Table 1. Most important character states of camaenid genera relevant for this study.
References Penis Inner structure of the penis Penial sheath Epiphallus Insertion of retractor muscle Penial caecum Flagellum Additional organ
Bouchetcamaena Thach, 2018 this study long, apically thickened parallel folds and a vestigial verge absent long, cylindrical on distal epiphallus absent long, slender absent
Continental " Chloritis " Sutcharit and Panha (2010), Páll-Gergely et al. (2020) long, apically thickened parallel folds and a large verge absent long, cylindrical on distal epiphallus absent medium length, gradually becoming slender absent
Neotrachia Schileyko, 2018 Schileyko (2018) short, thick longitudinal pilasters, broken into series of tubercles absent swollen, ovoid penis-epiphallus transition absent medium length, gradually becoming slender absent
Trachia Martens, 1860 (based on T. vittata) Schileyko (2003) short, swollen chaotically arranged pilasters covers entire penis rather short, thick incorporated into penial sheath absent rather long, gradually becoming slender -
Trichochloritis Pilsbry, 1891 Collinge (1903), Páll-Gergely and Neubert (2019) long, apically thickened unknown absent long, cylindrical on distal epiphallus moderately long, slender very short, pointed absent
Burmochloritis Godwin-Austen, 1920 Godwin-Austen (1920), unpublished information long, thick, cylindrical wavy folds, verge absent absent long, cylindrical bounds penis and epiphallus at some distance from their junction short, pointed long, slender long, cylindrical, derives from wall of vagina
Satsuma A. Adams, 1868 Wang et al. (2014), Zhang et al. (2020) long, cylindrical wavy folds, verge absent absent long, cylindrical on distal epiphallus well-developed, tapering long to short absent
Sinochloritis M. Wu & Z. Chen, 2019 Wu & Chen, 2019 thick, cylindrical parallel folds, verge absent absent long, cylindrical on distal epiphallus, and also covers proximal part of penis large, internally with "peach shaped epiphallic papilla" long, slender, tapering absent
Bellatrachia Schileyko, 2018 Schileyko (2018), Páll-Gergely and Neubert (2019) long, cylindrical parallel folds absent long, cylindrical penis-epiphallus transition absent thick, somewhat swollen, with slender tip absent
Planispira Beck, 1837 Schileyko (2003) short, apically thickened folds and an ovoid, large verge absent long, cylindrical middle of epiphallus absent short, conical absent