| Body length (mm) |
55-86 |
110-340 |
45-170 |
116 |
39-63 |
72-97 |
| Body width (mm) |
2.5-2.8 |
4.0-9.0 |
3.0-6.0 |
5.2 |
1.9-2.6 |
2-2.8 |
| Pigment |
|
|
|
|
|
|
| dorsum |
Purple brown before clitellum, light purple brown after |
Dark brown or purple |
Greenish brown |
Dark chocolate on anterior, grey on other parts |
Dark reddish brown |
No pigment |
| ventrum |
Light purple brown before clitellum, no after |
Dark brown or purple |
No |
Grey |
Light gray on ventrum |
No pogment |
| First dorsal pore |
13/14 |
12/13 |
10/11 or 12/13, usually at 11/12 |
12/13 |
6/7 |
11/12 or 12/13 |
| Clitellum locality |
XIV–XVI |
XIV–XVI |
XIV–XVI, occasionally shorter |
XIV–XVI |
XIV–XVI |
XIV–XVI |
| Setal formula |
AA=1.2-2.0AB, ZZ=1.4-2.0ZY |
|
- |
|
|
|
| Spermathecal pores |
4 pairs, in 5/6-8/9, 0.25C |
4 pairs in 5/6-8/9 or 3 pairs in 6/7-8/9, 0.33C |
4 pairs, 5/6-8/9, 0.33C |
4 pairs, in 5/6-8/9, 0.25C |
4 pairs, 5/6-8/9, ventral, 0.45C |
4 pairs, 5/6-8/9, 0.40C |
| Male pores |
Middle, round, surrounded by 3-4 rhombic ridges, 0.25C |
Round or elliptic |
Small, circular to transverse elliptical di sc,0.24-0.30C |
Roundish glandular area, about 1.5mm in diameter, 0.25C |
Round, smooth, slightly elevated with a male aperture inconspicuous on lateral concave area, 0.23-0.30C |
on a coniform glandular disc surrounded by a round pad, 0.33C |
| Papillae preclitellum |
Four pairs of postsetal genital papillae in VI–IX |
8 papillae just overhead 8 spermathecal pores, 2 pairs preclitellar arranged on VIII and IX |
Paired presetal and/or postsetal in some or all, near spermathecal pores |
Invisible |
presetal and postsetal, widely paired in 7, 8 and 9, number highly variable |
Invisible |
| Papillae postclitellum |
2 presetal crescent indented-topped genital papilla medial of male pores |
2 paired presetal genital on XVIII and IX, 1 pair postsetal genital on XVIII |
Present or absent, occasionally one or more near male pore |
Invisible |
presetal and postsetal, widely paired in XVII, XVIII and XIX, number highly variable |
postsetal, single or paired in XVII, XIX and XX |
| Prostate glands |
XVII–XXII |
Well developed |
XVII–XX, rudimentary or absent |
XVI–XXI |
XVI–XX, wrinkled |
XVI–XX, coarsely lobate |
| Spermathecae |
About 2.4mm long, ampulla slender heart-shaped; duct short |
Ampulla oval or pear-shaped, duct equal to or slightly shorter than ampulla |
Ampulla ovoid |
- |
Ampulla peach-shaped, stalk straight, much shorter than ampulla |
About 1.6mm long; ampulla heart-shaped, gradually slender duct as long as ampulla |
| Diverticulum |
About 2.0mm long, terminal 4/5, swollen, S-shaped twisted seminal chamber |
One-third to half of ampulla, slender stalk and a wider seminal chamber |
Blunt ovioid, straight stalk |
Shorter, seminal chamber ovoid and whitish |
Shorter, seminal chamber rudimentary or absent, straight or slightly bent |
As long as main spermathecal axis, slender, terminal 0.4 dilated into a band shaped chamber |
| Accessory glands |
1 stalked accessory gland observed near the ental part of each spermatheca |
- |
Stalked, coelomic, bound down to parietes or retained within body wall |
Invisible |
round, stalked observed in spermathecal pores and male pores region |
Invisible |