TABLE 1. A detailed comparison of the features of the Barwelli species group, including the new species
| Characters | D. barwelli (Beddard, 1886) 1, 2,3, 4 | D. beddardii (Rosa, 1890) 2, 4, 5, 6,7, 8 | D. rara Gates, 1925 6, 9 | D. constricta Gates, 1929 6, 7, 10 | D. spissata Gates, 1930 6, 11 | D. delicata * Gates, 1962 6, 11, 12 | D. vazhania Ardra & Narayanan, sp. nov. |
|---|---|---|---|---|---|---|---|
| Length | 25–44 mm | 24–120 mm | 35–80 mm | 73–110 mm | 90–130 mm | 23–27 mm | 21–39 mm |
| Width | 1.35–2.23 mm | 2–5 mm | 3–4 mm | 3–4½ mm | 5–7 mm | 2 mm | 2.5–4.5 mm |
| Segments | 89–119 | 117–188 | 154 | 148–196 | 159 | ? | 106–146 |
| Colour | Unpigmented | Yellow-brown or white (unpigmented) | White (unpigmented) | White (unpigmented) | Unpigmented | Greyish or blueish | Light grey dorsum, whitish ventrum (unpigmented) |
| Dorsal pores | Absent | Present | Absent | ? | Absent | ? | Absent |
| Clitellum | 10–13 (= 4) | 9–14 (= 6) | 10–13 (= 4) | 10–13, 14 (= 4, 5) | ? | ? | ½9–½14 (= 5) |
| Spermathecal pores | Concealed in furrow at c | Large slits at or median to cd | Median to c | Median to c | At c | Close to c (?) | Large slits at c |
| Male pore location | On small, eversible penes, penes usually retracted male apertures, between bc | On short tubular penes pointing forward (length a little more than half the segment), between bc | Usually on slight teat-like protuberances, between bc | On rather conical porophores, between bc | On conical anteriorly directed porophores, between bc | On anteriomesially directed porophores, between bc | On short tubular penes, usually concealed within aperture; between bc |
| Genital markings/ tumescence | Whitened pads anterior and posterior to secondary male aperture | Whitened areas of epidermal thickenings, in front and behind male pores | Present, usually whitened epidermal thickenings in segments 7, 8, 9, 10 and 11 | Present, whitened areas of epidermal thickenings in segments 10, 11, occasionally on 7, 8, 12. | Present, in segments 8 and circular lips on 10 marked by a deep groove | Absent, epidermis whitened in bc between equators of 10/11 | Present, irregularshaped epidermal thickening, together a flower-like impression, with grooves, in 10 and 11 |
| No. of gizzards (within segments) | Three (in 14–16) | Three to six (in 12–20) | Three to five (in 12–18) | Two to four (in 13–18) | Four to five (in 17–21) | Four (in 12–15) | Three to four (in 12–16) |
| Intestine begins | ? | 24±1 | 23±1 24, 25 | ? | ? | 20±1 | |
| Vas deferens; joins prostate at its | Thin delicate and much coiled; middle | Quite short, S-shaped and not twisted; subentally | Rather short, slender; about at the centre of prostate | Rather short, slender; ental end | Very long, thickened portion in a cluster of long loops, three times larger than testis sac; ental end | Long, in a cluster of loops, as large as testis sac; towards ectal end | Very long, coiled into a large mass, smaller than testis sac; middle portion |
| Prostates | Glandular, erect, cotton-like with penial chamber | Glandular, digitiform, not usually long, gland lack at ectal end | Sessile, circular, discoidal to domes-shaped | Short, erect | Glandular, very long, coiled, looped | Glandular, erect, berry-shaped with fine duct | Glandular, kidney-shaped, dome-shaped muscular ectal end (penial chamber) |
| Prostatic capsule | Digitiform | Digitiform, erect or bent | Small, spindle-shaped in parietes | Club-shaped, narrowed ectally, bent | Digitiform | Digitiform or club-shaped | Narrow, round tubular, Cshaped |
| Spermathecal duct | Long, slender, coiled and contorted | Simple and quiet straight | Duct slightly thickened ectally | Duct slightly thickened ectally | Very long, muscular, ectal portion strongly muscular | Short, straight with only one loop, thickness three to four times towards ectal end | Fairly short, thick with two or three loops |
| Spermathecal atrium | Small, spherical | Small simple widening of ectal end of duct, conical-, pinecone-shaped, mostly confined to parietes | Absent, slight enlargement duct ectally | Absent, slight enlargment duct ectally | Absent | Absent | Small, sessile on body wall, about oval-shaped, longitudinally placed |
| Ovisacs | Present, extends in three to eight segments in 11–18 | Present, in segment 12 | ? | ? | ? | Present, in 12 | Present, extends in segments 12–13 |
| Distribution | Australia, Bahamas, Chile, Cuba, Fiji, India, Indonesia, Jamaica, Mexico, Micronesia, Nicaragua, New Caledonia, Philippines, Puerto Rico, Samoa, São Tomé and Príncipe, Singapore, Taiwan, Tonga | India, Myanmar, South Korea, Thailand, Vietnam | Myanmar | India, Myanmar | Myanmar | Myanmar | India |
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