Karyotype: 2n = 32 and FN = 50. Autosomal complement: eight metacentric and submetacentric pairs large to medium decreasing in size, two subtelocentric pairs (one large and one medium), and five acrocentric pairs (one medium and four small). Sex chromosomes: X, a medium acrocentric; Y, a small acrocentric (Gardner & Patton 1976, pp. 10, Fig. 4C).