| Rhytidomes |
Short, thin, papyraceous, dark brown to reddish brown |
Long, thick, dark brown to reddish brown |
Long, thin, orange to dark brown |
| Leaves |
Obovate |
Simple triangular |
Orbicular, obovate |
| Petioles |
0.7–1.0 cm long, lanuginose with sparse filiform glandular trichomes of golden-yellowish resin and a 2 mm long tuft of hairs at the insertion of the petiole to the twig |
0.7–1.2 cm long, lanuginose, without glandular trichomes, with a tuft of hairs 2.4–2.7 m long at the insertion of the petiole to the twig |
0.5–1.5 cm long, sericeous with glandular reddish-yellow resin trichomes, with a tuft of hairs 2.0–3.0 mm long, at the insertion of the petiole to the twig |
| Stipular sheath |
3.0 × 0.8 cm with 2 persistent spurs, covered by sericeous indumentum 2.5–3.0 mm long on the external surface |
2.2–2.3 × 0.4–0.5 cm with 2 deciduous spurs, covered by sericeous indumentum 3.0–4.0 mm long on both sides |
1.5–1.8 cm long with 2 spurs, covered by sericeous indumentum on the external surface |
| Leaflets |
Imparipinnate with 1 pair, oblong-elliptic, adaxial surface sparsely lanuginose dark green, secondary venation inconspicuous, abaxial surface densely lanuginose with glandular filiform hairs of golden-yellowish resin, tridentate emarginate apex. Central, 2.9–4.0 × 0.6–0.8 cm, base obtuse, symmetrical. Lateral, 2.4–3.4 × 0.6–0.8 cm sessile, bases asymmetrical |
Imparipinnate with 1 pair, oblong-elliptic to obovate-elliptic, adaxial surface sparsely lanuginose dark green, secondary venation strongly printed conspicuous, abaxial surface densely lanuginose without glandular trichomes, apex emarginate. Central 3.5–4.5 × 0.8– 1.3 cm, base obtuse, symmetrical. Lateral, 3.0–4.5 × 0.8–1.0 cm, sessile, bases asymmetrical |
Imparipinnate with 1–2 pairs, oblong-lanceolate, adaxial surface glabrescent, light olive green, with inconspicuous secondary venation, abaxial surface densely sericeous without glandular trichomes, apex emarginate. Central 2.8–4.9 × 0.4–1.0 cm, base attenuated, symmetrical. Lateral, 2.3–4.5 × 0.4–0.9 cm, sessile, asymmetric bases strongly auriculate |
| Inflorescence |
4.0–8.0 cm long, lanuginose, semi-pendulous, rigid, with 5–8 flowers |
6.0–7.0 cm long, lanuginose, rigid pendulous, with 6–12 flowers |
8.0–14.0 cm long, sericeous, pendulous, with 6–13 flowers |
| Floral bract |
5–8, concave-cymbiform, 8.2–8.3 × 1.2–1.3 mm, densely lanuginose on the outer surface |
6–12, ovate-elliptic, acuminate, 7.0–9.0 × 2.2–2.7 mm, densely sericeous on the outer surface |
6–13, narrowly triangular, 0.8–1.7 × 0.2–0.3 cm slightly sericeous to glabrescent on the outer surface |
| Sepals |
3–4, mostly 4, concave cymbiform, densely lanuginose on the outer surface |
3–4, ovoid triangle, erect, concave, densely lanuginose on the outer surface |
3-4, rarely 5 ovate, green, densely sericeous on the outer surface |
| Androecium |
18–19 stamens |
6–18 stamens |
17–24 stamens |
| Gynoecium |
Style 1.8–2.0 mm in length |
Style 1.7–1.8 mm in length |
Style 1.5–2.8 mm in length |
| Fruits |
Rounded-ovoid 3.0–3.2 × 3.3–3.5 mm to cylindrical 4.0–4.2 × 1.7–1.8 mm, densely lanuginose, with 7–8 short spines, with blunt apex |
Rounded-ovoid, 3–3.5 × 2.8–3.2 mm, densely lanuginose, with 1–7 short spines with blunt apex. |
Turbinate, 2.5–6.5 × 4.5–10.0 mm, densely sericeous with 10–15 long pointed spines |
| Divergence of lateral leaflets (°) in mature leaves |
30–45° |
75–90° |
65–85° |