Two new species of Geophilus Leach, 1814, are identified and described from Romania. The examined material is compared with morphologically similar congeners, suggesting the presence of a complex of closely related species within the highly diverse genus Geophilus, characterized by a unique combination of diagnostic morphological and molecular characters. Two nuclear (18S, 28S) and one mitochondrial (COI) markers are sequenced for Geophilus tenuis sp. nov., and COI sequence data are provided for Geophilus captiosus sp. nov. Morphological data, phylogenetic reconstruction, and three molecular species delimitation tools (ASAP, bPTP, and GMYC) confirm their validity, with diagnostic nucleotide combinations, based on the COI barcode, additionally provided for them. Their placement in Geophilus is assessed through the most complete phylogeny of the genus hitherto constructed, using sequence data of 18S, 28S, and COI markers from all species present in public repositories. The extent of potential undescribed species diversity in Geophilus is discussed, highlighting the need for comprehensive sampling of small endogeic members of this genus.
", "publication_date": "2026-04-03", "publisher": "Pensoft Publishers", "references": [ { "reference": "Andersson G, Meidell BA, Scheller U, Djursvoll P, Budd GE, G\u00e4rdenfors U (2005) M\u00e5ngfotingar - Myriapoda - Nationalnyckeln till Sveriges flora och fauna. - ArtDatabanken, Uppsala, SLU: 351 pp." }, { "reference": "Attems C (1895) Die Myriopoden Steiermarks. Sitzungsberichten der Kaiserlichen Akademie der Wissenschaften in Wien 104: 117\u2013238." }, { "reference": "Attems C (1901) Myriopoden. - In: Botanik und Zoologie in \u00d6sterreich w\u00e4hrend der letzten f\u00fcnfzig Jahre. Alfred H\u00f6lder, Wien, 280\u2013288." }, { "reference": "Attems C (1903) Synopsis der Geophiliden. ZoologischeJahrb\u00fccher. Abteilung f\u00fcr Systematik 18: 155\u2013302." }, { "reference": "Attems C (1928) The Myriopoda of South Africa. Annals of the South African Museum 26: 1\u2013431." }, { "reference": "Attems C (1929) Geophilomorpha. Das Tierreich. 52. W. de Gruyter & Co., Berlin/Leipzig, 388 pp." }, { "reference": "Baba \u015eC, Giurginca A, Murariu D (2019) Extent of Sampling Effort of the Romanian Centipedes (Myriapoda: Chilopoda): Issues and Advances. Acta Zoologica Bulgarica 71: 29\u201336." }, { "reference": "Baba \u015eC, Giurginca A, Popovici G, Sarbu SM, Popa OP (2024) A new cave\u2013dwelling species of the genus Geophilus Leach, 1814 (Chilopoda: Geophilomorpha) from the chemoautotrophic\u2013based Movile Cave (Southern Dobrogea, Romania). The European Zoological Journal 91(2): 855\u2013868. https://doi.org/10.1080/24750263.2024.2378828" }, { "reference": "Barber AD (2009) Centipedes. Synopses of the British Fauna (N. S.) 58: 1\u2013228." }, { "reference": "Barber AD, Jones RE (1999) A description of Geophilus proximus C.L. Koch, 1847 (Chilopoda, Geophilomorpha). Bulletin of the British Myriapod Group 15: 19\u201325." }, { "reference": "Berlese A (1903) Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Acari, Miriapodi e Scorpioni Italiani. Fascicolo XCIX. Acari, Myriapoda et Scorpiones huscque in Italia reperta, 99. Padova. https://doi.org/10.5962/bhl.title.1589" }, { "reference": "Blower JG (1961) On some new and little known British centipedes. Annals and Magazine of Natural History 4(39): 183\u2013187. https://doi.org/10.1080/00222936108655800" }, { "reference": "Bonato L, Drago L, Murienne J (2014a) Phylogeny of Geophilomorpha (Chilopoda) inferred from new morphological and molecular evidence. Cladistics 30: 485\u2013507. https://doi.org/10.1111/cla.12060" }, { "reference": "Bonato L, Minelli A, Lopresti M, Cerretti P (2014b) ChiloKey, an interactive identification tool for the geophilomorph centipedes of Europe (Chilopoda, Geophilomorpha). ZooKeys 443: 1\u20139. https://doi.org/10.3897/zookeys.443.7530" }, { "reference": "Bonato L, Minelli A, Spungis V (2005) Geophilomorph centipedes of Latvia. Latvijas Entomologs 42: 5\u201315." }, { "reference": "Bonato L, Edgecombe GD, Lewis JGE, Minelli A, Pereira L, Shelley R, Zapparoli M (2010) A common terminology for the external anatomy of centipedes (Chilopoda). ZooKeys 69: 17\u201351. https://doi.org/10.3897/zookeys.69.737" }, { "reference": "Bonato L, Iorio \u00c9, Minelli A (2011) The centipede genus Clinopodes C. L. Koch, 1847 (Chilopoda, Geophilomorpha, Geophilidae): reassessment of species diversity and distribution, with a new species from the Maritime Alps (France). Zoosystema 33(2): 175\u2013205. https://doi.org/10.5252/z2011n2a3" }, { "reference": "Bonato L, Edgecombe GD, Lewis JG, Minelli A, Pereira LA, Shelley RM, Zapparoli M (2010) A common terminology for the external anatomy of centipedes (Chilopoda). ZooKeys 69: 17\u201351. https://doi.org/10.3897/zookeys.69.737" }, { "reference": "Bonato L, Minelli A (2009) Geophilomorph centipedes in the Mediterranean region: revisiting taxonomy opens new evolutionary vistas. Soil Organisms 81: 489\u2013503." }, { "reference": "Bonato L (2011) Chilopoda \u2013 Taxonomic Overview, Order Geophilomorpha. In: Minelli A (Eds) Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Vol. 1. Brill, Leiden, 407\u2013443. https://doi.org/10.1163/9789004188266" }, { "reference": "Bonato L, Minelli A (2014) Chilopoda Geophilomorpha of Europe: A revised list of species, with taxonomic and nomenclatorial notes. Zootaxa 3770: 1\u2013136. https://doi.org/10.11646/zootaxa.3770.1.1" }, { "reference": "Bouckaert R, Vaughan TG, Barido-Sottani J, Duch\u00eane S, Fourment M, Gavryushkina A, Heled J, Jones G, K\u00fchnert D, De Maio N, Matschiner M, Mendes FK, M\u00fcller NF, Ogilvie HA, du Plessis L, Popinga A, Rambaut A, Rasmussen D, Siveroni I, Suchard MA, Wu C-H, Xie D, Zhang C, Stadler T, Drummond AJ (2019) BEAST 2.5: An advanced software platform for Bayesian evolutionary analysis. PLOS Computational Biology 15(4): e1006650. https://doi.org/10.1371/journal.pcbi.1006650" }, { "reference": "Br\u00f6lemann HW (1909) Mat\u00e9riaux pour servir \u00e0 une faune des myriapodes de France - Feuille des Jeunes Naturalistes 39: 208\u2013212." }, { "reference": "Br\u00f6lemann HW (1924) Trois g\u00e9ophiliens (Myriapodes) nouveaux ou peu connus. Bulletin de la Societe d'Histoire naturelle de Toulouse 52: 14\u201320." }, { "reference": "Br\u00f6lemann HW (1930) \u00c9l\u00e9ments d'une faune des Myriapodes de France - Chilopodes. Faune de France 25: 1\u2013405." }, { "reference": "Castresana J (2000) Selection of Conserved Blocks from Multiple Alignments for Their Use in Phylogenetic Analysis. Molecular Biology and Evolution 17: 540\u2013552. https://doi.org/10.1093/oxfordjournals.molbev.a026334" }, { "reference": "Chalande J (1909) Nouvelle esp\u00e8ce Pyr\u00e9n\u00e9enne du genre Geophilus. Feuille des jeunes naturalistes 39(461): 89\u201392." }, { "reference": "Chamberlin RV (1912) Notes on Geophiloidea from Iowa and some neighbouring states. Canadian Entomologist 44: 65\u201372. https://doi.org/10.4039/Ent4465-3" }, { "reference": "Christian E (1996) Die Erdl\u00e4ufer (Chilopoda-Geophilida) des Wiener Stadtgebietes. Verhandlungen der Zoologisch-Botanischen Gesellschaft in \u00d6sterreich 133: 107\u2013132." }, { "reference": "Crabill REJ (1954) A conspectus of the northeastern North American species of Geophilus (Chilopoda, Geophilomorpha, Geophilidae). Proceedings of the Entomological Society of Washington 56: 172\u2013188." }, { "reference": "Demange JM (1959) Myriapodes Chilopodes de Mad\u00e8re (Mission de M. le Pr. A. Vandel). Revue Fran\u00e7aise d'Entomologie 26: 157\u2013166." }, { "reference": "Doni\u0163\u0103 N, Ivan D, Coldea G, Sandal\u0103 V, Popescu A, Chifu T, Pauc\u0103 Com\u0103nescu M, Mititelu D, Bo\u015fcaiu N (1992) Vegeta\u0163ia Rom\u00e2niei. Editura Tehnic\u0103 Agricol\u0103, Bucure\u015fti, 407 pp." }, { "reference": "Drummond AJ, Rambaut A (2007) BEAST: Bayesian evolutionary analysis by sampling trees. BMC Evolutionary Biology 7: 214. https://doi.org/10.1186/1471-2148-7-214" }, { "reference": "Eason EH (1964) Centipedes of the British Isles. Frederick Warne, London, 294 pp." }, { "reference": "Enghoff H (1971) Geophilus insculptus en hidt il up\u00e5agtet dansk skolopender samt bemaerkninger om G. proximus C. L. Koch (Chilopoda). Entomologiske Meddelelser 39: 119\u2013121." }, { "reference": "Ernst A (2000) Struktur und Verbreitung verschiedener Cuticularsensillen bei Geophilus longicornis LEACH (Chilopoda, Geophilomorpha: Geophilidae). Fragmenta Faunistica 43(Suppl.): 113\u2013129." }, { "reference": "Fedosov A, Achaz G, Gontchar A, Puillandre N (2022) MOLD, a novel software to compile accurate and reliable DNA diagnoses for taxonomic descriptions. Molecular Ecology Resources 22: 2038\u20132053. https://doi.org/10.1111/1755-0998.13590" }, { "reference": "Folmer OM, Black M, Hoeh R, Lutz R, Vrijehoek R (1994) DNA primers for amplification of mithocondrial cytochromec oxidase subunit I from diverse metazoan invertebrates. Molecular Phylogenetics and Evolution 2: 304\u2013313." }, { "reference": "Fujisawa T, Barraclough TG (2013) Delimiting Species Using Single-Locus Data and the Generalized Mixed Yule Coalescent Approach: A Revised Method and Evaluation on Simulated Data Sets. Systematic Biology 62(5): 707\u2013724. https://doi.org/10.1093/sysbio/syt033" }, { "reference": "Fourcroy AFde (1785) Entomologia Parisiensis; sive Catalogus Insectorum quae in Agro Parisiensi reperiuntur. Via et Aedibus Serpentineis, Paris, 544 pp. https://doi.org/10.5962/bhl.title.36528" }, { "reference": "Gava R (2009) Ecologia miriapodelor din p\u0103durile F\u0103get, Z\u0103voi \u015fi Trivale \u2013 Pite\u015fti. Editura Academiei Rom\u00e2ne, Bucure\u015fti, 214 pp." }, { "reference": "Gervais P (1837) \u00c9tudes pour servir \u00e0 l'histoire naturelle des Myriapodes. Annales des Sciences Naturelles, Paris (2)7: 35\u201360." }, { "reference": "Gervais P (1847) Myriapodes. In: Walckenaer CAde, Gervais FLP (Eds) Histoire naturelle des Insectes Apt\u00e8res, Tome 4. Librairie Encyclop\u00e9dique de Roret, Paris, 1\u201357, 210\u2013623. https://doi.org/10.5962/bhl.title.47512" }, { "reference": "Gregory SJ, Barber AD (2010) Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire. Bulletin of the British Myriapod and Isopod Group 24: 2\u201315." }, { "reference": "Guindon S, Dufayard J-F, Lefort V, Anisimova M, Hordijk W, Gascuel O (2010) New algorithms and methods to estimate maximum-likelihood phylogenies: assessing the performance of PhyML 3.0. Systematic Biology 59: 307\u2013321. https://doi.org/10.1093/sysbio/syq010" }, { "reference": "Hammer P (1931) Tusindben (Miriopoda). Denmarks Fauna 35: 1\u2013175." }, { "reference": "Horneland EO, Meidell B (2009) Postembryonic development of Strigamia maritima (Leach, 1817) (Chilopoda: Geophilomorpha, Linotaeniidae) with emphasis on how to separate the different stadia. Soil Organisms 81(3): 373\u2013386. https://soil-organisms.org/index.php/SO/article/view/34" }, { "reference": "Ion C (2016) A Catalogue of the Geophilomorpha Species (Myriapoda: Chilopoda) of Romania. Travaux Du Mus\u00e9um National d'Histoire Naturelle \"Grigore Antipa\" 58(1\u20132): 17\u201332. https://doi.org/10.1515/travmu-2016-0001" }, { "reference": "Iorio E (2006) La faune des Chilopodes du Massif Armoricain. Biologie, liste pr\u00e9liminaire et d\u00e9termination des esp\u00e8ces (Chilopoda). M\u00e9moires de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bourdeaux 7: 72 pp." }, { "reference": "Jawlowski H (1949) Wije (Myriapoda) Bialowieskiego Parku Narodowego-Myriapoda of the National Park in Bialowieza. Annales Universitatis Mariae Curie-Sklodowska, Lublin 4c: 309\u2013323." }, { "reference": "Kaczmarek J (1979) Pareczniki (Chilopoda) Polski. Wydawnictwo Naukowe Uniwersytetu im. A. Mickiewicza. Seria Zoologia 9: 1\u201399." }, { "reference": "Kalyaanamoorthy S, Minh BQ, Wong TKF, von Haeseler A, Jermiin LS (2017) ModelFinder: fast model selection for accurate phylogenetic estimates. Nature Methods 14(6): 587\u2013589. https://doi.org/10.1038/nmeth.4285" }, { "reference": "Katoh K, Rozewicki J, Yamada KD (2019) MAFFT online service: Multiple sequence alignment, interactive sequence choice and visualization. Briefings in Bioinformatics 20(4): 1160\u20131166. https://doi.org/10.1093/bib/bbx108" }, { "reference": "Koch CL (1835) Deutschlands Crustaceen Myriapoden und Arachniden. In: (Panzer) Herrich-Schaeffer's Deutschlands Insecten. Pustet, Regensburg, Heft 136." }, { "reference": "Koch CL (1847) System der Myriapoden. In: Herrich-Sch\u00e4ffer L (Ed.) Kritische Revision der Insectenfauna Deutschlands. Pustet, Regensburg 3: 1\u2013270." }, { "reference": "Kumar S, Stecher G, Li M, Knyaz C, Tamura K (2018) MEGA X: Molecular Evolutionary Genetics Analysis across Computing Platforms. Molecular Biology and Evolution 35(6): 1547\u20131549. https://doi.org/10.1093/molbev/msy096" }, { "reference": "Kuraku S, Zmasek CM, Nishimura O, Katoh K (2013) aLeaves facilitates on-demand exploration of metazoan gene family trees on MAFFT sequence alignment server with enhanced interactivity. Nucleic Acids Research 41(W1): W22\u2013W28. https://doi.org/10.1093/nar/gkt389" }, { "reference": "Latzel R (1880) Die Myriopoden der \u00d6sterreichisch-Ungarischen Monarchie. H\u00f6lder, Wien 1: [XV+]228 pp." }, { "reference": "Leach WE (1814) Crustaceology. In: Brewster D (Ed.) The Edinburgh Encyclopaedia. Blackwood, Edinburgh 7(2): 383\u2013437." }, { "reference": "Lewis JGE (1961) On Schendyla peyerimhoffi Br\u00f6lemann & Ribaut, and Geophilus pusillifrater Verhoeff, two geophilomorph centipedes new to the British Isles. Annals and Magazine of Natural History 4(43): 393\u2013399. https://doi.org/10.1080/00222936108651126" }, { "reference": "Lewis JGE (1962) The ecology, distribution and taxonomy of the centipedes found on the shore in the Plymouth area. Journal of the Marine Biological Association of the United Kingdom 42(3): 655\u2013664. https://doi.org/10.1017/S0025315400054333" }, { "reference": "Lignau N (1933) Einiges \u00fcber die Myriapodenfauna der Ukraine. Entomologicheskoe Obozrenie 25: 134\u2013144." }, { "reference": "Linnaeus C (1758) Systema naturae. Editio Decima. Holmiae: 1\u2013824." }, { "reference": "Matic Z (1972) Clasa Chilopoda, subclasa Epimorpha. Fauna Republicii Socialiste Rom\u00e2nia 6(2). Editura Republicii Socialiste Rom\u00e2nia, Bucure\u015fti, 224 pp. [in Romanian] https://doi.org/10.1016/0016-7185(71)90018-2" }, { "reference": "Meinert F (1870) Myriapoda Musaei Hauniensis: bidrag til Myriapodernes morphologi og systematik. Naturhistorisk Tidsskrift (3)7: 1\u2013128." }, { "reference": "Minelli A (1978) Note tassonomiche su Geophilus insculptus Att. e forme affini (Chilopoda). Animalia 5: 295\u2013305." }, { "reference": "Minh BQ, Nguyen MA, von Haeseler A (2013) Ultrafast approximation for phylogenetic bootstrap. Molecular Biology and Evolution 30(5): 1188\u20131195. https://doi.org/10.1093/molbev/mst024" }, { "reference": "Misioch M (1978) Variation of characters in some geophilid chilopods. Abhandlungen und Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 21\u201322: 55\u201362." }, { "reference": "Newport G (1843) On some new genera of the class Myriapoda. Proceedings of the Zoological Society of London 10(1842): 177\u2013181." }, { "reference": "Nguyen LT, Schmidt HA, von Haeseler A, Minh BQ (2015) IQ-TREE: a fast and effective stochastic algorithm for estimating maximum-likelihood phylogenies. Molecular Biology and Evolution 32(1): 268\u2013274. https://doi.org/10.1093/molbev/msu300" }, { "reference": "Palm\u00e9n E (1949) The Chilopoda of Eastern Fennoscandia. Annales Zoologici Societatis Zoologicae-Botanicae Fennicae 13: 1\u201354." }, { "reference": "Pereira LA (2000) The preparation of centipedes for microscopical examination with particular reference to the Geophilomorpha. Bulletin of the British Myriapod Group 16: 22\u201325." }, { "reference": "Pereira LA (2008) On the identity of Geophilus armatus Silvestri, 1895, a junior synonym of Plateurytion tenebrosus (Meinert, 1886), and other Neotropical members of the genus Plateurytion Attems, 1909, including Plateurytion mauryi n. sp. from the Andes of Argentina (Myriapoda: Chilopoda: Geophilomorpha). International Journal of Myriapodology 1(1): 55\u201396. https://doi.org/10.1163/187525408X316758" }, { "reference": "Pereira LA (2017) A redescription of the large geophilid centipede Chilenophilus corralinus (Attems, 1903) (Myriapoda: Chilopoda: Geophilomorpha), with new distribution records from southern Chile. Studies on Neotropical Fauna and Environment 52(2): 131\u2013145. https://doi.org/10.1080/01650521.2017.1299441" }, { "reference": "Peretti E, Bonato L (2016) Geophilus pygmaeus (Chilopoda: Geophilidae): clarifying morphology, variation and geographic distribution. Zootaxa 4139(4): 499\u2013514. https://doi.org/10.11646/zootaxa.4139.4.3" }, { "reference": "Pocock RI (1891) Descriptions of some new Geophilid\u00e6 in the collection of the British Museum. Annals And Magazine of Natural History 8(45): 215\u2013227. https://doi.org/10.1080/00222939109460423" }, { "reference": "Popovici G (2024) A review of the genus Pachymerium C. L. Koch, 1847 (Geophilomorpha, Geophilidae) from Romania. Zootaxa 5406(4): 588\u2013600. 588\u2013600. https://doi.org/10.11646/zootaxa.5406.4.7" }, { "reference": "Puillandre N, Brouillet S, Achaz G (2021) ASAP: assemble species by automatic partitioning. Molecular Ecology Resources 21(2): 609\u2013620. https://doi.org/10.1111/1755-0998.13281" }, { "reference": "Rambaut A, Drummond AJ, Xie D, Baele G, Suchard MA, Susko E (2018) Posterior Summarization in Bayesian Phylogenetics Using Tracer 1.7. Systematic Biology 67(5): 901\u2013904. https://doi.org:/10.1093/sysbio/syy032" }, { "reference": "Salinas JA (1990) Contribuci\u00f3n al conocimiento de los quil\u00f3podos de Navarra Espana (Myriapoda: Chilopoda). Publicaciones de biologia de la Universidad de Navarra Serie zoologica 20: 1\u201370." }, { "reference": "Silvestri F (1895) Chilopodi e diplopodi raccolti dal capitano G. Bove e dal Prof. L. Balzan nell'America meridionale. Annali del Museo civico di storia naturale di Genova, serie 2 14: 764\u2013783." }, { "reference": "Silvestri F (1896) Nuovi Diplopodi e Chilopodi dell'Italia settentrionale. Bollettino dei Musei di Zoologia ed Anatomia comparata della Regia Universit\u00e0 di Torino 11 : 1\u20135." }, { "reference": "Simaiakis SM, Djursvoll P, Bergersen R (2013) Influence of Climate on Segment Number in Geophilus flavus, a Centipede Species Inhabiting Sognefjord in Western Norway. Annales Zoologici Fennici 50(5): 247\u2013255. https://doi.org/10.5735/085.050.0507" }, { "reference": "Simaiakis SM, Iorio E, Djursvoll P, Meidell BA, Andersson G, Kirkendall LR (2010) A study of the diversity and geographical variation in numbers of leg-bearing segments in centipedes (Chilopoda: Geophilomorpha) in north-western Europe. Biological Journal of the Linnean Society 100: 899\u2013909. https://doi.org/10.1111/j.1095-8312.2010.01467.x" }, { "reference": "Stojanovi\u0107 DZ, Miti\u0107 BM, Gedged AM, Anti\u0107 D\u017d, Makarov SE (2019) Geophilus serbicus sp. nov., a new species from the Balkan Peninsula (Chilopoda: Geophilomorpha: Geophilidae). Zootaxa 4658(3): 556\u2013570. https://doi.org/10.11646/zootaxa.4658.3.7" }, { "reference": "Stojanovi\u0107 DZ, Miti\u0107 BM, Dudi\u0107 BD, Gedged AM, Tomi\u0107 VT, Anti\u0107 D\u017d, Makarov SE (2020) Early development of the centipede Geophilus serbicus (Chilopoda: Geophilomorpha: Geophilidae) from the Balkan Peninsula. Invertebrate Reproduction & Development 64(2): 115\u2013125. https://doi.org/10.1080/07924259.2020.1726514" }, { "reference": "Stojanovi\u0107 DZ, \u0160evi\u0107 M, Makarov SE (2024) A new dwarf schendylid centipede (Chilopoda: Geophilomorpha: Schendylidae) with a low number of legs from Serbia, Balkan Peninsula. Zootaxa 5419(3): 401\u2013418. https://doi.org/10.11646/zootaxa.5419.3.5" }, { "reference": "Talavera G, Castresana J (2007) Improvement of Phylogenies after Removing Divergent and Ambiguously Aligned Blocks from Protein Sequence Alignments. Systematic Biology 56: 564\u2013577. https://doi.org/10.1080/10635150701472164" }, { "reference": "Tamura K, Stecher G, Kumar S (2021) MEGA11: molecular evolutionary genetics analysis version 11. Molecular Biology and Evolution 38(7): 3022\u20133027. https://doi.org/10.1093/molbev/msab120" }, { "reference": "Th\u00e9nard J, Quindroit C, Caro G (2024) Red\u00e9couverte de Geophilus proximus C.L. Koch, 1847 en France (Geophilomorpha, Geophilidae), premi\u00e8re observation pour le d\u00e9partement de la Moselle (R\u00e9gion Grand Est). Bulletin de la Soci\u00e9t\u00e9 d'Histoire naturelle et d'Ethnographie de Colmar 80(15): 107\u2013111. https://doi.org/10.5281/zenodo.14054614" }, { "reference": "Turk FA (1947) Myriapodological Notes II. Northwestern Naturalist, September-December 1947: 226\u2013234." }, { "reference": "Verhoeff KW (1895) Beitr\u00e4ge zur Kenntnis pal\u00e4arktischer Myriopoden. (II. Aufsatz: \u00dcber mitteleurop\u00e4ischen Geophiliden). Archiv f\u00fcr Naturgeschichte 61: 346\u2013356." }, { "reference": "Verhoeff KW (1896) Geophiliden und Scolopendriden aus Portugal und Tabelle europ\u00e4ischer Geophilus-Arten. Zoologischer Anzeiger 19(74\u201379): 81\u201389." }, { "reference": "Verhoeff KW (1898) Beitr\u00e4ge zur Kenntniss pal\u00e4arktischer Myriopoden. VI. Aufsatz: Ueber pal\u00e4arktische Geophiliden. Archiv f\u00fcr Naturgeschichte 64(1): 335\u2013362. https://doi.org/10.5962/bhl.part.6891" }, { "reference": "Verhoeff KW (1925) Chilopoda. In: Bronn HG (Ed.) Klassen und Ordnungen des Tierreiches. Winter, Leipzig 5(2): 725 pp." }, { "reference": "Verhoeff KW (1928) Geophilomorphen-Beitr\u00e4ge und eine Lithobius-Form. Mitteilungen aus dem Zoologischen Museum in Berlin 14: 229\u2013286. https://doi.org/10.1002/mmnz.4830140202" }, { "reference": "Verhoeff KW (1934) Beitr\u00e4ge zur Systematik und Geographie der Chilopoden. Zoologische Jahrb\u00fccher, Abteilung f\u00fcr Systematik, \u00d6kologie und Geographie der Tiere 66: 1\u2013112." }, { "reference": "Verhoeff KW (1938) Chilopoden-Studien, zur Kenntnis der Epimorphen. Zoologische Jahrb\u00fccher, Abteilung f\u00fcr Systematik, \u00d6kologie und Geographie der Tiere 71: 340\u2013385." }, { "reference": "Verhoeff KW (1940) Chilopoden von K\u00e4rnten und Tauern, ihre Beziehungen zu europ\u00e4ischen und mediterranen L\u00e4ndern und \u00fcber allgemeine geographische Verh\u00e4ltnisse. Abhandlungen der Preussischen Akademie der Wissenschaften, Mathematisch Naturwissenschaftliche Klasse 5: 3\u201339." }, { "reference": "Whiting MF (2002) Mecoptera is paraphyletic: multiple genes and phylogeny of Mecoptera and Siphonaptera. Zoologica Scripta 31(1): 93\u2013104. https://doi.org/10.1046/j.0300-3256.2001.00095.x" }, { "reference": "Wieczorek J, Bloom D, Guralnick R, Blum S, D\u00f6ring M, Giovanni R, Robertson T, Vieglais D (2012) Darwin Core: An Evolving Community-Developed Biodiversity Data Standard. PLoS ONE 7(1): e29715. https://doi.org/10.1371/journal.pone.0029715" }, { "reference": "Zalesskaja NT, Titova LP, Golovatch SI (1982) The myriapod fauna (Myriapoda) of the Moscow area. In: Ghilarov MS (Ed.) Soil Invertebrates of the Moscow Region. Nauka, Moskow: 179\u2013200. [in Russian]" }, { "reference": "Zhang J, Kapli P, Pavlidis P, Stamatakis A (2013) A general species delimitation method with applications to phylogenetic placements. Bioinformatics 29(22): 2869\u20132876. https://doi.org/10.1093/bioinformatics/btt499" } ], "related_identifiers": [ { "identifier": "10.3897/zse.102.177928.figure9", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure1", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure10", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure4", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure2", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure3", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure6", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure7", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure8", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure11", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure5", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.figure12", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "image-figure", "title": { "de": "Abbildung", "en": "Figure" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.suppl4", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "other", "title": { "de": "Sonstige", "en": "Other" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.suppl1", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "other", "title": { "de": "Sonstige", "en": "Other" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.suppl3", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "other", "title": { "de": "Sonstige", "en": "Other" } }, "scheme": "doi" }, { "identifier": "10.3897/zse.102.177928.suppl2", "relation_type": { "id": "haspart", "title": { "de": "Umfasst folgenden Teil", "en": "Has part" } }, "resource_type": { "id": "other", "title": { "de": "Sonstige", "en": "Other" } }, "scheme": "doi" }, { "identifier": "10.1080/24750263.2024.2378828", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5962/bhl.title.1589", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/00222936108655800", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1111/cla.12060", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.3897/zookeys.443.7530", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.3897/zookeys.69.737", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5252/z2011n2a3", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.3897/zookeys.69.737", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1163/9789004188266", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.11646/zootaxa.3770.1.1", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1371/journal.pcbi.1006650", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/oxfordjournals.molbev.a026334", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.4039/Ent4465-3", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1186/1471-2148-7-214", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1111/1755-0998.13590", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/sysbio/syt033", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5962/bhl.title.36528", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5962/bhl.title.47512", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/sysbio/syq010", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1515/travmu-2016-0001", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1038/nmeth.4285", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/bib/bbx108", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/molbev/msy096", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/nar/gkt389", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/00222936108651126", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1017/S0025315400054333", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1016/0016-7185(71)90018-2", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/molbev/mst024", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/molbev/msu300", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1163/187525408X316758", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/01650521.2017.1299441", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.11646/zootaxa.4139.4.3", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/00222939109460423", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.11646/zootaxa.5406.4.7", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1111/1755-0998.13281", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/sysbio/syy032", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5735/085.050.0507", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1111/j.1095-8312.2010.01467.x", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.11646/zootaxa.4658.3.7", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/07924259.2020.1726514", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.11646/zootaxa.5419.3.5", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1080/10635150701472164", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/molbev/msab120", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5281/zenodo.14054614", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.5962/bhl.part.6891", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1002/mmnz.4830140202", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1046/j.0300-3256.2001.00095.x", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1371/journal.pone.0029715", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" }, { "identifier": "10.1093/bioinformatics/btt499", "relation_type": { "id": "cites", "title": { "de": "Zitiert", "en": "Cites" } }, "resource_type": { "id": "publication", "title": { "de": "Publikation", "en": "Publication" } }, "scheme": "doi" } ], "resource_type": { "id": "publication-article", "title": { "de": "Zeitschriftenartikel", "en": "Journal article" } }, "rights": [ { "description": { "en": "The Creative Commons Attribution license allows re-distribution and re-use of a licensed work on the condition that the creator is appropriately credited." }, "icon": "cc-by-icon", "id": "cc-by-4.0", "props": { "scheme": "spdx", "url": "https://creativecommons.org/licenses/by/4.0/legalcode" }, "title": { "en": "Creative Commons Attribution 4.0 International" } } ], "subjects": [ { "subject": "Centipedes" }, { "subject": "molecular diagnosis" }, { "subject": "Southeast Europe" }, { "subject": "species delimitation" }, { "subject": "systematics" } ], "title": "Integrative taxonomy uncovers hidden diversity in Romanian Geophilus Leach, 1814 (Geophilomorpha, Geophilidae)" }, "parent": { "access": { "owned_by": { "user": "22270" }, "settings": { "accept_conditions_text": null, "allow_guest_requests": false, "allow_user_requests": false, "secret_link_expiration": 0 } }, "communities": { "default": "c529f97d-f8cb-4c13-a439-9e36891694c2", "entries": [ { "access": { "member_policy": "open", "members_visibility": "public", "record_submission_policy": "open", "review_policy": "open", "visibility": "public" }, "children": { "allow": true }, "created": "2013-11-11T14:14:18+00:00", "custom_fields": {}, "deletion_status": { "is_deleted": false, "status": "P" }, "id": "c529f97d-f8cb-4c13-a439-9e36891694c2", "links": {}, "metadata": { "curation_policy": "The Biodiversity Literature Repository (BLR) is to share publications related to bio-systematics. The goal is to provide
\nFor additional search functionality refindit can be used. This includes also searches in CrossRef, DataCite, PubMed, RefBank, GNUB and Mendeley.
\nUpload guideline
\nA provisional guideline how to upload is here.
\n\n
\n
\n
\n
", "title": "Biodiversity Literature Repository", "type": { "id": "topic" }, "website": "https://www.biolitrepo.org" }, "revision_id": 15, "slug": "biosyslit", "theme": { "brand": "biosyslit", "enabled": true, "style": { "font": { "family": "Arial, sans-serif", "size": "16px", "weight": 600 }, "mainHeaderBackgroundColor": "#FFFFFF", "primaryColor": "#636363", "primaryTextColor": "#FFFFFF", "secondaryColor": "#7eb13e", "secondaryTextColor": "#FFFFFF", "tertiaryColor": "#7eb13e", "tertiaryTextColor": "#FFFFFF" } }, "updated": "2025-04-07T12:05:37.615917+00:00" } ], "ids": [ "c529f97d-f8cb-4c13-a439-9e36891694c2" ] }, "id": "19426500", "pids": {} }, "pids": { "doi": { "identifier": "10.3897/zse.102.177928", "provider": "external" }, "oai": { "identifier": "oai:zenodo.org:19426501", "provider": "oai" } }, "revision_id": 5, "stats": { "all_versions": { "data_volume": 372726464.0, "downloads": 26, "unique_downloads": 22, "unique_views": 30, "views": 31 }, "this_version": { "data_volume": 372726464.0, "downloads": 26, "unique_downloads": 22, "unique_views": 30, "views": 31 } }, "status": "published", "swh": {}, "updated": "2026-04-05T07:31:16.405520+00:00", "versions": { "index": 1, "is_latest": true } }