TABLE 1. Comparison of species of the Philippine sangirensis group and P. sangirensis of Indonesia. Setal counts are given as a pair of numbers separated by a comma, the first of the pair being the count in segment vii, and the second being the number in xx. Setal gaps are represented as D for dorsal gap, either present (+D) or absent (–D), and likewise V for ventral. Reproductive pore spacings are given as the estimated fraction of body circumference. Typhlosole presence accompanied by the segment number of its origin if not in XXVII. Septal presence and absence are indicated by + or –, the first of each pair being for septum 8/9. For bursal pads, those found on the floor of the bursa are represented by a number followed by F (e.g. 4F), and those attached to the roof of the bursa are represented by a number followed by R (e.g. 4R). Missing data are shown by a question mark (?). Data from Aspe & James (2014; 2016; 2017), Hong & James (2008; 2011; 2021), – James (2004b), and Michaelsen (1896). *individuals with spermathecae; some are athecal.
quincunxia diesmosi rugosa monoporata vicinipora baungon-ensis pauci-setosa alba rubida virgata sangirensis apoensis baracatan-ensis camiguin-ensis floresi sibucal-ensis solisi timpoong-ensis malindang-ensis misamis-ensis wati nunezae tigris longi-prostata maculo-dorsalis immanis lago boniao mariae lagunaensis vergrandis concepcionensis riparian-ensis bontoc-ensis banahaw-ensis sp. nov. Pheretima sp 2 aurora sancta sp. nov. taceae sp. nov. apolinarioi sp. nov. madii sp. nov.
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setae vi, xxv 18, 25 28, 38 24, 32 30, 34 52, 40 38, 50 22, 28 38, 36 34, 36 76, 80 ? 38–48, 50–58 28, 38 32, 42 37, 45 70, 90 52, 40 29–43, 50–72 44–48, 50–58 42–51, 43–48 60–70, 52–60 46, 51 38, 48–60 36–40, 38–48 74, 64–74 62–68, 64–68 49, 54 60–78, 62–68 32, 32 46, 52 37–42; 50–54 40–50; 56–68 42, 52 40, 62 46, 64 ? 28, 38 31, 36 39, 45 52, 56
setal gaps D, V Irreg. +D, –V +D, –V +D, –V +D, –V +D, –V +D, –V +D, –V +D, –V –D, –V +D,?V +D,–V +D, –V +D, –V +D, –V +D, –V +D, –V +D, –V –D, +V +D, –V +D, –V +D, –V +D, –V +D, –V +D, –V –D, –V +D, –V +D, –V +D, –V +D, –V +D, +V +D, +V +D, +V +D, +V +D, +V -D, –V –D, +V +D, –V +D, +V +D, +V
male pore setae 0–2 0 0 0 0 8 2 0 4 4–5 6 4–5 4–5 1–4 5 9 3–5 2–5 0 6–7 0 9 0 0–5 2–4 5 0–2 0 1 0 – 2 5–7 4–6 4–7 5 6–7 1 0 0
male pore spacing 0.13 0.15 0.15 0.05 0.04 0.19 0.13 0.10 0.12 0.09 0.17–0.25 0.15 0.22 0.17 0.20 0.17 0.15 0.23 0.11 0.23 0.08 0.22 0.14 0.16 0.13 0.14 0.15 0.03 0.09 0.10 – 0.14 0.11 0.10 0.09–0.13 0.23 0.16 0.13 0.12 0.04
sp. pore spacing 0.16 0.23 0.20 0 0.08 0.38 0.16 0.13 0.20 0.11 0.25–0.28 0.16* 0.32 0.17 0.21 0.21 0.18 0.20 0.16 0.30 0.17 0.28 0.13 0.16 0.09 0.12 0.18–0.24 0.14 0.04 0.03 – – 0.07 0.09 0.09–0.13 0.14 0.17–0.22 0.12 0 0.04
dorsal pores 12/13 12/13 11/12 12/13 13/14 13/14 11/12 13/14 11/12 12/13 ? 12/13 12/13 12/13 12/13 12/13 12/13 12/13 12/13 12/13 12/13 12/13 12/13 11/12 12/13 12/13 12/13 ? 12/13 12/13 13/14 12/13 12/13 12/13 4/5 11/12 12/13 5/6 11/12 12/13
dorsal pigment – brown brown brown brown brown – – red-brown brown stripes ? violet brown brown red-brown red- brown gray brown brown violet brown dark brown black brown gray-brown red - purple yellowbrown red dark violet dark brown violet brown dark violet brown light brown dark brown violet brown brown brown, red brown brown brown brown
equatorial pigment none + + + + + none none + none ? none + none + none + none none none + none none none none none none none + + present present + none none none + none + none
length mm 65–86 128 70 60–70 50 107 115 49 72 290–360 ? 160–350 100 63–122 48–67 >165 68–86 145–225 60–80 55–65 67–75 >116 230–280 37–41 226–235 365 233–315 101–133 55 40 35–75 >89 75–105 68–83 130 90 65 80 33 54–60
intestinal origin ½xvi xvii xvii xvii xvi xv ½xvi xvi xvi xvi ? xv xv xv xv or xvi xiv xv xv or xvi xvi xv xv xv xvi xv xvii xvi xiv xiv xvi xvi xv xvi xvi xvi xvi xvii xvi xvi xvi xvii
typhlosole – + + + + + – + + + ? + + + + + + + + + + + + + + + + + + + + + + + + + + + 33 + 33 +
esoph. lamellae + + + + + + + + + + ? + + + + + + + + + + + + + + + + + ? ? – – ? ? + ? + + ? ?
intestinal vessels ? ? ? 18–20 26 32 34 24 26 32–42 ? 32–38 28 26–30 35 38 29 31–44 30 32 34–38 20–23 56–58 20 50–54 28–32 36–38 33–45 34–36 28–30 24–28 30–35 ? 28–30 30 ? ? ? ? ?
hearts x–xiii x– xiii x–xiii x–xiii xi–xiii x–xiii x–xiii x–xiii x–xiii x–xiii ? x–xiii x– xiii x–xiii x–xiii x–xiii xi–xiii x–xiii xi–xiii x–xiii x–xiii xi–xiii x–xiii x–xiii x–xiii x–xiii x–xiii xi–xiii x–xiii x–xiii x–xiii x–xiii x–xiii x–xiii x–xiii x–xiii x– xiii x–xiii x–xiii x–xiii
septa 8/9/10 +, + +, + +, + +, + +, – +, – +, + –, – +, + +, – ? +, – +, – +, – +, – +, – +, – –, – –, – +, + –, + +, – +, – –, – +, – +, – +, – +, – –, – –, – –,– –,– –, – –, – –, – –, – –, – –, – –, – +, +
bithecate vs mono bi bi bi mono bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi bi mono mono bi bi bi bi bi bi mono bi
thecal segm. vii vii vii vii viii vii vii vii vii viii ? viii* viii viii viii vii viii viii viii vii viii vii viii vii viii viii vii viii vii vii vii viii vii vii vii viii vii vii viii vii
penes – – + + – – + + + + + – – + – – – + + + + + – + + – + – + + + + + + + + + + + +
bursal pads 4F 2F 3F 0 2R 2R 3F 0 0 5F, 4R ? 1R 3R 1R 2R 1R 2R 1F 2R 1R, 2F 1F 1R 2R, 2F 5F 0 2R 2F 2R,1F 1R 0 0 0 0 ring of wedges 4F wedges 5F, 1R 2F oval pads 4F, 1R ring F 5–6F wedges 4–5F wedges