Processing of file "matrix 3.0 nexus.txt" begins... Data matrix has 27 taxa, 57 characters Valid character-state symbols: 012345 Missing data identified by '?' Gaps identified by '-' "Equate" macros in effect: a ==> (01) Outgroup status changed: 1 taxon transferred to outgroup Total number of taxa now in outgroup = 1 Number of ingroup taxa = 26 Character types changed: Of 57 total characters: 13 characters are of type 'ord' (Wagner) 44 characters are of type 'unord' All characters have equal weight Processing of input file "matrix 3.0 nexus.txt" completed. paup> Bootstrap; Bootstrap method with heuristic search: Number of bootstrap replicates = 500 Starting seed = 1310 Number of characters resampled in each replicate = 57 Optimality criterion = parsimony Character-status summary: Of 57 total characters: 13 characters are of type 'ord' (Wagner) 44 characters are of type 'unord' All characters have equal weight 3 characters are parsimony-uninformative Number of parsimony-informative characters = 54 Gaps are treated as "missing" Multistate taxa interpreted as polymorphism Starting tree(s) obtained via stepwise addition Addition sequence: simple (reference taxon = Xyela) Number of trees held at each step = 1 Branch-swapping algorithm: tree-bisection-reconnection (TBR) with reconnection limit = 8 Steepest descent option not in effect 'Maxtrees' setting = 1000 (will not be increased) Branches collapsed (creating polytomies) if maximum branch length is zero 'MulTrees' option in effect No topological constraints in effect Trees are unrooted 500 bootstrap replicates completed Time used = 00:02:58 (CPU time = 00:02:58.2) Bootstrap 50% majority-rule consensus tree (plus other groups compatible with this tree) /------------------------------------------------------------------------------- Xyela(1) | | /------------------------------------------------------------------------- Xiphydria(3) | | | | /-------------------------------------------------------------------- Urocerus(2) \-----+ | | | /-------------------------------------------------------------- Syrista(4) | | | \-29-+ | /----------- Karatavites(5) | | | | | /---------------------32---------------------+ /------ Praeratavites(6) \-17--+ | \-54-+ | | \------ Postxiphidria(7) | | | | /--------------------------------------- Burmorussus(8) | | | | | /----51-----+ /---------------------------------- Praeoryssus(11) \-48--+ | | | | | \-74-+ /---------------------------- Paroryssus(9) | | | | | | \-30--+ /----------------------- Microryssus(10) | | | | | | \-36-+ /----------------- Minyorussus(14) | | | | \-85-+ \-87--+ /----------- Baltorussus(15) | | | | \-76--+ /------ Orussus(12) | \-69-+ | \------ Chalinus(13) | | /--------------------------------------------- Kuafua(16) | | \-65--+ /--------------------------------------- Leptephialtites(22) | | | | /---------------------------------- Anthobosca(26) \-54--+ | | | /------ Amplicella(24) \-69-+ /---------77----------+ | | \------ Xanthopimpla(25) | | \-49--+ /----------------------- Orthogonalys(20) | | | | /----------- Evania(19) \-50-+ | | /-61--+ /------ Pristaulacus(17) | | \-56-+ | | \------ Gasteruption(18) \-69--+ | /----------- Megalyra(21) | | \-24--+ /------ Schlettererius(23) \-32-+ \------ Dendrocerus(27) Bipartitions found in one or more trees and frequency of occurrence (bootstrap support values): 1 2 2 123456789012345678901234567 Freq % -------------------------------------------------- ...........****............ 435.51 87.10% .......******************** 423.13 84.63% .......................**.. 387.30 77.46% ...........**.*............ 380.02 76.00% ........*******............ 368.56 73.71% ................*****.***** 346.20 69.24% ................***.*.*...* 346.11 69.22% ...........**.............. 343.46 68.69% ...............************ 322.93 64.59% ................***........ 304.20 60.84% ................**......... 282.43 56.49% .....**.................... 269.50 53.90% ................*********** 267.81 53.56% .......********............ 252.82 50.56% ................*****.*...* 249.02 49.80% ................*****.***.* 245.83 49.17% ....*********************** 242.46 48.49% .........*.****............ 179.09 35.82% ......................*...* 160.48 32.10% ....***.................... 157.71 31.54% ........**................. 151.77 30.35% ........**.****............ 150.51 30.10% .*.************************ 144.31 28.86% .**.*********************** 143.65 28.73% ................***.*.....* 133.10 26.62% ....*..******************** 129.26 25.85% ....................*.*...* 121.69 24.34% ...............******.***** 121.01 24.20% .......*.......************ 116.92 23.38% ................***.*.***.* 114.04 22.81% ................***...*...* 112.53 22.51% ....................*.....* 110.89 22.18% ................***.......* 104.19 20.84% ........***................ 100.40 20.08% .........******............ 88.62 17.72% ...************************ 86.16 17.23% ................***...*.... 82.25 16.45% ...................*.....*. 67.67 13.53% ..*.*********************** 66.70 13.34% .*.*....................... 65.91 13.18% ...**..******************** 63.71 12.74% ...........***............. 63.16 12.63% .......********.*********** 59.89 11.98% .***...******************** 56.14 11.23% ................*****.....* 53.59 10.72% ................**....*...* 53.49 10.70% .......................***. 52.28 10.46% .*.**.********************* 51.78 10.36% ...*...******************** 51.10 10.22% ................**....*.... 49.59 9.92% ....*.********************* 48.77 9.75% .................*....*...* 48.59 9.72% .................*........* 47.58 9.52% .......*...****............ 46.36 9.27% .......*...**************** 43.28 8.66% .*.*...******************** 41.25 8.25% ................*****..**** 40.41 8.08% .................**........ 40.09 8.02% ...........*..*............ 38.39 7.68% .****..******************** 37.69 7.54% ................*.*........ 37.64 7.53% .*.**..******************** 37.43 7.49% ........*****.*............ 36.26 7.25% ...............******.***.* 36.01 7.20% ................*****.*..** 35.59 7.12% ...*.**.................... 34.27 6.85% .**........................ 33.14 6.63% ..........*****............ 33.03 6.61% ................*****.*.*.* 31.97 6.39% .***....................... 31.81 6.36% ....*.*.................... 31.66 6.33% ........*.*................ 29.45 5.89% ................*****..**.* 29.13 5.83% ...............******..**** 26.35 5.27% .................*....*.... 26.32 5.26% .****.********************* 26.16 5.23% .......................**.* 25.89 5.18% ............**............. 25.13 5.03% 759 groups at (relative) frequency less than 5% not shown paup> DescribeTrees; Tree description: Unrooted tree(s) rooted using outgroup method Optimality criterion = parsimony Character-status summary: Of 57 total characters: 13 characters are of type 'ord' (Wagner) 44 characters are of type 'unord' All characters have equal weight 3 characters are parsimony-uninformative Number of parsimony-informative characters = 54 Gaps are treated as "missing" Multistate taxa interpreted as polymorphism ("min" values for CI, RI, and RC are minimum- possible character lengths) Character-state optimization: Accelerated transformation (ACCTRAN) Tree 1 (rooted using user-specified outgroup) Branch lengths and linkages for tree #1 (unrooted) Assigned Minimum Maximum Connected branch possible possible Node to node length length length ------------------------------------------------------------------------- Xyela (1)* 52 4 4 4 51 52 1 1 1 Urocerus (2) 51 7 7 7 50 51 1 1 1 Syrista (4) 50 6 6 6 49 50 3 2 3 29 49 1 1 1 Karatavites (5) 29 1 1 1 28 29 2 2 2 Praeratavites (6) 28 1 1 1 Postxiphidria (7) 28 0 0 0 48 49 6 3 7 36 48 5 2 7 Burmorussus (8) 36 4 4 7 35 36 8 5 10 34 35 1 0 6 Paroryssus (9) 34 0 0 2 33 34 2 1 6 Microryssus (10) 33 1 1 3 32 33 8 3 12 31 32 1 1 7 30 31 2 1 7 Orussus (12) 30 0 0 1 Chalinus (13) 30 3 2 3 Baltorussus (15) 31 0 0 0 Minyorussus (14) 32 1 1 1 Praeoryssus (11) 35 0 0 0 47 48 7 3 9 Kuafua (16) 47 3 2 6 46 47 5 4 8 45 46 5 3 9 44 45 7 4 8 42 44 6 4 8 41 42 4 4 9 38 41 6 5 7 37 38 4 4 4 Pristaulacus (17) 37 3 2 3 Gasteruption (18) 37 4 4 5 Evania (19) 38 2 2 3 40 41 4 2 5 Megalyra (21) 40 6 5 7 39 40 4 4 7 Schlettererius (23) 39 9 8 11 Dendrocerus (27) 39 13 13 14 Orthogonalys (20) 42 6 2 6 43 44 3 1 7 Amplicella (24) 43 2 2 2 Xanthopimpla (25) 43 1 1 2 Anthobosca (26) 45 4 4 6 Leptephialtites (22) 46 2 0 2 Xiphydria (3) 52 2 2 2 ------------------------------------------------------------------------- Sum 181 Possible character-state assignments to internal nodes: 1 2 3 4 5 5 Node 123456789012345678901234567890123456789012345678901234567 -------------------------------------------------------------------------- 28 0a0000100101000a12000000000000000000000000000000001000000 29 0a0000100101010a12000000000100000000000000000000001000000 30 101043100100000112000101012201101011011112010000001101212 1 3 31 1010431001000001120001010121011010110a111001000000110a010 1 2 2 2 32 10004310010000011200010101210a1010110a111001000000110a010 1 2 2 2 33 00000310010a00011200000101210010a011001110010000001a0a010 1 2 2 34 00000010010100011200000101210010a011001010010000001a0a010 3 1 2 2 35 00000010010100011200000101210010a011001010010000001a0a010 1 2 2 36 00000010010101011200000000210000a001001010010000001a0a010 1 2 2 37 0a0a21110100111202010020112000122111111113111111011121110 38 0a0a20110100111202010020112100122011111213111111011121110 39 0a0000110100011212010a20122201122111011213a11111121111110 40 0a0000110100011202010a201121011220a1011213a11111121111110 41 0a00001101001112020100201121001220a1011213111111121111110 42 0a00001101001112020100201001000220a1011013111011121111010 1 1 2 2 43 0a0a00111100110222010020102100022011011010110011121110010 44 0a000011a1001102a20100201001000220a1011010110011121110010 1 2 45 0a000010a1001102a2010010100100022001011010010001121110010 46 0a0000100101010112010010100100022001001010010001a01110010 2 2 47 0a00001001010101120000001001000000010010100a0001a01110010 2 1 2 2 48 0a000010010101011200000000010000a0010010100a0000001a00010 49 0a0000100101010a12000000000100000001000000000000001000000 50 0a0000100101010010000000000100000001000000000000000000000 51 0a0000100101000010000000000100000001000000000000000000000 52 0a0000100101000000000000000100000001000000000000000000000 Data matrix and reconstructed states for internal nodes: 1 2 3 4 5 5 Taxon/Node 123456789012345678901234567890123456789012345678901234567 -------------------------------------------------------------------------- Xyela 0-0000-00001000000000000011100000000000000000000000000000 Urocerus 0-0000-01100000010010000000132000001010a01000000000000000 Xiphydria 0-0000-10101000001000000000100000001000000000000000000000 Syrista 0-0000-000010100101-0000100000020001000000000000000000000 Karatavites 0-0?0??0010101??12000000000100000?000000001000??00100??00 Praeratavites 0-0??0-00101?0??02000000000000000?000000000000??00100??00 Postxiphidria 0-0000-0010100??12000000000000000?0000000000000?00100???? Burmorussus 10011??00101011?120000000021000010010010110100??00110???? Paroryssus 0-?????00101?0??12000001-12100100?1100101????0??00100??10 Microryssus 0-??031001???0??12000001-22100100?1100?11???????00100??10 Praeoryssus 0-???0?001???0??12000001-121001???110?1?1???????001?0??10 Orussus 1010431001a0000112000101-12211101011011112010000001101212 Chalinus 1010431001000001121-0101-12232101011011112010000001101212 Minyorussus 10004??00100000112000101-02???101???????1??????000110???? Baltorussus 1-104??00100000112000101-12101101?1?????1??????000110???? Kuafua 0-0??0?1010101??12000000100100001???0010101000?112111??10 Pristaulacus 0-002111010011?202010020-110000221111-111311-111011121110 Gasteruption 0-0121111101111212010020-220001221111-111311-111011121110 Evania 0-0120110100111202010020-121001220111-1213-1-101011121111 Orthogonalys 0-0000-10100111202010020-00100022001011013111011121112212 Megalyra 0-001101011-011202010120-12111122001011213-1-111121111110 Leptephialtites 0-0?00?00101010?12010010100100022?010010a00100??00111??10 Schlettererius 110000-00100011212010020-22201122111011110011011001121110 Amplicella 0-0?00-1110111?-2-010020-021000120110110101100??12111??10 Xanthopimpla 0-0100-11100110-2--11020-02100022011011010110011121110010 Anthobosca 0-0023101100110202010010100100022001011010010001121010012 Dendrocerus 0-0145110100011222111120-222321221111--213-1-111121113312 28 000000100101000112000000000000000000000000000000001000000 29 000000100101010112000000000100000000000000000000001000000 30 101043100100000112000101012211101011011112010000001101212 31 101043100100000112000101012101101011011112010000001101212 32 100043100100000112000101012101101011011112010000001101212 33 000003100100000112000001012100100011001112010000001001210 34 000003100101000112000001012100100011001012010000001001210 35 000000100101000112000001012100100011001012010000001001210 36 000000100101010112000000002100001001001011010000001101210 37 010121110100111202010020112000122111111113111111011121110 38 010120110100111202010020112100122011111213111111011121110 39 010000110100011212010120122201122111011213011111121111110 40 010000110100011202010120112101122011011213011111121111110 41 010000110100111202010020112100122011011213111111121111110 42 010000110100111202010020102100022011011013111011121111110 43 010100111100110222010020102100022011011010110011121110010 44 010000111100110202010020102100022011011010110011121110010 45 010000101100110202010010100100022001011010010001121110010 46 010000100101010212010010100100022001001010010001121110010 47 010000100101010212000000100100001001001010010001121110010 48 000000100101010112000000000100001001001010010000001100010 49 000000100101010112000000000100000001000000000000001000000 50 000000100101010010000000000100000001000000000000000000000 51 000000100101000010000000000100000001000000000000000000000 52 000000100101000000000000000100000001000000000000000000000 Tree length = 184 Consistency index (CI) = 0.4620 Homoplasy index (HI) = 0.5543 CI excluding uninformative characters = 0.4500 HI excluding uninformative characters = 0.5500 Retention index (RI) = 0.7332 Rescaled consistency index (RC) = 0.3387 f value = 2910 f-ratio = 0.4145 (multistate unordered and stepmatrix characters excluded from f-value calculations) /---- Xyela | |/-- Xiphydria || ||/--------- Urocerus 52| ||/-------- Syrista ||| 51| /-- Karatavites || | || 29 /- Praeratavites 50 |\-28 | | \ Postxiphidria | | | | /----- Burmorussus | | | | | /----36 / Praeoryssus \--49 | | | | | \--------35/ Paroryssus | | || | | 34 /- Microryssus | | | | | | \-33 /-- Minyorussus | | | | \------48 \-------32 / Baltorussus | | | | \31 / Orussus | \30 | \---- Chalinus | | /---- Kuafua | | \------47 /-- Leptephialtites | | | | /----- Anthobosca \-----46 | | | /--- Amplicella \----45 /--43 | | \- Xanthopimpla | | \------44 /------- Orthogonalys | | | | /--- Evania \------42 | | /-----38 /---- Pristaulacus | | \---37 | | \----- Gasteruption \---41 | /------- Megalyra | | \---40 /----------- Schlettererius \---39 \---------------- Dendrocerus /----------------------------------------------------------------------------------- Xyela | | /----------------------------------------------------------------------------- Xiphydria | | | | /----------------------------------------------------------------------- Urocerus \----52 | | | /----------------------------------------------------------------- Syrista | | | \----51 | /------------ Karatavites | | | | | /---------------------------------------------29 /------ Praeratavites \----50 | \----28 | | \------ Postxiphidria | | | | /----------------------------------------- Burmorussus | | | | | /----------36 /------------------------------------ Praeoryssus \----49 | | | | | \---35 /------------------------------ Paroryssus | | | | | | \----34 /------------------------ Microryssus | | | | | | \----33 /------------------ Minyorussus | | | | \----48 \----32 /------------ Baltorussus | | | | \----31 /------ Orussus | \----30 | \------ Chalinus | | /----------------------------------------------- Kuafua | | \----47 /----------------------------------------- Leptephialtites | | | | /------------------------------------ Anthobosca \----46 | | | /------ Amplicella \---45 /----------------------43 | | \------ Xanthopimpla | | \----44 /------------------------ Orthogonalys | | | | /------------ Evania \----42 | | /----38 /------ Pristaulacus | | \----37 | | \------ Gasteruption \----41 | /------------ Megalyra | | \----40 /------ Schlettererius \----39 \------ Dendrocerus Character change lists: Character CI Steps Changes --------------------------------------------------------------------- 1 0.333 1 node_36 0 ==> 1 Burmorussus 1 node_33 0 ==> 1 node_32 1 node_39 0 ==> 1 Schlettererius 2 1.000 1 node_48 0 --> 1 node_47 3 1.000 1 node_32 0 ==> 1 node_31 4 0.200 1 node_36 0 ==> 1 Burmorussus 1 node_41 0 --> 1 node_38 1 node_37 1 --> 0 Pristaulacus 1 node_39 0 ==> 1 Dendrocerus 1 node_44 0 --> 1 node_43 5 0.500 1 node_36 0 ==> 1 Burmorussus 1 node_33 0 ==> 4 node_32 1 node_41 0 ==> 2 node_38 1 node_40 0 ==> 1 Megalyra 1 node_39 0 ==> 4 Dendrocerus 1 node_45 0 ==> 2 Anthobosca 6 0.600 1 node_35 0 --> 3 node_34 1 node_38 0 ==> 1 node_37 1 node_40 0 ==> 1 Megalyra 1 node_39 0 ==> 5 Dendrocerus 1 node_45 0 ==> 3 Anthobosca 7 1.000 1 node_40 1 ==> 0 Megalyra 8 0.250 1 node_47 0 ==> 1 Kuafua 1 node_45 0 ==> 1 node_44 1 node_39 1 ==> 0 Schlettererius 1 node_52 0 ==> 1 Xiphydria 9 0.250 1 node_51 0 ==> 1 Urocerus 1 node_46 0 --> 1 node_45 1 node_44 1 --> 0 node_42 1 node_37 0 ==> 1 Gasteruption 10 0.500 1 Xyela 0 <=> 1 node_52 1 node_50 1 ==> 0 Syrista 11 1.000 1 0 --> (01)=a within Orussus 1 node_40 0 ==> 1 Megalyra 12 0.200 1 node_51 1 ==> 0 Urocerus 1 node_34 1 --> 0 node_33 1 node_46 1 ==> 0 node_45 1 node_37 0 ==> 1 Gasteruption 1 node_43 0 ==> 1 Amplicella 13 0.500 1 node_46 0 ==> 1 node_45 1 node_41 1 ==> 0 node_40 14 0.333 1 node_51 0 ==> 1 node_50 1 node_29 1 ==> 0 node_28 1 node_36 1 ==> 0 node_35 15 0.500 1 node_36 0 ==> 1 Burmorussus 1 node_44 0 ==> 1 node_42 16 1.000 1 node_50 0 --> 1 node_49 1 node_48 1 --> 2 node_47 17 0.250 1 node_52 0 ==> 1 node_51 1 node_28 1 ==> 0 Praeratavites 1 node_46 1 --> 0 node_45 1 node_37 0 ==> 1 Gasteruption 1 node_40 0 ==> 1 node_39 1 node_39 1 ==> 2 Dendrocerus 2 node_44 0 ==> 2 node_43 18 1.000 1 node_50 0 ==> 2 node_49 1 node_52 0 ==> 1 Xiphydria 19 0.333 1 node_50 0 ==> 1 Syrista 1 node_30 0 ==> 1 Chalinus 1 node_39 0 ==> 1 Dendrocerus 20 0.500 1 node_51 0 ==> 1 Urocerus 1 node_47 0 ==> 1 node_46 21 0.500 1 node_39 0 ==> 1 Dendrocerus 1 node_43 0 ==> 1 Xanthopimpla 22 0.333 1 node_33 0 ==> 1 node_32 1 node_41 0 --> 1 node_40 1 node_39 1 --> 0 Schlettererius 23 1.000 1 node_47 0 ==> 1 node_46 1 node_45 1 ==> 2 node_44 24 1.000 1 node_36 0 ==> 1 node_35 25 0.500 1 node_50 0 ==> 1 Syrista 1 node_48 0 ==> 1 node_47 26 0.286 1 Xyela 1 <=> 0 node_52 1 node_36 0 ==> 1 node_35 1 node_33 1 ==> 2 Microryssus 1 node_32 1 ==> 0 Minyorussus 1 node_42 0 ==> 1 node_41 1 node_37 1 ==> 2 Gasteruption 1 node_40 1 ==> 2 node_39 27 0.250 1 Xyela 1 <=> 0 node_52 2 node_48 0 ==> 2 node_36 2 node_45 0 --> 2 node_44 1 node_37 2 ==> 1 Pristaulacus 2 node_42 2 --> 0 Orthogonalys 28 0.400 1 node_50 1 ==> 0 Syrista 1 node_29 1 ==> 0 node_28 1 node_31 1 ==> 2 node_30 1 node_38 1 ==> 0 node_37 1 node_40 1 ==> 2 node_39 29 0.400 1 node_51 0 ==> 3 Urocerus 1 node_31 0 --> 1 node_30 1 node_30 1 --> 3 Chalinus 1 node_40 0 ==> 1 Megalyra 1 node_39 0 ==> 3 Dendrocerus 30 0.400 1 node_51 0 ==> 2 Urocerus 1 node_33 0 --> 1 node_32 1 node_30 1 ==> 2 Chalinus 1 node_41 0 ==> 1 node_40 1 node_39 1 ==> 2 Dendrocerus 31 0.333 1 node_36 0 ==> 1 node_35 1 node_42 0 ==> 1 node_41 1 node_37 1 ==> 0 Pristaulacus 32 0.400 2 node_50 0 ==> 2 Syrista 2 node_47 0 ==> 2 node_46 1 node_43 2 ==> 1 Amplicella 33 0.500 1 node_49 0 --> 1 node_48 1 node_36 1 --> 0 node_35 1 node_33 0 --> 1 node_32 1 node_47 1 --> 2 node_46 34 0.500 1 node_38 0 ==> 1 node_37 1 node_40 0 ==> 1 node_39 35 0.250 1 node_36 0 ==> 1 node_35 1 node_45 0 --> 1 node_44 1 node_40 1 --> 0 Megalyra 1 node_42 1 --> 0 Orthogonalys 36 0.500 1 Xyela 0 <=> 1 node_52 1 node_49 1 ==> 0 node_29 37 0.500 1 node_41 0 ==> 1 node_38 1 node_39 0 ==> 1 Dendrocerus 38 0.333 1 node_51 0 ==> 1 Urocerus 1 node_33 0 --> 1 node_32 1 node_46 0 ==> 1 node_45 39 1.000 1 node_49 0 ==> 1 node_48 40 0.600 1 0 --> (01)=a within Urocerus 1 node_34 0 ==> 1 node_33 1 node_42 0 ==> 2 node_41 1 node_38 2 ==> 1 node_37 1 node_39 2 ==> 1 Schlettererius 41 1.000 1 node_49 0 ==> 1 node_48 1 1 --> (01)=a within Leptephialtites 42 0.600 1 node_51 0 ==> 1 Urocerus 1 node_48 0 --> 1 node_36 1 node_36 1 --> 2 node_35 1 node_44 0 ==> 3 node_42 1 node_39 3 ==> 0 Schlettererius 43 0.250 1 node_29 0 ==> 1 Karatavites 1 node_47 0 ==> 1 Kuafua 1 node_45 0 ==> 1 node_44 1 node_41 1 --> 0 node_40 44 0.500 1 node_49 0 --> 1 node_48 1 node_47 1 --> 0 Kuafua 45 1.000 1 node_44 0 ==> 1 node_42 46 0.500 1 node_42 0 ==> 1 node_41 1 node_39 1 ==> 0 Schlettererius 47 0.500 1 node_45 0 ==> 1 node_44 1 node_38 1 ==> 0 Evania 48 1.000 1 node_48 0 ==> 1 node_47 49 0.250 1 node_48 0 --> 1 node_47 1 node_41 1 ==> 0 node_38 1 node_39 1 ==> 0 Schlettererius 1 node_46 1 --> 0 Leptephialtites 50 0.500 1 node_48 0 --> 2 node_47 1 node_41 2 ==> 1 node_38 1 node_39 2 ==> 0 Schlettererius 1 node_46 2 --> 0 Leptephialtites 51 1.000 1 node_50 0 ==> 1 node_49 52 0.250 1 node_49 0 --> 1 node_48 1 node_36 1 --> 0 node_35 1 node_33 0 --> 1 node_32 1 node_45 1 ==> 0 Anthobosca 53 0.667 1 node_48 0 ==> 1 node_47 1 node_41 1 ==> 2 node_38 1 node_39 1 ==> 2 Schlettererius 54 0.600 1 node_48 0 --> 1 node_36 1 node_44 0 ==> 1 node_42 2 node_39 1 ==> 3 Dendrocerus 1 node_42 1 ==> 2 Orthogonalys 55 0.750 1 node_48 0 --> 2 node_36 1 node_44 0 --> 1 node_42 1 node_39 1 ==> 3 Dendrocerus 1 node_42 1 --> 2 Orthogonalys 56 1.000 1 node_49 0 ==> 1 node_48 57 0.400 1 node_33 0 --> 2 node_32 1 node_38 0 ==> 1 Evania 1 node_39 0 ==> 2 Dendrocerus 1 node_42 0 ==> 2 Orthogonalys 1 node_45 0 ==> 2 Anthobosca Apomorphy lists: Branch Character Steps CI Change ------------------------------------------------------------------------ node_52 <-> Xyela 10 1 0.500 1 <=> 0 26 1 0.286 0 <=> 1 27 1 0.250 0 <=> 1 36 1 0.500 1 <=> 0 node_52 --> node_51 17 1 0.250 0 ==> 1 node_51 --> Urocerus 9 1 0.250 0 ==> 1 12 1 0.200 1 ==> 0 20 1 0.500 0 ==> 1 29 1 0.400 0 ==> 3 30 1 0.400 0 ==> 2 38 1 0.333 0 ==> 1 40 1 0.600 0 --> (01)=a 42 1 0.600 0 ==> 1 node_51 --> node_50 14 1 0.333 0 ==> 1 node_50 --> Syrista 10 1 0.500 1 ==> 0 19 1 0.333 0 ==> 1 25 1 0.500 0 ==> 1 28 1 0.400 1 ==> 0 32 2 0.400 0 ==> 2 node_50 --> node_49 16 1 1.000 0 --> 1 18 1 1.000 0 ==> 2 51 1 1.000 0 ==> 1 node_49 --> node_29 36 1 0.500 1 ==> 0 node_29 --> Karatavites 43 1 0.250 0 ==> 1 node_29 --> node_28 14 1 0.333 1 ==> 0 28 1 0.400 1 ==> 0 node_28 --> Praeratavites 17 1 0.250 1 ==> 0 node_49 --> node_48 33 1 0.500 0 --> 1 39 1 1.000 0 ==> 1 41 1 1.000 0 ==> 1 44 1 0.500 0 --> 1 52 1 0.250 0 --> 1 56 1 1.000 0 ==> 1 node_48 --> node_36 27 2 0.250 0 ==> 2 42 1 0.600 0 --> 1 54 1 0.600 0 --> 1 55 1 0.750 0 --> 2 node_36 --> Burmorussus 1 1 0.333 0 ==> 1 4 1 0.200 0 ==> 1 5 1 0.500 0 ==> 1 15 1 0.500 0 ==> 1 node_36 --> node_35 14 1 0.333 1 ==> 0 24 1 1.000 0 ==> 1 26 1 0.286 0 ==> 1 31 1 0.333 0 ==> 1 33 1 0.500 1 --> 0 35 1 0.250 0 ==> 1 42 1 0.600 1 --> 2 52 1 0.250 1 --> 0 node_35 --> node_34 6 1 0.600 0 --> 3 node_34 --> node_33 12 1 0.200 1 --> 0 40 1 0.600 0 ==> 1 node_33 --> Microryssus 26 1 0.286 1 ==> 2 node_33 --> node_32 1 1 0.333 0 ==> 1 5 1 0.500 0 ==> 4 22 1 0.333 0 ==> 1 30 1 0.400 0 --> 1 33 1 0.500 0 --> 1 38 1 0.333 0 --> 1 52 1 0.250 0 --> 1 57 1 0.400 0 --> 2 node_32 --> node_31 3 1 1.000 0 ==> 1 node_31 --> node_30 28 1 0.400 1 ==> 2 29 1 0.400 0 --> 1 node_30 --> Orussus 11 1 1.000 0 --> (01)=a node_30 --> Chalinus 19 1 0.333 0 ==> 1 29 1 0.400 1 --> 3 30 1 0.400 1 ==> 2 node_32 --> Minyorussus 26 1 0.286 1 ==> 0 node_48 --> node_47 2 1 1.000 0 --> 1 16 1 1.000 1 --> 2 25 1 0.500 0 ==> 1 48 1 1.000 0 ==> 1 49 1 0.250 0 --> 1 50 1 0.500 0 --> 2 53 1 0.667 0 ==> 1 node_47 --> Kuafua 8 1 0.250 0 ==> 1 43 1 0.250 0 ==> 1 44 1 0.500 1 --> 0 node_47 --> node_46 20 1 0.500 0 ==> 1 23 1 1.000 0 ==> 1 32 2 0.400 0 ==> 2 33 1 0.500 1 --> 2 node_46 --> node_45 9 1 0.250 0 --> 1 12 1 0.200 1 ==> 0 13 1 0.500 0 ==> 1 17 1 0.250 1 --> 0 38 1 0.333 0 ==> 1 node_45 --> node_44 8 1 0.250 0 ==> 1 23 1 1.000 1 ==> 2 27 2 0.250 0 --> 2 35 1 0.250 0 --> 1 43 1 0.250 0 ==> 1 47 1 0.500 0 ==> 1 node_44 --> node_42 9 1 0.250 1 --> 0 15 1 0.500 0 ==> 1 42 1 0.600 0 ==> 3 45 1 1.000 0 ==> 1 54 1 0.600 0 ==> 1 55 1 0.750 0 --> 1 node_42 --> node_41 26 1 0.286 0 ==> 1 31 1 0.333 0 ==> 1 40 1 0.600 0 ==> 2 46 1 0.500 0 ==> 1 node_41 --> node_38 4 1 0.200 0 --> 1 5 1 0.500 0 ==> 2 37 1 0.500 0 ==> 1 49 1 0.250 1 ==> 0 50 1 0.500 2 ==> 1 53 1 0.667 1 ==> 2 node_38 --> node_37 6 1 0.600 0 ==> 1 28 1 0.400 1 ==> 0 34 1 0.500 0 ==> 1 40 1 0.600 2 ==> 1 node_37 --> Pristaulacus 4 1 0.200 1 --> 0 27 1 0.250 2 ==> 1 31 1 0.333 1 ==> 0 node_37 --> Gasteruption 9 1 0.250 0 ==> 1 12 1 0.200 0 ==> 1 17 1 0.250 0 ==> 1 26 1 0.286 1 ==> 2 node_38 --> Evania 47 1 0.500 1 ==> 0 57 1 0.400 0 ==> 1 node_41 --> node_40 13 1 0.500 1 ==> 0 22 1 0.333 0 --> 1 30 1 0.400 0 ==> 1 43 1 0.250 1 --> 0 node_40 --> Megalyra 5 1 0.500 0 ==> 1 6 1 0.600 0 ==> 1 7 1 1.000 1 ==> 0 11 1 1.000 0 ==> 1 29 1 0.400 0 ==> 1 35 1 0.250 1 --> 0 node_40 --> node_39 17 1 0.250 0 ==> 1 26 1 0.286 1 ==> 2 28 1 0.400 1 ==> 2 34 1 0.500 0 ==> 1 node_39 --> Schlettererius 1 1 0.333 0 ==> 1 8 1 0.250 1 ==> 0 22 1 0.333 1 --> 0 40 1 0.600 2 ==> 1 42 1 0.600 3 ==> 0 46 1 0.500 1 ==> 0 49 1 0.250 1 ==> 0 50 1 0.500 2 ==> 0 53 1 0.667 1 ==> 2 node_39 --> Dendrocerus 4 1 0.200 0 ==> 1 5 1 0.500 0 ==> 4 6 1 0.600 0 ==> 5 17 1 0.250 1 ==> 2 19 1 0.333 0 ==> 1 21 1 0.500 0 ==> 1 29 1 0.400 0 ==> 3 30 1 0.400 1 ==> 2 37 1 0.500 0 ==> 1 54 2 0.600 1 ==> 3 55 1 0.750 1 ==> 3 57 1 0.400 0 ==> 2 node_42 --> Orthogonalys 27 2 0.250 2 --> 0 35 1 0.250 1 --> 0 54 1 0.600 1 ==> 2 55 1 0.750 1 --> 2 57 1 0.400 0 ==> 2 node_44 --> node_43 4 1 0.200 0 --> 1 17 2 0.250 0 ==> 2 node_43 --> Amplicella 12 1 0.200 0 ==> 1 32 1 0.400 2 ==> 1 node_43 --> Xanthopimpla 21 1 0.500 0 ==> 1 node_45 --> Anthobosca 5 1 0.500 0 ==> 2 6 1 0.600 0 ==> 3 52 1 0.250 1 ==> 0 57 1 0.400 0 ==> 2 node_46 --> Leptephialtites 41 1 1.000 1 --> (01)=a 49 1 0.250 1 --> 0 50 1 0.500 2 --> 0 node_52 --> Xiphydria 8 1 0.250 0 ==> 1 18 1 1.000 0 ==> 1 Character diagnostics: Min Tree Max G- Character Range steps steps steps CI RI RC HI fit -------------------------------------------------------------------- 1 1 1 3 6 0.333 0.600 0.200 0.667 0.600 2 1 1 1 1 1.000 0/0 0/0 0.000 1.000 3 1 1 1 3 1.000 1.000 1.000 0.000 1.000 4 1 1 5 5 0.200 0.000 0.000 0.800 0.429 5 3 3 6 11 0.500 0.625 0.313 0.500 0.500 6 3 3 5 8 0.600 0.600 0.360 0.400 0.600 7 1 1 1 1 1.000 0/0 0/0 0.000 1.000 8 1 1 4 10 0.250 0.667 0.167 0.750 0.500 9 1 1 4 5 0.250 0.250 0.063 0.750 0.500 10 1 1 2 2 0.500 0.000 0.000 0.500 0.750 11 1 2 2 2 1.000 0/0 0/0 0.500 0.750 12 1 1 5 12 0.200 0.636 0.127 0.800 0.429 13 1 1 2 7 0.500 0.833 0.417 0.500 0.750 14 1 1 3 12 0.333 0.818 0.273 0.667 0.600 15 1 1 2 7 0.500 0.833 0.417 0.500 0.750 16 2 2 2 12 1.000 1.000 1.000 0.000 1.000 17 2 2 8 11 0.250 0.333 0.083 0.750 0.333 18 2 2 2 4 1.000 1.000 1.000 0.000 1.000 19 1 1 3 3 0.333 0.000 0.000 0.667 0.600 20 1 1 2 12 0.500 0.909 0.455 0.500 0.750 21 1 1 2 2 0.500 0.000 0.000 0.500 0.750 22 1 1 3 6 0.333 0.600 0.200 0.667 0.600 23 2 2 2 11 1.000 1.000 1.000 0.000 1.000 24 1 1 1 7 1.000 1.000 1.000 0.000 1.000 25 1 1 2 4 0.500 0.667 0.333 0.500 0.750 26 2 2 7 17 0.286 0.667 0.190 0.714 0.375 27 2 2 8 22 0.250 0.700 0.175 0.750 0.333 28 2 2 5 9 0.400 0.571 0.229 0.600 0.500 29 2 2 5 5 0.400 0.000 0.000 0.600 0.500 30 2 2 5 7 0.400 0.400 0.160 0.600 0.500 31 1 1 3 12 0.333 0.818 0.273 0.667 0.600 32 2 2 5 23 0.400 0.857 0.343 0.600 0.500 33 2 2 4 15 0.500 0.846 0.423 0.500 0.600 34 1 1 2 4 0.500 0.667 0.333 0.500 0.750 35 1 1 4 12 0.250 0.727 0.182 0.750 0.500 36 1 1 2 4 0.500 0.667 0.333 0.500 0.750 37 1 1 2 4 0.500 0.667 0.333 0.500 0.750 38 1 1 3 9 0.333 0.750 0.250 0.667 0.600 39 1 1 1 7 1.000 1.000 1.000 0.000 1.000 40 2 3 5 10 0.600 0.714 0.429 0.600 0.500 41 1 2 2 8 1.000 1.000 1.000 0.500 0.750 42 3 3 5 10 0.600 0.714 0.429 0.400 0.600 43 1 1 4 7 0.250 0.500 0.125 0.750 0.500 44 1 1 2 8 0.500 0.857 0.429 0.500 0.750 45 1 1 1 2 1.000 1.000 1.000 0.000 1.000 46 1 1 2 5 0.500 0.750 0.375 0.500 0.750 47 1 1 2 7 0.500 0.833 0.417 0.500 0.750 48 1 1 1 8 1.000 1.000 1.000 0.000 1.000 49 1 1 4 7 0.250 0.500 0.125 0.750 0.500 50 2 2 4 10 0.500 0.750 0.375 0.500 0.600 51 1 1 1 4 1.000 1.000 1.000 0.000 1.000 52 1 1 4 10 0.250 0.667 0.167 0.750 0.500 53 2 2 3 16 0.667 0.929 0.619 0.333 0.750 54 3 3 5 9 0.600 0.667 0.400 0.400 0.600 55 3 3 4 9 0.750 0.833 0.625 0.250 0.750 56 1 1 1 6 1.000 1.000 1.000 0.000 1.000 57 2 2 5 6 0.400 0.250 0.100 0.600 0.500 Note: Multistate unordered and/or stepmatrix characters excluded from patristic distance calculations. Patristic distance matrix Below diagonal: Adjusted character distances Above diagonal: Patristic distances 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 1 Xyela - 9 5 12 10 12 11 19 22 24 22 29 30 28 28 22 49 50 45 41 2 Urocerus 9 - 6 11 9 11 10 18 21 23 21 28 29 27 27 21 48 49 44 40 3 Xiphydria 5 6 - 9 7 9 8 16 19 21 19 26 27 25 25 19 46 47 42 38 4 Syrista 10 11 9 - 10 12 11 19 22 24 22 29 30 28 28 22 49 50 45 41 5 Karatavites 8 9 7 10 - 4 3 13 16 18 16 23 24 22 22 16 43 44 39 35 6 Praeratavites 6 9 5 10 4 - 1 15 18 20 18 25 26 24 24 18 45 46 41 37 7 Postxiphidria 7 8 6 9 3 1 - 14 17 19 17 24 25 23 23 17 44 45 40 36 8 Burmorussus 17 18 16 19 13 15 14 - 9 11 9 16 17 15 15 15 42 43 38 34 9 Paroryssus 14 17 15 20 14 14 13 9 - 2 0 7 8 6 6 18 45 46 41 37 10 Microryssus 16 17 17 22 16 16 15 11 2 - 2 7 8 6 6 20 47 48 43 39 11 Praeoryssus 14 17 15 20 14 14 13 9 0 2 - 7 8 6 6 18 45 46 41 37 12 Orussus 21 20 22 27 21 21 20 12 7 7 7 - 1 3 1 25 52 53 48 44 13 Chalinus 22 21 23 26 22 22 21 13 8 8 8 1 - 4 2 26 53 54 49 45 14 Minyorussus 20 17 19 24 18 18 17 9 6 6 6 3 4 - 2 24 51 52 47 43 15 Baltorussus 20 19 21 26 20 20 19 11 6 6 6 1 2 2 - 24 51 52 47 43 16 Kuafua 20 19 15 18 12 16 15 15 18 20 18 23 24 20 22 - 33 34 29 25 17 Pristaulacus 33 32 32 31 31 31 32 26 27 27 27 28 29 27 27 21 - 7 6 16 18 Gasteruption 38 35 37 34 34 36 35 25 26 26 26 29 30 28 28 24 7 - 7 17 19 Evania 33 32 32 33 31 33 34 22 23 23 23 24 25 23 23 21 6 7 - 12 20 Orthogonalys 31 26 26 27 25 27 28 22 27 27 27 28 29 25 27 13 10 17 12 - 21 Megalyra 32 31 31 32 32 32 33 23 24 24 24 22 24 22 22 20 13 16 11 11 22 Leptephialtites 20 17 17 14 14 16 15 14 17 19 17 22 23 19 21 7 18 21 18 12 23 Schlettererius 33 30 34 31 31 33 32 20 21 19 21 18 19 19 19 23 12 11 12 16 24 Amplicella 30 25 27 28 24 28 27 19 22 24 22 25 26 22 24 12 17 14 15 13 25 Xanthopimpla 33 26 30 29 27 31 30 22 25 25 25 26 27 23 25 15 16 15 14 12 26 Anthobosca 24 17 21 20 20 20 21 21 22 22 22 25 26 22 24 12 17 22 17 9 27 Dendrocerus 42 39 41 38 40 42 41 29 30 28 30 27 26 28 28 28 17 14 15 19 Patristic distance matrix (continued) Below diagonal: Adjusted character distances Above diagonal: Patristic distances 21 22 23 24 25 26 27 1 Xyela 46 23 53 38 37 28 54 2 Urocerus 45 22 52 37 36 27 53 3 Xiphydria 43 20 50 35 34 25 51 4 Syrista 46 23 53 38 37 28 54 5 Karatavites 40 17 47 32 31 22 48 6 Praeratavites 42 19 49 34 33 24 50 7 Postxiphidria 41 18 48 33 32 23 49 8 Burmorussus 39 16 46 31 30 21 47 9 Paroryssus 42 19 49 34 33 24 50 10 Microryssus 44 21 51 36 35 26 52 11 Praeoryssus 42 19 49 34 33 24 50 12 Orussus 49 26 56 41 40 31 57 13 Chalinus 50 27 57 42 41 32 58 14 Minyorussus 48 25 55 40 39 30 56 15 Baltorussus 48 25 55 40 39 30 56 16 Kuafua 30 7 37 22 21 12 38 17 Pristaulacus 15 28 22 21 20 23 23 18 Gasteruption 16 29 23 22 21 24 24 19 Evania 11 24 18 17 16 19 19 20 Orthogonalys 13 20 20 13 12 15 21 21 Megalyra - 25 13 18 17 20 14 22 Leptephialtites 17 - 32 17 16 7 33 23 Schlettererius 13 16 - 25 24 27 13 24 Amplicella 18 13 19 - 3 12 26 25 Xanthopimpla 17 14 18 3 - 11 25 26 Anthobosca 16 7 19 12 11 - 28 27 Dendrocerus 14 25 13 20 17 26 - Pairwise homoplasy matrix 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 1 Xyela - 2 Urocerus 0 - 3 Xiphydria 0 0 - 4 Syrista 2 0 0 - 5 Karatavites 2 0 0 0 - 6 Praeratavites 6 2 4 2 0 - 7 Postxiphidria 4 2 2 2 0 0 - 8 Burmorussus 2 0 0 0 0 0 0 - 9 Paroryssus 8 4 4 2 2 4 4 0 - 10 Microryssus 8 6 4 2 2 4 4 0 0 - 11 Praeoryssus 8 4 4 2 2 4 4 0 0 0 - 12 Orussus 8 8 4 2 2 4 4 4 0 0 0 - 13 Chalinus 8 8 4 4 2 4 4 4 0 0 0 0 - 14 Minyorussus 8 10 6 4 4 6 6 6 0 0 0 0 0 - 15 Baltorussus 8 8 4 2 2 4 4 4 0 0 0 0 0 0 - 16 Kuafua 2 2 4 4 4 2 2 0 0 0 0 2 2 4 2 - 17 Pristaulacus 16 16 14 18 12 14 12 16 18 20 18 24 24 24 24 12 - 18 Gasteruption 12 14 10 16 10 10 10 18 20 22 20 24 24 24 24 10 0 - 19 Evania 12 12 10 12 8 8 6 16 18 20 18 24 24 24 24 8 0 0 - 20 Orthogonalys 10 14 12 14 10 10 8 12 10 12 10 16 16 18 16 12 6 0 0 - 21 Megalyra 14 14 12 14 8 10 8 16 18 20 18 27 26 26 26 10 2 0 0 2 22 Leptephialtites 3 5 3 9 3 3 3 2 2 2 2 4 4 6 4 0 10 8 6 8 23 Schlettererius 20 22 16 22 16 16 16 26 28 32 28 38 38 36 36 14 10 12 6 4 24 Amplicella 8 12 8 10 8 6 6 12 12 12 12 16 16 18 16 10 4 8 2 0 25 Xanthopimpla 4 10 4 8 4 2 2 8 8 10 8 14 14 16 14 6 4 6 2 0 26 Anthobosca 4 10 4 8 2 4 2 0 2 4 2 6 6 8 6 0 6 2 2 6 27 Dendrocerus 12 14 10 16 8 8 8 18 20 24 20 30 32 28 28 10 6 10 4 2 Pairwise homoplasy matrix (continued) 21 22 23 24 25 26 27 21 Megalyra - 22 Leptephialtites 8 - 23 Schlettererius 0 16 - 24 Amplicella 0 4 6 - 25 Xanthopimpla 0 2 6 0 - 26 Anthobosca 4 0 8 0 0 - 27 Dendrocerus 0 8 0 6 8 2 -