei Cappuccini at Torino (Italy) (Bellardi 1887a: 84, pl. 4, fig. 52). The Italian species, however, differs in its more convex, faintly shouldered whorls and narrower spiral cords.

The Miocene Mitra aequopersulcata Sacco 1904 from Viale (Italy) is based on two syntypes, which are probably not conspecific. The specimen illustrated in Sacco (1904: 83 pl. 18, fig. 44) is reminiscent of D. pulchra, but differs in its more slender shape and higher spire whorls.

Palaeoenvironment. Unknown, probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus, Coşteiu de Sus (own data).