Table 1. Morphometric measurements (average ± standard deviation) of the Ptychadena neumanni complex.
| Species |
Sex |
N |
SVL |
HW |
HL |
SL |
NS |
IND |
EN |
IOD |
ETD |
TD |
ED |
UEW |
FLL |
HAL |
FinDW |
THL |
TL |
FL |
Toe4DW |
MTL |
| P. amharensis |
M |
6 |
36.9 ± 2.7 |
13.9 ± 1.4 |
13.2 ± 0.6 |
5.5 ± 0.3 |
2.8 ± 0.2 |
2.9 ± 0.3 |
2.5 ± 0.3 |
2.1 ± 0.2 |
1.2 ± 0.2 |
2.9 ± 0.3 |
4.2 ± 0.6 |
2.7 ± 0.4 |
6.8 ± 0.3 |
8.3 ± 0.5 |
0.6 ± 0.1 |
17.4 ± 1.2 |
18.8 ± 1.3 |
18.9 ± 0.9 |
0.6 ± 0.1 |
1.6 ± 0.3 |
| F |
7 |
42.5 ± 2 |
13.9 ± 0.9 |
14.2 ± 0.9 |
6.1 ± 0.3 |
3 ± 0.5 |
3.3 ± 0.2 |
2.6 ± 0.3 |
2.4 ± 0.3 |
1.5 ± 0.3 |
2.9 ± 0.3 |
4.3 ± 0.4 |
2.9 ± 0.2 |
7.6 ± 0.6 |
8.5 ± 0.5 |
0.6 ± 0.1 |
19.2 ± 1.3 |
21.2 ± 1.3 |
21.1 ± 1.3 |
0.5 ± 0.2 |
1.7 ± 0.2 |
| P. beka |
M |
9 |
37.9 ± 2.6 |
14.5 ± 1 |
14.2 ± 1.3 |
5.7 ± 0.4 |
2.9 ± 0.4 |
3 ± 0.2 |
2.6 ± 0.3 |
2.7 ± 0.3 |
1.1 ± 0.4 |
2.9 ± 0.2 |
4 ± 0.5 |
2.6 ± 0.4 |
6.7 ± 0.5 |
8.3 ± 0.5 |
0.6 ± 0.1 |
17.6 ± 1.2 |
19.5 ± 1 |
19.5 ± 1 |
0.6 ± 0.1 |
1.7 ± 0.2 |
| F |
8 |
43.2 ± 2.9 |
14.7 ± 1.2 |
15.6 ± 1.3 |
6.7 ± 0.5 |
3.4 ± 0.4 |
3.7 ± 0.3 |
3.2 ± 0.3 |
3.2 ± 0.5 |
1.5 ± 0.2 |
3.2 ± 0.3 |
4.5 ± 0.7 |
2.9 ± 0.3 |
8 ± 0.7 |
9.5 ± 0.5 |
0.6 ± 0.1 |
20.9 ± 1.8 |
24.7 ± 1.9 |
23.6 ± 1.4 |
0.7 ± 0.2 |
1.9 ± 0.1 |
| P. cooperi |
M |
8 |
45 ± 2.7 |
17.7 ± 1.1 |
16.5 ± 1.5 |
7.3 ± 0.5 |
4.2 ± 0.5 |
3.3 ± 0.4 |
2.9 ± 0.4 |
2.9 ± 0.5 |
1.7 ± 0.3 |
3.6 ± 0.4 |
4.9 ± 0.5 |
3 ± 0.3 |
9.7 ± 1.1 |
11.5 ± 1.2 |
0.7 ± 0.2 |
22.5 ± 2.7 |
26.4 ± 1.5 |
25.3 ± 1.5 |
0.7 ± 0.1 |
1.9 ± 0.2 |
| F |
1 |
61.2 |
19.9 |
20.7 |
7.6 |
3.8 |
4 |
3.1 |
3.4 |
2.2 |
4.6 |
5.8 |
4.8 |
11.4 |
12.6 |
0.9 |
29.4 |
31.9 |
31.1 |
1 |
2.5 |
| P. delphina |
M |
6 |
40.1 ± 2.9 |
14.7 ± 0.4 |
14.3 ± 0.8 |
6.2 ± 0.3 |
3 ± 0.2 |
3.2 ± 0.4 |
2.8 ± 0.6 |
2.5 ± 0.3 |
1.3 ± 0.2 |
3 ± 0.4 |
4.2 ± 0.6 |
2.9 ± 0.2 |
8.1 ± 0.5 |
8.9 ± 0.9 |
0.6 ± 0.1 |
18.8 ± 0.8 |
21.4 ± 1.5 |
21.6 ± 2.6 |
0.5 ± 0 |
1.5 ± 0.2 |
| F |
6 |
47.2 ± 2.3 |
15.9 ± 1.4 |
16.3 ± 0.8 |
6.9 ± 0.4 |
3.3 ± 0.3 |
3.4 ± 0.2 |
3.2 ± 0.2 |
3.2 ± 0.3 |
1.6 ± 0.2 |
3.4 ± 0.3 |
5.2 ± 0.4 |
3.3 ± 0.3 |
9.3 ± 0.7 |
10.1 ± 0.6 |
0.6 ± 0.1 |
22.9 ± 1.9 |
25.9 ± 2.2 |
25.7 ± 1.6 |
0.7 ± 0.1 |
1.8 ± 0.1 |
| P. doro |
M |
13 |
35.8 ± 2.2 |
12.6 ± 0.8 |
13.1 ± 1.1 |
5.6 ± 0.4 |
2.7 ± 0.3 |
3.1 ± 0.2 |
2.5 ± 0.2 |
2.8 ± 0.4 |
1.1 ± 0.2 |
2.5 ± 0.3 |
4 ± 0.4 |
2.5 ± 0.2 |
7.1 ± 0.5 |
8.5 ± 0.7 |
0.6 ± 0.2 |
17.5 ± 1.6 |
20.7 ± 1.4 |
21.1 ± 2.2 |
0.5 ± 0.1 |
1.5 ± 0.3 |
| F |
6 |
42.4 ± 5.7 |
14.2 ± 0.9 |
15 ± 0.4 |
6.7 ± 0.5 |
3.4 ± 0.5 |
3.5 ± 0.4 |
2.8 ± 0.5 |
3.2 ± 0.5 |
1.3 ± 0.3 |
2.9 ± 0.4 |
4.6 ± 0.4 |
3 ± 0.4 |
8.9 ± 0.9 |
10.4 ± 1 |
0.6 ± 0.1 |
22.6 ± 2.4 |
26.6 ± 1.1 |
26.9 ± 1.6 |
0.8 ± 0.2 |
1.8 ± 0.3 |
| P. erlangeri |
M |
35 |
34.1 ± 2.5 |
11.7 ± 1 |
12.2 ± 1.3 |
5 ± 0.5 |
2.6 ± 0.3 |
2.7 ± 0.3 |
2.2 ± 0.2 |
2.5 ± 0.4 |
1 ± 0.2 |
2.4 ± 0.3 |
3.5 ± 0.4 |
2.4 ± 0.4 |
6.3 ± 0.6 |
7.2 ± 0.7 |
0.6 ± 0.1 |
15.8 ± 1.3 |
17.9 ± 2 |
18.3 ± 1.8 |
0.5 ± 0.1 |
1.4 ± 0.3 |
| F |
10 |
34.6 ± 1.7 |
11.6 ± 1 |
12.2 ± 1.3 |
5.2 ± 0.5 |
2.7 ± 0.3 |
2.9 ± 0.2 |
2.5 ± 0.2 |
2.5 ± 0.4 |
1 ± 0.2 |
2.3 ± 0.3 |
3.6 ± 0.5 |
2.4 ± 0.3 |
6.2 ± 0.5 |
7.5 ± 0.8 |
0.5 ± 0.1 |
16.9 ± 2.1 |
18.7 ± 2.5 |
19.2 ± 1.8 |
0.5 ± 0.1 |
1.4 ± 0.3 |
| P. goweri |
M |
15 |
42.4 ± 2.7 |
15.7 ± 0.9 |
16.2 ± 1.4 |
6.7 ± 0.5 |
3.1 ± 0.3 |
3.5 ± 0.4 |
3.1 ± 0.4 |
3.2 ± 0.4 |
1.5 ± 0.2 |
3.1 ± 0.4 |
4.3 ± 0.7 |
2.9 ± 0.2 |
8.5 ± 0.9 |
10.2 ± 0.8 |
0.7 ± 0.2 |
21.9 ± 1.5 |
25.4 ± 1.5 |
25.7 ± 1.2 |
0.7 ± 0.1 |
2 ± 0.3 |
| F |
6 |
51.2 ± 4.2 |
17.6 ± 2 |
18.6 ± 2.1 |
7.5 ± 0.4 |
3.7 ± 0.3 |
3.9 ± 0.5 |
3.5 ± 0.3 |
3.5 ± 0.4 |
1.8 ± 0.3 |
3.5 ± 0.4 |
5 ± 0.5 |
3.3 ± 0.2 |
9.8 ± 1 |
11.8 ± 1 |
0.7 ± 0.1 |
25.7 ± 2.7 |
30.4 ± 2.3 |
29.4 ± 2 |
0.8 ± 0.1 |
2.5 ± 0.3 |
| P. harenna |
M |
4 |
36.3 ± 2.3 |
13.6 ± 1.2 |
14.2 ± 1.6 |
5.8 ± 0.3 |
3.1 ± 0.2 |
3.4 ± 0.5 |
2.4 ± 0.3 |
2.4 ± 0.5 |
1.2 ± 0.4 |
3.1 ± 0.6 |
4.8 ± 0.6 |
2.8 ± 0.5 |
7.6 ± 0.4 |
8.1 ± 0.5 |
0.6 ± 0.1 |
18.4 ± 2 |
20.6 ± 1.3 |
20.7 ± 1.7 |
0.5 ± 0.1 |
1.4 ± 0.1 |
| F |
2 |
43.3 ± 0.3 |
15.4 ± 1.2 |
16 ± 0.8 |
7 ± 0.4 |
3.4 ± 0.3 |
3.8 ± 0.1 |
3 ± 0.1 |
4.2 ± 0.2 |
1.4 ± 0.1 |
3.1 ± 0.4 |
4.7 ± 0.1 |
2.7 ± 0.6 |
7.8 ± 0.5 |
9.8 ± 0.6 |
0.8 ± 0.4 |
21.8 ± 0.4 |
24.9 ± 0.4 |
24 ± 0.5 |
0.8 ± 0.3 |
2 ± 0.5 |
| P. levenorum |
M |
8 |
34.3 ± 2.3 |
11.7 ± 1.3 |
11.9 ± 1.1 |
4.6 ± 0.5 |
2.4 ± 0.3 |
2.4 ± 0.3 |
1.8 ± 0.4 |
2.2 ± 0.3 |
1 ± 0.3 |
2.2 ± 0.2 |
3.2 ± 0.4 |
2.2 ± 0.4 |
6.5 ± 1 |
7 ± 0.7 |
0.6 ± 0.1 |
14.2 ± 1.4 |
16.9 ± 1.8 |
18 ± 2 |
0.5 ± 0.1 |
1.6 ± 0.2 |
| F |
4 |
37 ± 2.5 |
12.2 ± 0.9 |
12.1 ± 0.5 |
4.8 ± 0.5 |
2.5 ± 0.3 |
2.7 ± 0.4 |
2.1 ± 0.2 |
2.6 ± 0.3 |
1.3 ± 0.2 |
2.4 ± 0.4 |
3.5 ± 0.6 |
2.5 ± 0.4 |
6.6 ± 0.8 |
7.5 ± 0.5 |
0.5 ± 0.2 |
16 ± 1.6 |
18.9 ± 2.1 |
20.1 ± 2.4 |
0.6 ± 0.1 |
1.8 ± 0.2 |
| P. nana |
M |
12 |
28.3 ± 3.3 |
10.2 ± 1.4 |
10.1 ± 0.7 |
4.2 ± 0.4 |
2.2 ± 0.2 |
2.3 ± 0.2 |
1.8 ± 0.2 |
2.2 ± 0.3 |
0.9 ± 0.1 |
1.8 ± 0.2 |
2.9 ± 0.5 |
2 ± 0.3 |
5 ± 0.6 |
5.6 ± 0.6 |
0.5 ± 0.1 |
11.7 ± 1.4 |
12.7 ± 0.9 |
13.8 ± 1 |
0.5 ± 0.1 |
1.2 ± 0.2 |
| F |
12 |
29.6 ± 3.2 |
9.5 ± 1.3 |
10 ± 0.8 |
4.2 ± 0.4 |
2.3 ± 0.3 |
2.5 ± 0.3 |
1.9 ± 0.2 |
2 ± 0.4 |
1 ± 0.2 |
1.8 ± 0.2 |
3.2 ± 0.4 |
2.1 ± 0.4 |
4.9 ± 0.3 |
5.8 ± 0.6 |
0.5 ± 0.1 |
12.2 ± 1.2 |
12.9 ± 1.1 |
14.3 ± 1.2 |
0.5 ± 0.1 |
1.2 ± 0.2 |
| P. neumanni |
M |
20 |
34.6 ± 2 |
12 ± 1.2 |
13.6 ± 0.8 |
5.6 ± 0.4 |
2.8 ± 0.4 |
3.2 ± 0.3 |
2.6 ± 0.3 |
3.1 ± 0.3 |
1 ± 0.1 |
2.7 ± 0.4 |
3.9 ± 0.2 |
2.3 ± 0.4 |
6.8 ± 0.4 |
8.1 ± 0.6 |
0.6 ± 0.2 |
17.9 ± 1.2 |
20.1 ± 1.1 |
20.6 ± 1.4 |
0.6 ± 0.1 |
1.4 ± 0.2 |
| F |
8 |
45.6 ± 1.4 |
15.2 ± 1.1 |
16.7 ± 1.6 |
7.1 ± 0.4 |
3.6 ± 0.1 |
4 ± 0.4 |
3.2 ± 0.3 |
3.6 ± 0.4 |
1.5 ± 0.3 |
3.4 ± 0.4 |
4.7 ± 0.5 |
3 ± 0.4 |
8.3 ± 0.7 |
9.8 ± 0.7 |
0.7 ± 0.2 |
21.8 ± 1.5 |
26.1 ± 0.8 |
26.4 ± 1.4 |
0.7 ± 0.2 |
1.9 ± 0.3 |
| P. robeensis |
M |
13 |
30.3 ± 0.7 |
10.7 ± 0.7 |
11.1 ± 0.8 |
4.3 ± 0.2 |
2.2 ± 0.2 |
2.4 ± 0.2 |
1.9 ± 0.1 |
2.4 ± 0.3 |
1.1 ± 0.1 |
2 ± 0.2 |
3 ± 0.4 |
2.1 ± 0.2 |
5.4 ± 0.5 |
6.4 ± 0.4 |
0.6 ± 0.2 |
12.4 ± 0.8 |
13.6 ± 0.8 |
15.1 ± 1 |
0.5 ± 0.1 |
1.2 ± 0.3 |
| F |
4 |
31.6 ± 2.9 |
11.1 ± 0.6 |
10.9 ± 0.2 |
4.3 ± 0.4 |
2.2 ± 0.5 |
2.5 ± 0.2 |
2 ± 0.1 |
2.5 ± 0.4 |
1.1 ± 0.2 |
2 ± 0.2 |
3.3 ± 0.1 |
2 ± 0.3 |
5.5 ± 0.5 |
6.5 ± 0.5 |
0.5 ± 0.1 |
12.2 ± 1.1 |
13.9 ± 0.8 |
15.4 ± 1.2 |
0.6 ± 0.1 |
1.3 ± 0.2 |