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TABLE 1. Comparison of species of the Philippine sangirensis group and P. sangirensis of Indonesia. Setal counts are given as a pair of numbers separated by a comma, the first of the pair being the count in segment vii, and the second being the number in xx. Setal gaps are represented as D for dorsal gap, either present (+D) or absent (–D), and likewise V for ventral. Reproductive pore spacings are given as the estimated fraction of body circumference. Typhlosole presence accompanied by the segment number of its origin if not in XXVII. Septal presence and absence are indicated by + or –, the first of each pair being for septum 8/9. For bursal pads, those found on the floor of the bursa are represented by a number followed by F (e.g. 4F), and those attached to the roof of the bursa are represented by a number followed by R (e.g. 4R). Missing data are shown by a question mark (?). Data from Aspe & James (2014; 2016; 2017), Hong & James (2008; 2011; 2021), – James (2004b), and Michaelsen (1896). *individuals with spermathecae; some are athecal.

quincunxiadiesmosirugosamonoporataviciniporabaungon-ensispauci-setosaalbarubidavirgatasangirensisapoensisbaracatan-ensiscamiguin-ensisfloresisibucal-ensissolisitimpoong-ensismalindang-ensismisamis-ensiswatinunezaetigrislongi-prostatamaculo-dorsalisimmanislagoboniaomariaelagunaensisvergrandisconcepcionensisriparian-ensisbontoc-ensisbanahaw-ensis sp. nov.Pheretima sp 2 aurorasancta sp. nov.taceae sp. nov.apolinarioi sp. nov.madii sp. nov.
setae vi, xxv18, 2528, 3824, 3230, 3452, 4038, 5022, 2838, 3634, 3676, 80?38–48, 50–5828, 3832, 4237, 4570, 9052, 4029–43, 50–7244–48, 50–5842–51, 43–4860–70, 52–6046, 5138, 48–6036–40, 38–4874, 64–7462–68, 64–6849, 5460–78, 62–6832, 3246, 5237–42; 50–5440–50; 56–6842, 5240, 6246, 64?28, 3831, 3639, 4552, 56
setal gaps D, VIrreg.+D, –V+D, –V+D, –V+D, –V+D, –V+D, –V+D, –V+D, –V–D, –V+D,?V+D,–V+D, –V+D, –V+D, –V+D, –V+D, –V+D, –V–D, +V+D, –V+D, –V+D, –V+D, –V+D, –V+D, –V–D, –V+D, –V+D, –V+D, –V+D, –V+D, +V+D, +V+D, +V+D, +V+D, +V-D, –V–D, +V+D, –V+D, +V+D, +V
male pore setae0–2000082044–564–54–51–4593–52–506–70900–52–450–2010–25–74–64–756–7100
male pore spacing0.130.150.150.050.040.190.130.100.120.090.17–0.250.150.220.170.200.170.150.230.110.230.080.220.140.160.130.140.150.030.090.10–0.140.110.100.09–0.130.230.160.130.120.04
sp. pore spacing0.160.230.2000.080.380.160.130.200.110.25–0.280.16*0.320.170.210.210.180.200.160.300.170.280.130.160.090.120.18–0.240.140.040.03––0.070.090.09–0.130.140.17–0.220.1200.04
dorsal pores12/1312/1311/1212/1313/1413/1411/1213/1411/1212/13?12/1312/1312/1312/1312/1312/1312/1312/1312/1312/1312/1312/1311/1212/1312/1312/13?12/1312/1313/1412/1312/1312/134/511/1212/135/611/1212/13
dorsal pigment–brownbrownbrownbrownbrown––red-brownbrown stripes?violet brownbrownred-brownred- browngray brownbrownviolet browndark brownblackbrowngray-brownred - purpleyellowbrownreddark violetdark brownviolet browndark violetbrownlight browndark brownviolet brownbrownbrown,redbrownbrownbrownbrown
equatorial pigmentnone+++++nonenone+none?none+none+none+nonenonenone+nonenonenonenonenonenonenone++presentpresent+nonenonenone+none+none
length mm65–861287060–70501071154972290–360?160–35010063–12248–67>16568–86145–22560–8055–6567–75>116230–28037–41226–235365233–315101–133554035–75>8975–10568–831309065803354–60
intestinal origin½xvixviixviixviixvixv½xvixvixvixvi?xvxvxvxv or xvixivxvxv or xvixvixvxvxvxvixvxviixvixivxivxvixvixvxvixvixvixvixviixvixvixvixvii
typhlosole–+++++–+++?+++++++++++++++++++++++++++ 33+ 33+
esoph. lamellae++++++++++?+++++++++++++++++??––??+?++??
intestinal vessels???18–20263234242632–42?32–382826–3035382931–44303234–3820–2356–582050–5428–3236–3833–4534–3628–3024–2830–35?28–3030?????
heartsx–xiiix– xiiix–xiiix–xiiixi–xiiix–xiiix–xiiix–xiiix–xiiix–xiii?x–xiiix– xiiix–xiiix–xiiix–xiiixi–xiiix–xiiixi–xiiix–xiiix–xiiixi–xiiix–xiiix–xiiix–xiiix–xiiix–xiiixi–xiiix–xiiix–xiiix–xiiix–xiiix–xiiix–xiiix–xiiix–xiiix– xiiix–xiiix–xiiix–xiii
septa 8/9/10+, ++, ++, ++, ++, –+, –+, +–, –+, ++, –?+, –+, –+, –+, –+, –+, ––, ––, –+, +–, ++, –+, ––, –+, –+, –+, –+, ––, ––, ––,––,––, ––, ––, ––, ––, ––, ––, –+, +
bithecate vs monobibibimonobibibibibibibibibibibibibibibibibibibibibibibibibibimonomonobibibibibibimonobi
thecal segm.viiviiviiviiviiiviiviiviiviiviii?viii*viiiviiiviiiviiviiiviiiviiiviiviiiviiviiiviiviiiviiiviiviiiviiviiviiviiiviiviiviiviiiviiviiviiivii
penes––++––+++++––+–––+++++–++–+–++++++++++++
bursal pads4F2F3F02R2R3F005F, 4R?1R3R1R2R1R2R 1F2R1R, 2F1F1R2R, 2F5F02R2F2R,1F1R0000ring of wedges4F wedges5F, 1R2F oval pads4F, 1Rring F5–6F wedges4–5F wedges

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(2008) Three new earthworms of the genus Pheretima (Oligochaeta: Megascolecidae) from Mt. Makiling, Luzon Island, Philippines. Zootaxa, 1695 (1), 45-52. https://doi.org/10.11646/zootaxa.1695.1.2" }, { "reference": "James, S. W. (2004 b) New species of Amynthas, Pheretima, Pleionogaster (Oligochaeta: Megascolecidae) of the Mt. Kitanglad Range, Mindanao Island, Philippines. Raffles Bulletin of Zoology, Singapore, 52 (2), 289-313." }, { "reference": "Michaelsen, W. (1896) Oligochaten Kukenthal - Ergebnisse einer zool. Forschungsreise in den Molukken und in Borneo. 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