internalTaxonId,scientificName,kingdomName,phylumName,className,orderName,familyName,genusName,speciesName,infraType,infraName,infraAuthority,subpopulationName,authority,taxonomicNotes 10030,Hexanchus griseus,ANIMALIA,CHORDATA,CHONDRICHTHYES,HEXANCHIFORMES,HEXANCHIDAE,Hexanchus,griseus,,,,,"(Bonnaterre, 1788)", 10041,Heosemys annandalii,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,GEOEMYDIDAE,Heosemys,annandalii,,,,,"(Boulenger in Annandale & Robinson, 1903)",Appeared on IUCN Red Lists prior to 2007 under the generic name Hieremys. 132523146,Hubbsina turneri,ANIMALIA,CHORDATA,ACTINOPTERYGII,CYPRINODONTIFORMES,GOODEIDAE,Hubbsina,turneri,,,,,"(de Buen, 1940)","Fernando de Buén described Hubbsina turneri after specimens he collected in August 1940 in the Presa Cointzio, Río Grande de Morelia drainage. In 1983, Uyeno et al. published studies on the Karyotypes of Goodeids and A. Radda attached importance to these results, According to identical Karyotypes, he transfered 1984 this species into the genus Girardinichthys. Though phylogenetic results since the early 2000's (Doadrio and Domínguez, 2004) supported the individuality of the genus Hubbsina, Radda held onto his system, in 2004 raising the stock from the Lago Zacapú to species rank describing it as Girardinichthys ireneae. This stock is now regarded as subpopulation of Hubbsina turneri. According to its affiliation to three drainages, three subpopulations are distinguished: The Río Grande de Morelia subpopulation (type subpopulation), the Laguna Yuriría subpopulation and the Río Angulo subpopulation. The first two subpopulations are regarded Extinct. This species is here considered valid in the genus Hubbsina de Buen, 1940 following Ramírez-Herrejón et al. (2010). This differs from Eschmeyer et al. (2018) which consider it meanwhile valid in the genus Girardinichthys Bleeker, 1860." 10267,Hungerfordia pelewensis,ANIMALIA,MOLLUSCA,GASTROPODA,ARCHITAENIOGLOSSA,DIPLOMMATINIDAE,Hungerfordia,pelewensis,,,,,"Beddome, 1889", 10767,Ictalurus australis,ANIMALIA,CHORDATA,ACTINOPTERYGII,SILURIFORMES,ICTALURIDAE,Ictalurus,australis,,,,,"(Meek, 1904)", 10769,Ictalurus mexicanus,ANIMALIA,CHORDATA,ACTINOPTERYGII,SILURIFORMES,ICTALURIDAE,Ictalurus,mexicanus,,,,,"(Meek, 1904)", 1078,Ambloplites cavifrons,ANIMALIA,CHORDATA,ACTINOPTERYGII,PERCIFORMES,CENTRARCHIDAE,Ambloplites,cavifrons,,,,,"Cope, 1868", 10803,Iguanognathus werneri,ANIMALIA,CHORDATA,REPTILIA,SQUAMATA,COLUBRIDAE,Iguanognathus,werneri,,,,,"Boulenger, 1898",Known from the type specimen only. 132521900,Ilyodon whitei,ANIMALIA,CHORDATA,ACTINOPTERYGII,CYPRINODONTIFORMES,GOODEIDAE,Ilyodon,whitei,,,,,(Meek 1904),"The types of this species were collected by E.W. Nelson from a tributary to the Río Balsas near Yautepec, state of Morelos, at the beginning of the last century and described by Meek as Goodea whitei in 1904. C. Hubbs transfered the species into the genus Balsadichthys, finally Miller and Fitzsimons (1971) into the genus Ilyodon. In 1983, Meyer and Förster described Ilyodon lennoni from the Río Chacembero, and in 2000 Paulo-Maya and Trujillo-Jiménez Ilyodon cortesae from the Río Tacambaro. Here we consider Ilyodon lennoni and Ilyodon cortesae synonyms of Ilyodon whitei following Beltrán-López et al. (2017). According to this paper, five subpopulations can be inferred: The Río Amacuzac subpopulation (type subpopulation), the Río Atoyac subpopulation, the Upper Río Cutzmala subpopulation, the Upper Río Coahuayana subpopulation and the Western Río Balsas subpopulation." 10825,Indotestudo forstenii,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,TESTUDINIDAE,Indotestudo,forstenii,,,,,"(Schlegel & Müller, 1845)", 10950,Pangshura sylhetensis,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,GEOEMYDIDAE,Pangshura,sylhetensis,,,,,"Jerdon, 1870","
Traditionally placed in the genus Kachuga, it was assigned to the subgenus Pangshura by Moll (1987); Pangshura was elevated to genus level by Das (2001).
" 11058,Kubaryia pilikia,ANIMALIA,MOLLUSCA,GASTROPODA,LITTORINIMORPHA,ASSIMINEIDAE,Kubaryia,pilikia,,,,,"Clench, 1948", 11150,Lagocephalus gloveri,ANIMALIA,CHORDATA,ACTINOPTERYGII,TETRAODONTIFORMES,TETRAODONTIDAE,Lagocephalus,gloveri,,,,,"Abe & Tabeta, 1983","Puffers of the genus Lagocephalus are globally distributed in warm shallow waters, however classification at the species level in Lagocephalus has not yet been studied in depth, causing confusion in the status of several species including L. cheesemanii, L. gloveri, L. spadiceus, and L. wheeleri (Matsuura et al. 2011). " 1117,Ameca splendens,ANIMALIA,CHORDATA,ACTINOPTERYGII,CYPRINODONTIFORMES,GOODEIDAE,Ameca,splendens,,,,,"Gartner, 1981","The male Holotype of this species was collected together with the female Allotype and 198 Paratopotypes by Robert R. Miller and James T. Greenbank in March 1955 from the Río Teuchitlán, near the road between HW 70 and Etzatlán. Nine Paratypes were collected by Arthur A. Alcorn from a tributary to the Presa La Vega in February 1949 about 2km ESE of Teuchitlán and ten more by James A. Peters and William B. McIntosh in April of the same year from the Río Ameca about 12km E of the town of Ameca. In recent years (2006), the species has been additionally reported from the Balneario Almoloya, endorheic Laguna Magdalena basin, and the El Molino pond near Cuyacapán, endorheic Laguna de Sayula basin. Affiliated to three distinct drainages, three subpopulations can be inferred. The Río Ameca subpopulation (type subpopulation), the Laguna Magdalena subpopulation and the Laguna de Sayula subpopulation. The last one disappeared from the wild a few years after it had been discovered and is regarded Extinct." 11200,Lamna nasus,ANIMALIA,CHORDATA,CHONDRICHTHYES,LAMNIFORMES,LAMNIDAE,Lamna,nasus,,,,,"(Bonnaterre, 1788)", 11242,Lampsilis rafinesqueana,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lampsilis,rafinesqueana,,,,,"Frierson, 1927", 11249,Lampsilis abrupta,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lampsilis,abrupta,,,,,"(Say, 1831)", 11250,Hamiota altilis,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Hamiota,altilis,,,,,"(Conrad, 1834)", 11256,Obovaria haddletoni,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Obovaria,haddletoni,,,,,"(Athearn, 1964)","This species is very similar in appearance to the sympatric Villosa choctawensis Athearn, 1964, so that the taxonomic status of this species requires verification (NatureServe 2010)." 11257,Lampsilis ornata,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lampsilis,ornata,,,,,"(Conrad, 1835)", 11263,Lampsilis splendida,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lampsilis,splendida,,,,,"(Lea, 1838)", 11361,Lasmigona subviridis,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lasmigona,subviridis,,,,,"Conrad, 1835", 11479,Lemiox rimosus,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Lemiox,rimosus,,,,,"(Rafinesque, 1831)", 11481,Lemmus lemmus,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,CRICETIDAE,Lemmus,lemmus,,,,,"(Linnaeus, 1758)",Will be transferred to family Cricetidae. 11686,Leptodea ochracea,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Leptodea,ochracea,,,,,"(Say, 1817)", 11785,Leptoxis taeniata,ANIMALIA,MOLLUSCA,GASTROPODA,SORBEOCONCHA,PLEUROCERIDAE,Leptoxis,taeniata,,,,,"(Conrad, 1834)", 11797,Lepus castroviejoi,ANIMALIA,CHORDATA,MAMMALIA,LAGOMORPHA,LEPORIDAE,Lepus,castroviejoi,,,,,"Palacios, 1977","Lepus castroviejoi was described relatively recently, being separated from L. granatensis, a split supported by genetic evidence. While the range of L. castroviejoi partially overlaps that of L. granatensis and L. europaeus on the Iberian Peninsula, no hybridization has been detected with these forms. Recent studies have shown that instead, based on ecological, molecular and morphological characteristics, it is most closely related to L. corsicanus; some believe these two are conspecific (Alves et al. 2008, Acevedo et al. 2014, Ballesteros 2018). Additionally, two different occurrences of introgression of mtDNA from L. timidus into L. castroviejoi have been documented – indicating that these two forms have come into contact and hybridized two different times in the past, most recently during the last deglaciation (Melo-Ferreira et al. 2005, 2012; Ballesteros 2018). No subspecies or geographic variation in L. castroviejoi have been described (Flux and Angermann 1990, Hoffmann and Smith 2005, Ballesteros 2018)." 11922,Pleuronaia dolabelloides,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Pleuronaia,dolabelloides,,,,,"(Lea, 1840)", 12124,Lissemys scutata,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,TRIONYCHIDAE,Lissemys,scutata,,,,,"(Peters, 1868)", 12137,Lithognathus lithognathus,ANIMALIA,CHORDATA,ACTINOPTERYGII,PERCIFORMES,SPARIDAE,Lithognathus,lithognathus,,,,,"(Cuvier, 1829)", 12395,Lucania interioris,ANIMALIA,CHORDATA,ACTINOPTERYGII,CYPRINODONTIFORMES,FUNDULIDAE,Lucania,interioris,,,,,"Hubbs & Miller, 1965", 12417,Lutjanus cyanopterus,ANIMALIA,CHORDATA,ACTINOPTERYGII,PERCIFORMES,LUTJANIDAE,Lutjanus,cyanopterus,,,,,"(Cuvier, 1828)", 12695,Malaclemys terrapin,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,EMYDIDAE,Malaclemys,terrapin,,,,,"(Schoepff, 1793)","Seven subspecies are generally accepted (TTWG 2017):Lindholm (1929) recognized Testudo tornieri Siebenrock, 1903 as the type of a new genus, for which he established the name Malacochersus. The genus is monotypic, with only one species currently recognized. Genetic studies have demonstrated that Malacochersus is most closely related to Asian Indotestudo and Eurasian Testudo (Le et al. 2006; Parham et al. 2006; Fritz and Bininda-Emonds 2007). Phylogeographic studies of the species have not yet been realized, but are clearly needed in view of its disjunct distribution. Vernacular names used for the species in East African Kiswahili are Kobe ya Mawe and Kobe kama Chapati.
" 12775,Manouria impressa,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,TESTUDINIDAE,Manouria,impressa,,,,,"(Günther, 1882)", 12776,Mantella aurantiaca,ANIMALIA,CHORDATA,AMPHIBIA,ANURA,MANTELLIDAE,Mantella,aurantiaca,,,,,"Mocquard, 1900","There are populations of ""golden"" mantellas that might belong to other species. The taxonomy of this group is uncertain but the definition of Mantella aurantiaca is clear and there seems to be little genetic subdivision within the species (M. Vences pers. comm.)." 12800,Margaritifera marrianae,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,MARGARITIFERIDAE,Margaritifera,marrianae,,,,,"Johnson, 1983", 12803,Margaritifera hembeli,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,MARGARITIFERIDAE,Margaritifera,hembeli,,,,,"Conrad, 1838", 12835,Marmota marmota,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,SCIURIDAE,Marmota,marmota,,,,,"(Linnaeus, 1758)", 12927,Medionidus parvulus,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Medionidus,parvulus,,,,,"I. Lea, 1860", 12930,Medionidus walkeri,ANIMALIA,MOLLUSCA,BIVALVIA,UNIONIDA,UNIONIDAE,Medionidus,walkeri,,,,,"(Wright, 1897)", 12953,Megalagrion jugorum,ANIMALIA,ARTHROPODA,INSECTA,ODONATA,COENAGRIONIDAE,Megalagrion,jugorum,,,,,"(Perkins, 1899)", 12958,Megalagrion oceanicum,ANIMALIA,ARTHROPODA,INSECTA,ODONATA,COENAGRIONIDAE,Megalagrion,oceanicum,,,,,"McLachlan, 1883", 12959,Megalagrion pacificum,ANIMALIA,ARTHROPODA,INSECTA,ODONATA,COENAGRIONIDAE,Megalagrion,pacificum,,,,,"(McLachlan, 1883)", 12960,Megalagrion xanthomelas,ANIMALIA,ARTHROPODA,INSECTA,ODONATA,COENAGRIONIDAE,Megalagrion,xanthomelas,,,,,"(Selys, 1876)", 12961,Megalagrion oahuense,ANIMALIA,ARTHROPODA,INSECTA,ODONATA,COENAGRIONIDAE,Megalagrion,oahuense,,,,,"(Blackburn, 1884)", 13013,Megupsilon aporus,ANIMALIA,CHORDATA,ACTINOPTERYGII,CYPRINODONTIFORMES,CYPRINODONTIDAE,Megupsilon,aporus,,,,,"Miller & Walters, 1972", 1301,Anhydrophryne rattrayi,ANIMALIA,CHORDATA,AMPHIBIA,ANURA,PYXICEPHALIDAE,Anhydrophryne,rattrayi,,,,,"Hewitt, 1919", 13038,Melanochelys tricarinata,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,GEOEMYDIDAE,Melanochelys,tricarinata,,,,,"(Blyth, 1856)", 13039,Melanochelys trijuga,ANIMALIA,CHORDATA,REPTILIA,TESTUDINES,GEOEMYDIDAE,Melanochelys,trijuga,,,,,"(Schweigger, 1812)","Six or seven subspecies are widely recognised: Melanochelys trijuga trijuga (Schweigger, 1812); M. t. coronata (Anderson, 1879); M. t. edeniana (Theobald, 1876); M. t. indopeninsularis (Annandale ,1913); M. t. parkeri (Deraniyagala, 1939); M. t. thermalis (Lesson, 1830). Subspecies wiroti (Reimann in Nutaphand, 1979) was synonymized with M. t. edeniana. The colour and size is considered as criteria for subspecies differentiation (Deraniyagala 1939, Das 1995, Das and Bhupathy 2009). The head of M. trijuga thermalis is highly ornamented with either red, orange, or yellow spots or blotches. During a study of 76 live M. t. thermalis from 19 different localities in Sri Lanka a high percentage (76.%) of specimens had orange, red, yellow and brown patches on the head. Head ornamentation was common and distinct in young (Abeysekera et al. 2015).Oreobates lehri was initially ascribed to two distant populations, one in Cordillera Vilcabamba, which is now considered typical O. lehri, and one in the Andean hills of the Kosñipata Valley, to which is now assigned to O. gemcare (Padial et al. 2012). This is a restricted concept of O. lehri following the split of the broader concept into this and O. gemcare (Padial et al. 2012).
" 136498,Microtus liechtensteini,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,CRICETIDAE,Microtus,liechtensteini,,,,,"(Wettstein, 1927)","Microtus liechtensteini is part of Microtus multiplex species-complex. Its formerly continuous range in the Alps was split into three parts during the last glaciation which led to the evolution of three species: Microtus liechtensteini (south of the Alps east of the river Adige and Balkan mountains), Microtus multiplex (south of the Alps west of the river Adige), and M. bavaricus (north of the Alps in a small refuge in Bavaria and Northern Tyrol) (Haring et al. 2000, Spitzenberger et al. 2000, Spitzenberger 2002, Musser and Carleton 2005, Martinkova et al. 2007)." 136503,Plecotus sardus,ANIMALIA,CHORDATA,MAMMALIA,CHIROPTERA,VESPERTILIONIDAE,Plecotus,sardus,,,,,"Mucedda, Kiefer, Pidinchedda, & Veith, 2002","Identified as a new species by genetic analyses, this species combines morphological features of both P. austriacus and P. auritus." 136536,Microtus brachycercus,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,CRICETIDAE,Microtus,brachycercus,,,,,"Lehmann, 1961","The validity of this species needs to be confirmed (Wilson and Reeder 2005, G. Amori pers. comm. 2006)." 136680,Myotis alcathoe,ANIMALIA,CHORDATA,MAMMALIA,CHIROPTERA,VESPERTILIONIDAE,Myotis,alcathoe,,,,,"von Helversen & Heller, 2001","Formerly included in Myotis mystacinus (Kuhl, 1817); the species was differentiated (von Helversen et al. 2001) on the base of karyological, genetic and echolocation characters." 136773,Uromys boeadii,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,MURIDAE,Uromys,boeadii,,,,,"Groves & Flannery, 1994", 136814,Sorex antinorii,ANIMALIA,CHORDATA,MAMMALIA,EULIPOTYPHLA,SORICIDAE,Sorex,antinorii,,,,,"Bonaparte, 1840","Formerly known as the Valais chromosome race of the Common Shrew S. araneus (2n = 24/25, FN = 40). Given species rank, diagnosed and reviewed by Brünner et al. (2002). Sorex arunchi may be related, if not conspecific (Hutterer in Wilson and Reeder 2005)." 136816,Apodemus epimelas,ANIMALIA,CHORDATA,MAMMALIA,RODENTIA,MURIDAE,Apodemus,epimelas,,,,,"(Nehring, 1902)",A. mystacinus has been split into two separate species: A. mystacinus and A. epimelas (Wilson and Reeder 2005). 137066,Catharsius rhinoceros,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Catharsius,rhinoceros,,,,,"Klug, 1855","The Oriental and Afrotropical genus Catharsius Hope 1837 is currently represented by 80 species in the Afrotropical region (ScarabNet 2009). It is probable that understanding of the genus, species, and their distributional ranges would benefit from a revision." 137070,Deltochilum icarus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Deltochilum,icarus,,,,,"(Olivier, 1789)","Deltochilum was described in 1822 by Eschscholtz. The main diagnostic characteristic is the presence of short carinae or tubercles at the apex of some elitral interstriae. The genus is composed by eight subgenera: Aganhyboma Kolbe, 1893; Calhyboma Kolbe, 1893; Deltochilum s. str.; Deltohyboma Lane, 1946; Euhyboma Kolbe, 1983; Hybomidium Shipp, 1897; Parahyboma Paulian, 1938; Rubrohyboma Paulian, 1938." 137072,Copris victorini,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,victorini,,,,,"Boheman, 1857","The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009). This species belongs to a group currently represented by six species that is restricted to the southeast coastline and adjoining highlands of South Africa. There has been much taxonomic confusion between these species." 137074,Paraphytus africanus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Paraphytus,africanus,,,,,"Boucomont, 1923","The genus Paraphytus Harold 1877 is shared between the rainforests of both the Afrotropical and Oriental regions, with six species considered valid in the Afrotropical region." 137075,Coptorhina nitefacta,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coptorhina,nitefacta,,,,,"Gillet, 1932","The genus Coptorhina is restricted to the Afrotropical region where it comprises seven species, all of which are specialized to feeding and breeding using buried fragments of basidomycete fungi." 137079,Anomiopus parallelus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,parallelus,,,,,"(Harold, 1862)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137080,Cheironitis indicus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cheironitis,indicus,,,,,"Lansberge, 1875","The genus Cheironitis van Lansberge 1875 is centred on the Palaearctic, western Oriental and Afrotropical regions, and is currently represented by eleven species in the Afrotropical region (ScarabNet 2009). The taxonomy of Cheironitis indicus is in need of revision in view of the claim that it has a disjunct distribution in India, Arabia and southern Africa.
" 137083,Ateuchus latus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,latus,,,,,"(Boucomont, 1928)","Ateuchus is a diverse genus, with approximately 70 species, distributed from the United States to Argentina and Lesser Antilles. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy of identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137084,Onthophagus manya,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,manya,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137086,Canthidium calidum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,calidum,,,,,"Harold, 1880","Canthidium is a diverse, species rich genus, with many species unknown. They are distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems. In addition to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137094,Onitis parvulus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,parvulus,,,,,"Balthasar, 1963","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 8 that is restricted to the tropical eastern highlands (Ferreira 1978)." 137095,Canthon divinator,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,divinator,,,,,"Gistel, 1857","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137097,Coptodactyla lesnei,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coptodactyla,lesnei,,,,,"Paulian, 1933","The genus Coptodactyla Burmeister, 1846 occurs in Australia and Papua New Guinea. The Australian fauna comprises 14 species (Reid 2000, Matthews 1974, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137098,Onitis granulisetosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,granulisetosus,,,,,"Ferreira, 1976","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species is in Onitis Group 11, which shows a distributional range in dry to moist savanna regions of southern Africa. The validity of this species needs to be assessed by comparison with the type material of Onitis aeruginosus Klug 1855, which occurs within the same ecoregion.
" 137099,Anomiopus bonariensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,bonariensis,,,,,"(Bruch, 1925)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137100,Cryptocanthon nebulinus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,nebulinus,,,,,"Howden, 1973","According to Cook (2002), Cryptocanthon is a genus with wide occurrence in the New World tropics. They inhabit the litter of wet forests, both lowland rain forests and higher elevation montane and cloud forests, and occur from Brazil to Mexico. Some species are flightless." 137103,Scarabaeus namibicus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,namibicus,,,,,"Krajcik, 2006","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2005): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Kheper, which is represented in both the Afrotropical and Oriental regions (ScarabNet 2009). The species selected for assessment was Scarabaeus namibicus Krajcik 2006. However, following its selection, S. namibicus was formally synonymized with Scarabeus cupreus (Castelnau 1840) (Deschodt et al. 2011). Hence the review of Scarabaeus (Kheper) cupreus (Castelnau)." 137106,Anisocanthon pygmaeus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anisocanthon,pygmaeus,,,,,"(Gillet, 1911)","This genus was created to group three species (two of them described as Canthon and the third as Deltochilum) and now contains four species described. Currently, its geographic distribution is restricted to the South American continent.
" 137107,Heliocopris faunus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Heliocopris,faunus,,,,,"Boheman, 1857","The genus Heliocopris Hope 1837 is found in the Afrotropical and Oriental regions, with one species shared between the Afrotropical and Palaearctic regions, possibly in a Saharo-Sindian pattern of distribution. Most of the taxa are endemic to the Afrotropical region where 44 species are currently recognized as valid (ScarabNet 2009)." 137108,Ateuchus euchalceus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,euchalceus,,,,,"(Balthasar, 1939)","Ateuchus is a diverse genus, with approximately 70 species, distributed from the United States to Argentina and Lesser Antilles. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy of identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137112,Canthidium luteum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,luteum,,,,,"Balthasar, 1939","Canthidium is a diverse genus, distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137118,Canthon gemellatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,gemellatus,,,,,"Erichson, 1847","
Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137119,Onthophagus lamgalio,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,lamgalio,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137121,Scatonomus xanthopygus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scatonomus,xanthopygus,,,,,"Pereira, 1954","Scatonomus is a small genus, containing only eight described species, of which little taxonomic and biological information is available. The genus is known from Brazil, Argentina, Uruguay and Paraguay, and its species are distributed throughout “Cerrado”, “Pampa” and Atlantic forest." 137122,Epirinus gratus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Epirinus,gratus,,,,,"Péringuey, 1901",The genus Epirinus currently comprises 29 species with a combined distribution restricted primarily to the coolest climates of South Africa (ScarabNet 2009). 137123,Neosisyphus macrorubrus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Neosisyphus,macrorubrus,,,,,"(Paschalidis, 1974)","The genus Neosisyphus Müller 1942 shows a widespread distribution in the Oriental and Afrotropical regions and is currently represented by 22 species in the Afrotropical region (ScarabNet 2009). It is in urgent need of revision. This species has frequently been cited as N. macroruber for reasons of gender in Latin usage, although no formal abridgement of the name has been published. It is a close relative of N. rubrus Paschalidis 1974, which has also been assessed for the IUCN Red List." 137131,Canthon corpulentus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,corpulentus,,,,,"Harold, 1868","A review of Canthon lamproderes by Fernando Vaz-de-Mello will soon be published which places some specimens previously identified as Canthon lamproderes within Canthon corpulentus - this has been taken into account in this assessment.
Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically relatively well known.
The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012).
" 137137,Thyregis relictus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Thyregis,relictus,,,,,"Matthews, 1976","The endemic Australian genus Thyregis Blackburn, 1904 comprises four described and one undescribed species (Matthews 1976, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Matthews (1976) considers Thyregis to be a relict genus and Thyregis relictus or a similar species to be ancestral to Coptodactyla. " 137139,Dicranocara deschodti,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dicranocara,deschodti,,,,,"Frolov & Scholtz, 2003",The genus Dicranocara was discovered in the late 1990s and described only in 2003. It now comprises three species (ScarabNet 2009) recorded from arid climate both on the South African and Namibian sides of the Lower Orange River. 137149,Sulcophanaeus actaeon,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Sulcophanaeus,actaeon,,,,,"(Erichson, 1847)","This genus was recently revised by Edmonds (2000). According to the author, Sulcophanaeus is a Neotropical genus comprising five species groups (faunus, carnifex, auricollis, imperator and menelas). Currently, the genus presents 15 valid species and some infraspecific taxa." 137150,Anomiopus smaragdinus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,smaragdinus,,,,,"(Westwood, 1842)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137151,Bdelyrus grandis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,grandis,,,,,"Cook, 1998","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
" 137154,Scarabaeus galenus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,galenus,,,,,"Westwood, 1847","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2006): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Scarabaeus. This species is currently considered to comprise two varieties, a typically finer punctuate variety that is widespread in southern Africa and a more coarsely punctate variety that occurs in the lowlands of southeast east Mozambique and northeast South Africa and may deserve species status." 137155,Frankenbergerius opacus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Frankenbergerius,opacus,,,,,"Frolov & Scholtz, 2005","The genus Frankenbergerius Balthasar, 1938 has been recently revalidated (Frolov and Scholtz 2005) after being in synonymy with the genus Coptorhina Hope 1830. It currently comprises seven species restricted to the eastern and southwest margins of South Africa (ScarabNet 2009)." 137159,Onthophagus villanuevai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,villanuevai,,,,,"Delgado & Deloya, 1990", 137160,Canthon podagricus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,podagricus,,,,,"Harold, 1868","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.
" 137161,Eodrepanus fastiditus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Eodrepanus,fastiditus,,,,,"(Péringuey, 1901)","This species has recently been transferred from the genus Drepanocerus Kirby 1828 to the new genus Eodrepanus Barbero, Pallestrini & Roggero 2009. It is represented in both the Afrotropical and Oriental regions with five species found in the Afrotropical region currently considered valid. Eodrepanus fastiditus (Péringuey, 1901) is often confused with a close relatives Eodrepanus bechynei (Janssens, 1953) and Eodrepanus parallelus (Raffray, 1877)." 137163,Dichotomius nimuendaju,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,nimuendaju,,,,,"Luederwaldt, 1925","The genus consists of approximately 160 valid species that are distributed throughout Americas. However, the greater diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: (Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia). Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus can be changed.
" 137164,Canthon indigaceus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,indigaceus,,,,,"Harold, 1868","Halffter (1961) proposed three subspecies: C. indigaceus indigaceus, C. indigaceus chaipas and C. indigaceus chevrolati." 137165,Cryptocanthon genieri,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,genieri,,,,,"Cook, 2002","According to Cook (2002), Cryptocanthon is a genus with wide occurrence in the New World tropics. They inhabit the litter of wet forests, both lowland rain forests and higher elevation montane and cloud forests, and occur from Brazil to Mexico. Some species are flightless.
" 137166,Frankenbergerius armatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Frankenbergerius,armatus,,,,,"(Boheman, 1857)","The genus Frankenbergerius Balthasar 1938 has been recently revalidated (Frolov and Scholtz 2005) after being in synonymy with the genus Coptorhina Hope, 1830. It currently comprises seven species restricted to the eastern and southwest margins of South Africa (ScarabNet 2009). Frankenbergerius armatus (Boheman, 1857) comprises two subspecies F. armatus armatus (Boheman, 1857) and F. armatus tuberculatus (Frolov & Scholtz, 2005).
" 137169,Coprophanaeus terrali,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coprophanaeus,terrali,,,,,"Arnaud, 2002","According to Edmonds and Zidek (2010), the Neotropical genus Coprophanaeus Olsoufieff (1924) is currently represented by 38 species, distributed among three subgenera (Megaphanaeus Olsoufieff, Metallophanaeus Olsoufieff, and Coprophanaeus s. str.) and eight species groups.
According to Edmonds and Zidek (2010), C. terrali can be near, or perhaps synonym to C. dardanus. Thus, the authors consider the status of C. terrali uncertain.
Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. Onthophagus lacustris is in Onthophagus Group 7 (d’Orbigny 1913), which may be equivalent to the Palaearctic subgenus Eremonthophagus. Group 7 is widely distributed across the savannas of the Afrotropical region. Onthophagus lacustris may have been confused with two close relatives that are inland shade specialists at higher altitude. One of these other species is O. discretus Péringuey. These two species occurred together at 1,000 m in Kyle Game Reserve in Zimbabwe (Australian CSIRO 1970-1986, unpublished records), but only O. discretus was recorded at 1,100 m near Tshwane / Pretoria where it was a soil generalist as well as a shade specialist (Davis 1996).
" 137177,Onitis aeruginosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,aeruginosus,,,,,"Klug, 1855","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species is in Onitis Group 11, which shows a distributional range primarily in dry to moist savanna regions of southern Africa. In view of the extremely widespread but limited data for this species, an assessment of its extinction risk should await a revision of Group 11 and a consideration of the validity of the various included taxa." 137178,Phanaeus daphnis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Phanaeus,daphnis,,,,,"Harold, 1863", 137183,Cryptocanthon rayonensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,rayonensis,,,,,"Cook, 2002", 137185,Demarziella storeyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Demarziella,storeyi,,,,,"Matthews, 1976","The endemic Australian genus Demarziella Balthasar, 1961 comprises 14 described and two undescribed species (Matthews 1976, Matthews 1986, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137186,Onthophagus pugionatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,pugionatus,,,,,"Fahraeus, 1857","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 24 (d’Orbigny 1913), which is widespread in the forest and savannas of the Afrotropical region.
" 137187,Canthidium viride,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,viride,,,,,"(Lucas, 1857)","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137191,Copris inhalatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,inhalatus,,,,,"Quedenfeldt, 1884","The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009). Copris inhalatus comprises four subspecies, Copris inhalatus inhalatus (Quedenfeldt, 1884), Copris inhalatus grandis Nguyen-Phung & Cambefort, 1986, Copris inhalatus perturbator Péringuey, 1901, and Copris inhalatus santaeluciae Nguyen-Phung & Cambefort, 1986." 137194,Onthophagus fuscivestis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,fuscivestis,,,,,"D'Orbigny, 1902","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 23 (d’Orbigny 1913), which is widespread in the forest, savanna and upland grassland of the Afrotropical region.
" 137195,Canthon simulans,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,simulans,,,,,"(Martinez, 1950)","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.
" 137198,Heliocopris japetus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Heliocopris,japetus,,,,,"Klug, 1855","The genus Heliocopris Hope 1837 is found in the Afrotropical and Oriental regions with one species shared between the Afrotropical and Palaearctic regions, possibly in a Saharo-Sindian pattern of distribution. Most of the taxa are endemic to the Afrotropical region where 44 species are currently recognized as valid (ScarabNet 2009).
" 137201,Onthophagus cavia,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,cavia,,,,,"Boucomont, 1914","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species (ScarabNet 2009). The taxonomical limits of the Oriental subgenus Micronthophagus are not defined, but presumably this taxon includes 10-20 species (ScarabNet 2009)." 137202,Onitis mniszechi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,mniszechi,,,,,"Lansberge, 1875","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 18, which is widespread throughout lowland to upland drier savannas of the Afrotropical region on both deep sands and finer-grained soils." 137205,Canthon coloratus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,coloratus,,,,,"Schmidt, 1922","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137209,Sauvagesinella palustris,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Sauvagesinella,palustris,,,,,"Matthews, 1974","The endemic Australian genus Sauvagesinella Paulian, 1934 comprises three species." 137213,Demarziella eungella,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Demarziella,eungella,,,,,"Matthews, 1976","The endemic Australian genus Demarziella Balthasar, 1961 comprises 14 described and two undescribed species (Matthews 1976, Matthews 1986, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Matthews (1986) ascribed specimens from Iron Range in far north Queensland to this species, but these are now considered to belong to an undescribed species." 137214,Bdelyrus paraensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,paraensis,,,,,"Cook, 1998","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
" 137217,Mimonthophagus limbibasis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Mimonthophagus,limbibasis,,,,,"(d'Orbigny, 1905)","Mimonthophagus Balthasar 1963 currently comprises four species with combined distribution in Madagascar and the west, west-central, and southern Afrotropical region, where the various species occur in forest or open savanna woodland. As this species has been recently transferred from Onthophagus Group 5, the generic status of the other group members should be reviewed. The distribution of these other known group members is restricted to southern and east Africa." 137222,Canthidium erythropterum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,erythropterum,,,,,"(Lucas, 1857)","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137227,Ontherus grandis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,grandis,,,,,"Luederwaldt, 1931","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastem Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus." 137236,Dichotomius dahli,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,dahli,,,,,"(Landin, 1956)","The genus consists of approximately 160 valid species that are distributed throughout the Americas. However, the greatest diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia. Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus may change. For this species, it is likely that it may be synonimized with Homocopris torulosus in the near future.
" 137241,Canthidium angulicolle,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,angulicolle,,,,,"Balthasar, 1939","Canthidium is a diverse, species rich genus, with many species unknown. They are distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems. In addition to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus." 137242,Temnoplectron cooki,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Temnoplectron,cooki,,,,,"Reid & Storey, 2000","The genus Temnoplectron Westwood, 1841 occurs in Australia and Papua New Guinea. The Australian fauna comprises 16 species (Matthews 1974, Reid and Storey 2000, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137247,Scarabaeus laevistriatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,laevistriatus,,,,,"Fairmaire, 1893","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2005): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Kheper that is represented in the Afrotropical and Oriental regions (ScarabNet 2009).
" 137248,Scarabaeus damarensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,damarensis,,,,,"Janssens, 1940","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2006): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus, Scarabeolus, which is primarily restricted to Africa." 137249,Onthophagus bubalus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,bubalus,,,,,"Harold, 1867","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 13 (d'Orbigny 1913), which is restricted to highland regions in west central, northeast, east, and southern Africa." 137250,Sisyphus fasciculatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Sisyphus,fasciculatus,,,,,"Boheman, 1857","The genus Sisyphus Latreille is biogeographically widespread across the Afrotropical, Palaearctic and Oriental regions, with a restricted occurrence in the Central American part of the Neotropical region. It is currently represented by 18 species in the Afrotropical region, but is in urgent need of revision. This species is a distinctive taxon that differs from a close relative in bearing discrete bundles of setae (fascicules) on both the dorsal disc of the prothorax and the elytra as opposed to the elytra only." 137252,Anomiopus howdeni,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,howdeni,,,,,"Canhedo, 2006","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137253,Onthophagus surdus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,surdus,,,,,"Boucomont, 1925","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide (ScarabNet 2009)." 137255,Gyronotus mulanjensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gyronotus,mulanjensis,,,,,"Davis & Scholtz & Harrison, 1999","The genus Gyronotus Lansberge, 1874 is a flightless genus, currently comprising six species that are restricted to forest patches on the east coast of the Afrotropical region from Kenya to South Africa (ScarabNet 2009)." 137266,Onthophagus subocelliger,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,subocelliger,,,,,"Blackburn, 1903","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972; T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Onthophagus waterhousei Gillet, 1927 is a junior synonym of this species (S. Spector pers. comm. 2012)." 137267,Temnoplectron rotundum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Temnoplectron,rotundum,,,,,"Westwood, 1841","The genus Temnoplectron Westwood, 1841 occurs in Australia and Papua New Guinea. The Australian fauna comprises 16 species (Matthews 1974, Reid and Storey 2000, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137272,Onthophagus alquirta,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,alquirta,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137273,Scarabaeus nigroaeneus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,nigroaeneus,,,,,"Boheman, 1857","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2005): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Kheper that is represented in the Afrotropical and Oriental regions (ScarabNet 2009)." 137274,Canthon luctuosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,luctuosus,,,,,"Harold, 1868","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon,Peltecantho, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137280,Onthophagus cyaneoniger,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,cyaneoniger,,,,,"D'Orbigny, 1902","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 23 (d'Orbigny 1913), which is widespread in the forest, savanna and upland grassland of the Afrotropical region." 137281,Pedaria ovata,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,ovata,,,,,"Boucomont, 1922", 137286,Caccobius ferrugineus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Caccobius,ferrugineus,,,,,"(Fåhraeus, 1857)","The genus Caccobius Thomson 1859 is widespread in the Afrotropical, Oriental and Palaearctic regions. In the Afrotropical region, it is currently represented by 42 species that are considered valid (ScarabNet 2009).
" 137293,Deltorrhinum batesi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Deltorrhinum,batesi,,,,,"Harold, 1867","Deltorhinum presents 8 described species, distributed through Amazonian Forest. Currently, the genus has been revised by F. Génier (2010). Might be associated with leafcutter ants." 137294,Canthidium bokermanni,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,bokermanni,,,,,"(Martinez, Halffter & Pereira, 1964)","Canthidium is a diverse, species rich genus, with many species unknown. They are distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems. In addition to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137295,Onthophagus vilis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,vilis,,,,,"Harold, 1877","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012).
" 137296,Epirinus sulcipennis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Epirinus,sulcipennis,,,,,"Boheman, 1857","The genus Epirinus currently comprises 29 species with a combined distribution restricted primarily to the coolest climates of South Africa (ScarabNet 2009). The identity of E. sulcipennis Boheman, 1857 is uncertain as material included in the revision of Epirinus (Scholtz and Howden 1987) comprises at least two different species. However, Scholtz and Howden (1987) state that E. sulcipennis has close relationships with E. validus Péringuey, 1901. Of the material available to this assessor that was cited by Scholtz and Howden (1987) as E. sulcipennis, specimens from 1,450 m on Grootberg (Western Cape) are close in appearance to E. validus. However, material from Queenstown (80 km N) is dissimilar to E. validus but similar to material from the Sani Pass cited as E. sulcipennis by Davis et al. (1999). This Queenstown and Sani Pass material is, in fact, close to E. striatus Scholtz & Howden, 1987. Here, E. sulcipennis is assumed to be represented by the material from Grootberg in which the interstriae show obtuse angular ridges compared to the non-ridged interstriae of E. validus." 137297,Onthophagus parallelicornis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,parallelicornis,,,,,"Macleay, 1887","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137301,Onthophagus azusae,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,azusae,,,,,"Ochi & Kon, 2006","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide, with about 500 species in the Oriental region (ScarabNet 2009).
" 137302,Dichotomius glaucus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,glaucus,,,,,"(Harold, 1869)","The genus consists of approximately 160 valid species that are distributed throughout the Americas. However, the greatest diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia. Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus may change.
" 137303,Phanaeus bispinus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Phanaeus,bispinus,,,,,"Bates, 1868","The genus Phanaeus was revised by Edmonds (1994). Its species are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in Mexico. The species that compose the genus are divided into two subgenera (Notiophanaeus and Phanaeus s. str.).
" 137309,Onthophagus muticus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,muticus,,,,,"Macleay, 1864","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137314,Onthophagus diversiformis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,diversiformis,,,,,"Boucomont, 1914","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species (ScarabNet 2009). The Afro-Oriental subgenus Parascatonomus, to which this species has been assigned (Ochi 2003), includes around 50 species (ScarabNet 2009). The majority of them feed on carrion." 137315,Onthophagus eschscholtzi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,eschscholtzi,,,,,"Boucomont, 1924","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide (ScarabNet 2009). Presumably, this species is close to the representatives of the subgenus Gibbonthophagus." 137318,Gymnopleurus andreaei,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gymnopleurus,andreaei,,,,,"Ferreira, 1954","The genus Gymnopleurus Illiger, 1803 shows a range in the Afrotropical, Oriental and Palaearctic regions. It is currently represented by 33 species in the Afrotropical region (ScarabNet 2009)." 137321,Pachysoma aesculapius,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pachysoma,aesculapius,,,,,"(Olivier, 1789)","The genus Pachysoma Macleay 1821 comprises thirteen flightless species with a combined range extending along the dry to arid coastal region in the southwest of Africa (ScarabNet 2009)." 137322,Onthophagus granulatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,granulatus,,,,,"Boheman, 1858","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137323,Copris amabilis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,amabilis,,,,,"Kolbe, 1914","
The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009). Citations for Copris amabilis from west (Côte d’Ivoire) and west central Africa (Gabon) (Cambefort 1991, Cambefort and Walter 1991) refer to the recently-described close relative, Copris phungae Cambefort, 1992.
" 137324,Allogymnopleurus chloris,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Allogymnopleurus,chloris,,,,,"Klug, 1855","The Oriental and Afrotropical genus Allogymnopleurus Janssens 1940 is currently represented by 18 Afrotropical species (ScarabNet 2009). However, Allogymnopleurus chloris is considered to represent an often more shiny, green colour variety of Allogymnopleurus thalassinus Klug 1855.
" 137326,Canthidium atomarium,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,atomarium,,,,,"Balthasar, 1939","Canthidium is a diverse, species rich genus, with many species unknown. They are distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems. In addition to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137327,Pedaria morettoi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,morettoi,,,,,"Josso & Prevost, 2003", 137330,Onthophagus kavirondus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,kavirondus,,,,,"D'Orbigny, 1915","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 16 (d’Orbigny 1913), which is widespread in forest, savanna and grassland of the Afrotropical region. Othophagus kavirondus is cited as the valid name for this species (Ferreira 1972, Schoolmeesters 2011) but its synonym, Onthophagus kavirondi d’Orbigny 1913, was described two years earlier.
" 137333,Canthidium latum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,latum,,,,,"(Blanchard, 1846)","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.The genus still presents a considerable number of species to describe.
" 137336,Dichotomius crinicollis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,crinicollis,,,,,"(Germar, 1824)","The genus consists of approximately 160 valid species that are distributed throughout the Americas. However, the greatest diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia. Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus may change.
" 137339,Uroxys variabilis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Uroxys,variabilis,,,,,"Robinson, 1951","Uroxys presents about 60 valid species, distributed from Mexico to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.
" 137340,Oxysternon macleayi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Oxysternon,macleayi,,,,,"(Nevison, 1892)","The genus Oxysternon was recently revised by Edmonds and Zídek (2004). It consists of approximately 11 species which are distributed throughout tropical South America. The species that are found within the genus are divided into two subgenera: Oxysternon s. str. and Mioxysternon. According to Edmonds and Zídek (2004), Mioxysternon species occur along the periphery (or outside) of the Amazon Basin, whist those of Oxysternon s. str. are found within." 137343,Gymnopleurus thelwalli,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gymnopleurus,thelwalli,,,,,"Waterhouse, 1890","The genus Gymnopleurus Illiger, 1803 shows a range in the Afrotropical, Oriental and Palaearctic regions. It is currently represented by 33 species in the Afrotropical region (ScarabNet 2009)." 137345,Onthophagus gulmarri,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,gulmarri,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137346,Cryptocanthon brevisetosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,brevisetosus,,,,,"Howden, 1973", 137351,Bdelyrus metaensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,metaensis,,,,,"Cook, 1998","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
" 137352,Canthidium flabellatum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,flabellatum,,,,,"Harold, 1883","Canthidium is a diverse genus, distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
Canthidium flabellatum requires taxonomic work, as it may be a complex of multiple undescribed taxa that may be a different genus altogether.
" 137353,Gymnopleurus aeruginosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gymnopleurus,aeruginosus,,,,,"Harold, 1867","The genus Gymnopleurus Illiger, 1803 shows a range in the Afrotropical, Oriental and Palaearctic regions. It is currently represented by 33 species in the Afrotropical region (ScarabNet 2009). There is need to verify the relationship between G. aeruginosus and a widespread west and north central African species, G. coerulescens (Olivier, 1789) (F. Génier pers. comm.). This latter species occupies a range primarily in wetter savanna regions, which may be reflected by its metallic blue colour compared to the metallic green colouration but similar appearance of G. aeruginosus from possibly drier climate in the Sudan, based on material that is held in the collection of the Natural History Museum, London. Change in exoskeleton colour of G. humanus Macleay, 1821 from metallic blue to green to cupreous red in southern Africa has been shown to be correlated to temperature (Davis et al. 2008).
" 137354,Onthophagus mundill,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,mundill,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Females can be separated from the closely related Onthophagus mamillatus Matthews, 1972, but the males are indistinguishable, but the two species are widely geographically separated." 137363,Dichotomius planicollis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,planicollis,,,,,"(Gillet, 1911)","The genus consists of approximately 160 valid species that are distributed throughout Americas. However, the greater diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: (Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia). Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus can be changed." 137365,Oniticellus pseudoplanatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Oniticellus,pseudoplanatus,,,,,"Balthasar, 1964","The genus Oniticellus Serville 1828 occurs in both the Afrotropical and the Oriental regions, with five species currently considered valid in the Afrotropical region (ScarabNet 2009). Oniticellus pseudoplanatus is very closely related to Oniticellus planatus Castelnau 1840." 137367,Scybalocanthon trimaculatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scybalocanthon,trimaculatus,,,,,"(Schmidt, 1922)","The genus Scybalocanthon Martínez (1948) is a small genus of dung beetle from Central and South America. It needs taxonomical revision and many of its species can not be securely identified at this moment. So far, Scybalocanthon has 18 described species." 137368,Anomiopus nigrocoeruleus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,nigrocoeruleus,,,,,"(Martinez, 1955)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137370,Copris maesi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,maesi,,,,,"Ratcliffe, 1998","This species has been described from a single disarticulated specimen: ""I describe here a highly unusual new species of Copris whose disarticulated remains were found beneath a log in Nicaragua by Jean-Michel Maes and A. de la Fuente. I had hoped that additional specimens would have been found in the intervening years since the specimen was found, but this has not happened. I have decided to describe the species now, without waiting for a complete body to be collected, because its remarkably distinctive features are diagnostic."" (Ratcliffe 1998)" 137371,Pedaria spinithorax,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,spinithorax,,,,,"Paulian & Cambefort & Mauchamp, 1982", 137375,Coptodactyla nitida,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coptodactyla,nitida,,,,,"Paulian, 1933","The genus Coptodactyla Burmeister, 1846 occurs in Australia and Papua New Guinea. The Australian fauna comprises 14 species (Reid 2000, Matthews 1974, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). This species was raised from synonymy with Coptodactyla subaenea by Reid (2000)." 137380,Canthidium subdopuncticolle,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,subdopuncticolle,,,,,"Howden & Young, 1981","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe.
" 137381,Lepanus pisoniae,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Lepanus,pisoniae,,,,,"Lea, 1923","The genus Lepanus Balthasar, 1966, as conceived by Matthews (1974) and subsequent workers is the most heterogeneous of the Australian Canthonini. It comprises 24 described and at least 60 undescribed species (Matthews 1974, T. Weir pers. comm, G.B. Monteith pers. comm. and Gunter pers. comm. 2012). Lepanus pisoniae as interperated by Matthews (1974) is now known to be a complex of species. This assessment refers to the current knowledge of the true Lepanus pisoniae" 137385,Coptodactyla storeyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coptodactyla,storeyi,,,,,"Reid, 2000","The genus Coptodactyla Burmeister, 1846 occurs in Australia and Papua New Guinea. The Australian fauna comprises 14 species (Reid 2000, Matthews 1974, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137390,Pedaria biseria,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,biseria,,,,,"Josso & Prevost, 2003", 137398,Catharsius pandion,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Catharsius,pandion,,,,,"Harold, 1877","The Oriental and Afrotropical genus Catharsius Hope 1837 is currently represented by 80 species in the Afrotropical region (ScarabNet 2009). There is undoubtedly a good deal of confusion in the taxonomy of Catharsius pandion and published distribution records need to be treated with caution. It appears that it is one of a complex of closely related species that are similar in appearance and difficult to separate since the diagnostic morphological features vary in prominence and shape according to body size. Although there is no doubt that the taxon treated here as Catharsius pandion is a valid species, the application of the name, Catharsius pandion, to this species needs to be validated by a revision of the group taxonomy." 137406,Canthon lituratus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,lituratus,,,,,"(Germar, 1813)","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137409,Demarziella imitatrix,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Demarziella,imitatrix,,,,,"Balthasar, 1967","The endemic Australian genus Demarziella Balthasar, 1961 comprises 14 described and two undescribed species (Matthews 1976, Matthews 1986, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137411,Onitis mendax,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,mendax,,,,,"Gillet, 1918","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 19, which is centred in dry to moist savannas throughout the Afrotropical region and includes a number of known elephant dung specialists occurring on either deep sands or finer-grained soils. As in all Onitis, there is variation in body size and the shape and prominence of diagnostic femoral armament in males leading to synonyms for this widespread species described from both Zimbabwe (Onitis cerrutii Müller 1949) and Mozambique (Onitis janssenii Gomes Alves 1954)." 137420,Onthophagus giraffa,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,giraffa,,,,,"Hausmann, 1807","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species belongs to Onthophagus group 17 (d’Orbigny 1913), which is dominated by species showing rain forest, highland grassland, or cooler climatic ranges and is widespread in the Afrotropical region.
" 137422,Pachysoma endroedyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pachysoma,endroedyi,,,,,"(Harrison & Scholtz & Chown, 2003)","The genus Pachysoma Macleay 1821 comprises thirteen flightless species with a combined range extending along the dry to arid coastal region in the southwest of Africa (ScarabNet 2009)." 137423,Canthon plagiatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,plagiatus,,,,,"Harold, 1880","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.
" 137424,Onthophagus humpatensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,humpatensis,,,,,"Lansberge, 1886","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 24 (d’Orbigny 1913), which is widespread in the forest and savannas of the Afrotropical region.
" 137427,Coprophanaeus gamezi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coprophanaeus,gamezi,,,,,"Arnaud, 2002","According to Edmonds and Zidek (2010), the Neotropical genus Coprophanaeus Olsoufieff (1924) is currently represented by 38 species, distributed among three subgenera (Megaphanaeus Olsoufieff, Metallophanaeus Olsoufieff, and Coprophanaeus s. str.) and eight species groups." 137428,Onthophagus australis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,australis,,,,,"Guerin-Meneville, 1830","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Matthews (1972) records hybridising between this species and Onthophagus nurubuan Matthews, 1972 in the northern part of its range." 137429,Onthophagus victoriensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,victoriensis,,,,,"Blackburn, 1903","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012).
" 137430,Gymnopleurus reichei,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gymnopleurus,reichei,,,,,"Waterhouse, 1890","The genus Gymnopleurus Illiger, 1803 shows a range in the Afrotropical, Oriental and Palaearctic regions. It is currently represented by 33 species in the Afrotropical region (ScarabNet 2009). Other than in colour, South African specimens do not appear to differ appreciably to G. reichei from Kenya." 137431,Ontherus dentatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,dentatus,,,,,"Luederwaldt, 1930","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastern Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus." 137433,Temnoplectron finnigani,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Temnoplectron,finnigani,,,,,"Reid & Storey, 2000","The genus Temnoplectron Westwood, 1841 occurs in Australia and Papua New Guinea. The Australian fauna comprises 16 species (Matthews 1974, Reid and Storey 2000, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137438,Pachysoma hippocrates,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pachysoma,hippocrates,,,,,"(Macleay, 1821)","The genus Pachysoma Macleay 1821 comprises thirteen flightless species with a combined range extending along the dry to arid coastal region in the southwest of Africa (ScarabNet 2009). The morphological characteristics of P. hippocrates populations show a distinct north / south cline (Harrison et al. 2003) that is also reflected by phylogenetic analysis using the mitochondrial cytochrome oxidase I (COI) gene (Sole et al. 2005). Although, in the past, northern populations have been separated at specific level under the name, P. hessei Ferreira, this name is currently synonymized with P. hippocrates, and all populations are considered to belong to a single variable coastal species.
" 137441,Onthophagus semimetallicus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,semimetallicus,,,,,"Lea, 1923","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012).
" 137442,Phanaeus tridens,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Phanaeus,tridens,,,,,"Castelnau, 1840", 137449,Copris morphaeus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,morphaeus,,,,,"Gillet, 1932","The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009)." 137451,Cryptocanthon andersoni,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,andersoni,,,,,"Cook, 2002","According to Cook (2002), Cryptocanthon is a genus of wide occurrence in the New World tropics. They inhabit the litter of wet forests, both lowland rain forests and higher elevation montane and cloud forests, and occur from Brazil to Mexico." 137459,Onitis nemoralis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,nemoralis,,,,,"Gillet, 1932","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 3, which is widespread throughout the savannas and forests of Africa." 137461,Onthophagus tamworthi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,tamworthi,,,,,"Blackburn, 1903","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012).
" 137462,Pedaria tibialis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,tibialis,,,,,"Josso & Prevost, 2006", 137463,Onthophagus depressus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,depressus,,,,,"Harold, 1871","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 32 (d'Orbigny 1913), which is widely distributed across the dry to moist savannas of the Afrotropical region.
" 137465,Canthon quinquemaculatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,quinquemaculatus,,,,,"Castelnau, 1840","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.
" 137468,Uroxys pygmaeus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Uroxys,pygmaeus,,,,,"Harold, 1883","Uroxys presents about 60 valid species, distributed from Mexico to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.
" 137469,Onthophagus sikkimensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,sikkimensis,,,,,"Gillet, 1925","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide (ScarabNet 2009)." 137471,Ateuchus guatemalensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,guatemalensis,,,,,"(Bates, 1887)","Ateuchus is a diverse genus, with over 90 species, distributed from the USA to Argentina, and the Antilles. There are several taxonomic and nomenclatural problems, and many species to describe." 137473,Dichotomius horridus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,horridus,,,,,"(Felsche, 1911)","The genus consists of approximately 160 valid species that are distributed throughout Americas. However, the greater diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: (Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia). Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus can be changed.
" 137476,Canthon smaragdulus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,smaragdulus,,,,,"(Fabricius, 1781)","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.
" 137477,Onthophagus aethiopicus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,aethiopicus,,,,,"D'Orbigny, 1902","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 31 (d'Orbigny 1913), which is widely distributed throughout the savanna and highland grassland of the Afrotropical region and is equivalent to the Palaearctic and Oriental subgenus Furconthophagus Zunino 1979." 137479,Bdelyrus howdeni,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,howdeni,,,,,"Cook, 1998","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
" 137480,Neosisyphus rubrus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Neosisyphus,rubrus,,,,,"(Paschalidis, 1974)","The genus Neosisyphus Müller 1942 shows a widespread distribution in the Oriental and Afrotropical regions and is currently represented by 22 species in the Afrotropical region (ScarabNet 2009). It is in urgent need of revision. This species has frequently been cited as N. ruber for reasons of gender in Latin usage although no formal abridgement of the name has been published. It is a close relative of N. macrorubrus Paschalidis 1974, which has also been assessed for the IUCN Red List.
" 137482,Onthophagus planifrons,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,planifrons,,,,,"Frey, 1962","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide (ScarabNet 2009). This species is known only from the type series of two females and the taxonomic status likely requires reconsideration because of the fact that species identification in Onthophagus is usually impossible without males." 137485,Dichotomius apicalis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,apicalis,,,,,"(Luederwaldt, 1931)","The genus consists of approximately 160 valid species that are distributed throughout the Americas. However, the greatest diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia. Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus may change." 137491,Canthochilum anacaona,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthochilum,anacaona,,,,,"Zayas & Matthews, 1966","According to Philips and Ivie (2008), Canthochilum Chapin is a Greater Antillean endemic genus of small to very small scarabaeines (2.3–6.0 mm) found in forested habitats across a wide variety of moisture and elevational gradients. This genus is composed by 23 species and it represents more than half of the known scarabaeines of the Greater Antilles (Philips and Ivie, 2008).
" 137496,Anomiopus pictus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,pictus,,,,,"(Harold, 1862)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137498,Canthidium flavipes,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,flavipes,,,,,"Harold, 1867","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137499,Copris capensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,capensis,,,,,"Waterhouse, 1891","The genus Copris Muller 1764 shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009)." 137501,Canthon edentulus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,edentulus,,,,,"Harold, 1868","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137504,Onthophagus nefarius,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,nefarius,,,,,"Balthasar, 1941","The mega-diverse genus Onthophagus is currently represented by approximately 2,000 species. The Oriental subgenus Indachorius includes approximately 30 species (ScarabNet 2009). The species of this subgenus are quite small onthophagines which are similar to each other in external morphology and can reliably identified only by male genitalia. The representatives of the subgenus feed on dung, some of the species roll dung balls. The majority of the species are strictly localized, displaying a high degree of endemism to a particular habitat, normally primary tropical forests." 137505,Coprophanaeus pertyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coprophanaeus,pertyi,,,,,"(d'Olsufieff, 1924)","According to Edmonds and Zidek (2010), the Neotropical genus Coprophanaeus Olsoufieff (1924) is currently represented by 38 species, distributed among three subgenera (Megaphanaeus Olsoufieff, Metallophanaeus Olsoufieff, and Coprophanaeus s. str.) and eight species groups. " 137507,Ateuchus murrayi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,murrayi,,,,,"(Harold, 1868)","Ateuchus is a diverse genus, with approximately 70 species, distributed from the United States to Argentina and Lesser Antilles. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy of identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137510,Dendropaemon tenuitarsis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dendropaemon,tenuitarsis,,,,,"Felsche, 1909","The genus is currently being reviewed by Patrick Arnaud and François Génier. Therefore, it is not safe now to report data on the number of species or internal structure of the genus. The species of Dendropaemon are restricted to South America, except Chile.
" 137515,Onthophagus propinquus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,propinquus,,,,,"Macleay, 1888","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972; T. Weir pers. comm. and G.B. Monteith pers. comm. 2012). Onthophagus helmsi is considered a junior synonym of O. propinquus (Matthews 1972)." 137519,Platyonitis bicuariensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Platyonitis,bicuariensis,,,,,"Ferreira, 1976","This species is one of only three described for the genus Platyonitis, one occurring in southwest Africa and the other two in northeast Africa." 137520,Dichotomius maya,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,maya,,,,,"Peraza & Deloya, 2006", 137526,Aulacopris matthewsi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Aulacopris,matthewsi,,,,,"Storey, 1986","The endemic Australian genus Aulacopris White, 1859 comprises three species. Aulacopris matthewsi is the only flightless species of the genus known." 137530,Onthophagus caesariatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,caesariatus,,,,,"Boucomont, 1921","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide, of which around 500 species occur in the Oriental region (ScarabNet 2009)." 137532,Canthon laminatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,laminatus,,,,,"Balthasar, 1939","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137533,Pedaria durandi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,durandi,,,,,"Paulian & Cambefort & Mauchamp, 1982", 137540,Scarabaeus costatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,costatus,,,,,"(Wiedemann, 1823)","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively (ScarabNet 2009). This species was formerly classified as one of two members of the genus Drepanopodus Wiedemann 1823, which was recently synonymised with the genus Scarabaeus (Forgie et al. 2005)." 137542,Ontherus azteca,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,azteca,,,,,"Harold, 1869","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastern Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus." 137546,Cryptocanthon campbellorum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cryptocanthon,campbellorum,,,,,"Howden, 1973","According to Cook (2002), Cryptocanthon is a genus of wide occurrence in the New World tropics. They inhabit the litter of wet forests, both lowland rain forests and higher elevation montane and cloud forests, and occur from Brazil to Mexico." 137547,Catharsius chinai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Catharsius,chinai,,,,,"Ferreira, 1960","The Oriental and Afrotropical genus Catharsius Hope 1837 is currently represented by 80 species in the Afrotropical region (ScarabNet 2009). It is probable that understanding of the genus, species, and their distributional ranges would benefit from a revision." 137555,Demarziella planitarsis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Demarziella,planitarsis,,,,,"Petrovitz, 1971","The endemic Australian genus Demarziella Balthasar, 1961 comprises 14 described and two undescribed species (Matthews 1976, Matthews 1986, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137559,Gyronotus pumilus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gyronotus,pumilus,,,,,"(Boheman, 1857)","The genus Gyronotus Lansberge, 1874 is a flightless genus, currently comprising six species that are restricted to forest patches on the east coast of the Afrotropical region from Kenya to South Africa (ScarabNet 2009). Although G. marginatus Péringuey, 1888 was synonymized with G. pumilus (Boheman, 1857) (Scholtz and Howden 1987), it apparently differs in showing consistently larger body size than G. pumilus and restriction to forests right on the Pondoland coastline as opposed to a distribution extending further inland. Here, it is assessed as a synonym.
" 137561,Metacatharsius pumilioniformis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Metacatharsius,pumilioniformis,,,,,"Ferreira, 1964","The genus Metacatharsius Paulian 1939 is predominantly restricted to the Afrotropical region where 62 species are currently considered valid (ScarabNet 2009). A single species, Metacatharsius inermis Castelnau, extends from the Afrotropical region across Arabia to the western edge of the Oriental Region in a Saharo-Sindian pattern of distribution." 137562,Scarabaeus canaliculatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,canaliculatus,,,,,"Fairmaire, 1888","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2006): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Scarabeolus, which is primarily restricted to Africa. Available reference material identified as Scarabaeus canaliculatus originates entirely from the southwest of the Western Cape and coastal regions of Little Namaqualand (South Africa), whereas material identified as its close relative, Scarabaeus fritschi Harold 1868, originates from Great Namaqualand (southern Namibia), inland regions of Little Namaqualand above the escarpment, and the Northern Cape (both in South Africa). Scarabaeus canaliculatus was described from Namaqua (Fairmaire 1888) whereas Scarabaeus fritschi was described from Africa Austral. Inter. The cited type locality of Scarabaeus canaliculatus (Namaqua) is insufficiently exact to determine if it represents the southern or the northern species. As Péringuey (1901) cites the type locality of Scarabaeus canaliculatus as Great Namaqualand, this would place it outside of the known range of the species treated here as Scarabaeus canaliculatus, but coincidental with the range of the species treated here as Scarabaeus fritschi. Clearly, a revision of the taxonomy of these species is required as the lack of coincidence between ranges and type localities may be indicative of confusion between Scarabaeus canaliculatus and its close relative, including the synonym Scarabaeus reichei Waterhouse 1890, described from an unspecified southern locality in the Cape of Good Hope." 137563,Zonocopris machadoi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Zonocopris,machadoi,,,,,"Vaz-de-Mello, 2007","Zonocopris is known for its relationships with giant South American land snails. The genus was revised by Vaz-de-Mello (2007). Currently, it consists of two species, Z. gibbicollis (Harold 1868), distributed in Bolivia, Argentina, Paraguay and Brazil and Z. machadoi Vaz-de-Mello, 2007, from Paraguay and Brazil.
" 137564,Copris wiesei,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,wiesei,,,,,"Kolbe, 1914","The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009)." 137565,Ontherus raptor,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,raptor,,,,,"Génier, 1996","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastern Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus." 137566,Trichillidium quadridens,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Trichillidium,quadridens,,,,,"(Arrow, 1932)","The genus Trichillidium was described and positioned in the subtribe Scatimina (Coleoptera: Scarabaeinae: Ateuchini) by Vaz-de-Mello (2008). It is comprised of four species that are distributed throughout southern Nicaragua to central Argentina (although they are absent in western Amazonia), central and northeastern Brazil. The species that compose the genus were originally described in Trichillum or Pedaridium." 137568,Onthophagus hoplocerus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,hoplocerus,,,,,"Lea, 1923","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137570,Garreta malleolus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Garreta,malleolus,,,,,"(Kolbe, 1895)","The genus Garreta Janssens, 1940 is shared between the Afrotropical and Oriental regions with eleven species currently recognized as valid in the Afrotropical region (ScarabNet 2009)." 137573,Ontherus laminifer,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,laminifer,,,,,"Balthasar, 1938","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastern Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus." 137574,Canthidium femoratum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,femoratum,,,,,"Boucomont, 1935","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe.
" 137578,Boletoscapter furcatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Boletoscapter,furcatus,,,,,"Matthews, 1974","The endemic Australian genus Boletoscapter Matthews, 1974 comprises two species (Matthews 1974, T. Weir pers. comm. and G.B. Monteith pers. comm.).
" 137581,Macroderes cornutus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Macroderes,cornutus,,,,,"Frolov & Scholtz, 2004",The genus Macroderes Westwood 1842 is restricted to the winter and bimodal rainfall regions on the west and southern coastal regions of South Africa where it is currently represented by fourteen valid flightless species and three with poor support (ScarabNet 2009). 137584,Canthon quadripunctatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,quadripunctatus,,,,,"Redtenbacher, 1867","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically relatively well known.
" 137585,Canthidium flavicorne,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,flavicorne,,,,,"(Blanchard, 1846)","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.The genus still presents a considerable number of species to describe.
" 137586,Bdelyrus apaporisae,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,apaporisae,,,,,"Cook, 1998","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137598,Ateuchus halffteri,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,halffteri,,,,,"Kohlmann, 1981","Ateuchus is a diverse genus, with over 90 species, distributed from the USA to Argentina, and the Antilles. There are several taxonomic and nomenclatural problems, and many species to describe. Ateuchus halffteri is endemic of Mexico (Kohlmann 1981, Kohlmann 2003). Ateuchus halffteri is the only Mexican species with an internal sac with two copulatory spines." 137604,Onthophagus depilatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,depilatus,,,,,"D'Orbigny, 1913","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 9 (d'Orbigny 1913), which is widely distributed throughout the forests, moist savannas and cooler climates of the Afrotropical region." 137607,Canthon lividus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,lividus,,,,,"Blanchard, 1846","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, and Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America.The genus is taxonomically well known.
" 137608,Onitis meyeri,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,meyeri,,,,,"Kolbe, 1891","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 3, which is widespread throughout the savannas and forests of Africa." 137609,Onthophagus bornemisszai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,bornemisszai,,,,,"Matthews, 1972","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137613,Bdelyrus cochabambae,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Bdelyrus,cochabambae,,,,,"Cook, 2000","The group was reviewed by Cook in 1998. According to Cook (1998), Bdelyrus is endemic in tropical America and is widespread throughout this region, ranging from Nicaragua to southern Brazil and occurring in humid forest at elevations between 10 and 1,800 m. Adults have been found in detritus accumulated in reservoirs of both terrestrial and arboreal bromeliads. Species of this group are recorded in dung-baited pitfall traps, carrion-baited pitfall traps, wet forest litter, and flowers of Heliconia (Cook 1998).
" 137615,Pedaria cambeforti,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,cambeforti,,,,,"Josso & Prevost, 2003", 137617,Anomiopus laetus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,laetus,,,,,"(Waterhouse, 1891)","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137618,Onthophagus mamillatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,mamillatus,,,,,"Lea, 1923","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137622,Onthophagus janthinus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,janthinus,,,,,"Harold, 1875","Onthophagus is a diverse and cosmopolitan genus, with approximately 2,000 species distributed worldwide. The great diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task." 137629,Onthophagus aequepubens,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,aequepubens,,,,,"D'Orbigny, 1905","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species belongs to Onthophagus Group 2 (d'Orbigny 1913), which is widely represented in dry to wet regions throughout the Afrotropical region.
" 137631,Onthophagus anchommatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,anchommatus,,,,,"Lea, 1923","The cosmopolitan genus Onthophagus Latreille, 1802 is the largest of the Australian dung beetle fauna and comprises 193 described and 41 undescribed species (Matthews 1972, Storey 1977, Storey and Weir 1990, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137634,Odontoloma pygidiale,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Odontoloma,pygidiale,,,,,"Peringuey, 1901","The genus Odontoloma Boheman 1857 is restricted to the Afrotropical region where most of the twenty species currently considered valid show southerly centres of distribution (ScarabNet 2009). Owing to the very small body size and great similarity in appearance between most species, the taxonomy of this genus is difficult and requires revision." 137635,Deltochilum batesi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Deltochilum,batesi,,,,,"Paulian, 1938","Deltochilum was described in 1822 by Eschscholtz. The main diagnostic characteristic is the presence of short carinae or tubercles at the apex of some elitral interstriae. The genus is composed by eight subgenera: Aganhyboma Kolbe, 1893; Calhyboma Kolbe, 1893; Deltochilum s. str.; Deltohyboma Lane, 1946; Euhyboma Kolbe, 1983; Hybomidium Shipp, 1897; Parahyboma Paulian, 1938; Rubrohyboma Paulian, 1938." 137639,Ontherus hadros,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,hadros,,,,,"Génier, 1996","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastem Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus.
" 137641,Macroderes endroedyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Macroderes,endroedyi,,,,,"Frolov & Scholtz, 2004",The genus Macroderes Westwood 1842 is restricted to the winter and bimodal rainfall regions on the west and southern coastal regions of South Africa where it is currently represented by fourteen valid flightless species and three with poor support (ScarabNet 2009). 137643,Onitis obscurus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onitis,obscurus,,,,,"Lansberge, 1886","The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species is classified in Onitis Group 17, which shows a widespread distribution in dry and moist savanna of the Afrotropical region. The group 17 species occupy distinct regional ranges but are difficult to separate leading to taxonomic confusion in the literature and, as a result, much erroneous distribution data." 137644,Canthon helleri,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,helleri,,,,,"Schmidt, 1922","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137645,Lepanus politus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Lepanus,politus,,,,,"Carter, 1936","The genus Lepanus Balthasar, 1966, as conceived by Matthews (1974) and subsequent workers is the most heterogeneous of the Australian Canthonini. It comprises 24 described and at least 60 undescribed species (Matthews 1974, T. Weir pers. comm, G.B. Monteith pers. comm. and Gunter pers. comm. 2012)." 137646,Proagoderus extensus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Proagoderus,extensus,,,,,"(Harold, 1878)","The genus Proagoderus van Lansberge 1883 shows a wide biogeographical distribution across the Afrotropical and Oriental regions and is currently represented by 107 species in the Afrotropical region. These are divided into nine species groups (Palestrini 1992).
" 137648,Frankenbergerius nitidus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Frankenbergerius,nitidus,,,,,"Frolov & Scholtz, 2005","The genus Frankenbergerius Balthasar, 1938 has been recently revalidated (Frolov and Scholtz 2005) after being in synonymy with the genus Coptorhina Hope 1830. It currently comprises seven species restricted to the eastern and southwest margins of South Africa (ScarabNet 2009)." 137652,Scarabaeus rixosus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,rixosus,,,,,"Peringuey, 1901","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2006): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Scarabaeus. Scarabaeus rixosus is one of two species formerly classified under the generic name Neateuchus Gillet 1911, which was recently synonymized with Scarabaeus (Forgie et al. 2006). A recently re-discovered museum specimen may be the type of this species as it matches the original description in bearing a monofid spur on the mentum and bears a label in the handwriting of the describing authority (L. Péringuey) showing it was collected near the type locality. Péringuey’s personal collection is known to have been purchased and donated to the museum in which this specimen is lodged (National Collection of Insects, South Africa). Examination of the specimen shows that it differs distinctly from its close relative, Scarabaeus proboscideus Guérin-Méneville 1844, the only other Scarabaeus species known to bear a spur on the mentum." 137654,Canthidium magnum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,magnum,,,,,"Harold, 1871","Canthidium is a diverse genus, distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137656,Canthidium flavum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,flavum,,,,,"Balthasar, 1939","Canthidium is a diverse genus, distributed from the United States to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems (Martínez and Halffter 1986). In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.The genus still presents a considerable number of species to describe." 137659,Anomiopus birai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Anomiopus,birai,,,,,"Canhedo, 2006","The genus is exclusively Neotropical and most species occur in South America (Canhedo 2006). The group was reviewed by Canhedo in 2006. However, ten new species were described by the same author in 2004 parallel to the process of publication of taxonomic revision, so that the works that complement the taxonomic revision were published even before the own revision of the genus (Canhedo 2004a, 2004b, 2006). After the publication of these papers the genus now includes 58 species. Although there is a taxonomic revision with a relatively recent identification key for the genus, it still presents a considerable number of species to describe." 137660,Ateuchus ambiguus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,ambiguus,,,,,"Martinez & Martinez, 1990","Ateuchus is a diverse genus, with over 90 species, distributed from the United States to Argentina and Lesser Antilles. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy of identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe.The genus Onitis Fabricius 1798 is widespread in the Afrotropical, Oriental and Palaearctic regions. It is divided into 20 species groups of which 13 groups and 130 species are represented in Africa (ScarabNet 2009). This species belongs to Onitis Group 9, which shows a wide distribution across the moist savannas of the Afrotropical region (Ferreira 1978).
" 137666,Canthon heyrovskyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,heyrovskyi,,,,,"Balthasar, 1939","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon,Peltecantho, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
The genus is endemic to South America and currently has five valid, described species. They are specialists of beaches (riverine and sea).
" 137680,Odontoloma endroedyi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Odontoloma,endroedyi,,,,,"Howden & Scholtz, 1987","The genus Odontoloma Boheman 1857 is restricted to the Afrotropical region where most of the twenty species currently considered valid show southerly centres of distribution (ScarabNet 2009). Owing to the very small body size and great similarity in appearance between most species, the taxonomy of this genus is difficult and requires revision." 137681,Onthophagus lemekensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,lemekensis,,,,,"D'Orbigny, 1915","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species remains unassigned to a group." 137685,Caccobius cavatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Caccobius,cavatus,,,,,"D'Orbigny, 1908","The genus Caccobius Thomson 1859 is widespread in the Afrotropical, Oriental and Palaearctic regions. In the Afrotropical region, it is currently represented by 42 species that are considered valid (ScarabNet 2009)." 137687,Onthophagus falculatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,falculatus,,,,,"Boucomont, 1914","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide (ScarabNet 2009)." 137688,Onthophagus pseudovirens,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,pseudovirens,,,,,"D'Orbigny, 1913","A new subgenus of Onthophagus Latreille 1802 was recently created for O. pseudovirens and allied species, which were transferred from Proagoderus van Lansberge 1883, a taxon that is variously cited as having subgeneric or generic status. Since its creation, the subgenus, Hyalonthophagus Palestrini & Giacone 1989, has also often been cited as having generic status. Whether cited as a genus or subgenus, it is restricted to the Afrotropical region where it currently comprises ten very closely related species. Owing to the extremely close relationships, a review of the taxonomy within this genus is recommended." 137693,Coprophanaeus ohausi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Coprophanaeus,ohausi,,,,,"(Felsche, 1911)","According to Edmonds and Zidek (2010), the Neotropical genus Coprophanaeus Olsoufieff (1924) is currently represented by 38 species, distributed among three subgenera (Megaphanaeus Olsoufieff, Metallophanaeus Olsoufieff, and Coprophanaeus s. str.) and eight species groups. Edmonds and Zidek (2010) synonymized Coprophanaeus florenti Arnaud under Coprophanaeus ohausi Felsche.
" 137695,Onthophagus hippopotamus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,hippopotamus,,,,,"Harold, 1869", 137699,Onthophagus impressicollis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,impressicollis,,,,,"Boheman, 1860","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 24 (d’Orbigny 1913), which is widespread in the forest and savannas of the Afrotropical region. This species is entirely metallic green. Another sympatric Group 24 species, Onthophagus leucopygus Harold 1867, in which the elytra are white rather than metallic, is thought to be a further synonym of Onthophagus impressicollis Boheman 1860." 137702,Onthophagus quadricuspis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,quadricuspis,,,,,"D'Orbigny, 1908","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas of Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australiasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups of which 29 currently remain classified within the genus. Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 12, which is widespread in the dry to moist savannas of the Afrotropical region, and is equivalent to the Palaearctic and Oriental subgenus Trichonthophagus." 137704,Scarabaeus karrooensis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scarabaeus,karrooensis,,,,,"Zur Strassen, 1961","The genus Scarabaeus Linnaeus 1758 ranges across the Palaearctic, Oriental and Afrotropical regions and is currently considered to comprise four subgenera (Forgie et al. 2006): Scarabaeus Linnaeus 1758, Scarabeolus Balthasar 1965, Kheper Janssens 1940, Sceliages Westwood 1837, with a representation in the Afrotropical region of 55, 27, 21 and seven species respectively. This species belongs to the subgenus Scarabeolus." 137705,Dichotomius alyattes,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,alyattes,,,,,"(Harold, 1880)","The genus consists of approximately 160 valid species that are distributed throughout the Americas. However, the greater diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia. Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus may change.
" 137710,Onthophagus nigriventris,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,nigriventris,,,,,"D'Orbigny, 1905","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species belongs to Onthophagus Group 4 (d’Orbigny 1913), which shows a distribution restricted to tropical east African highlands and cool temperate highlands of South Africa.
" 137711,Onthophagus carpophilus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,carpophilus,,,,,"Pereira & Halffter, 1961", 137715,Lepanus gelasinus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Lepanus,gelasinus,,,,,"Matthews, 1974","The genus Lepanus Balthasar, 1966, as conceived by Matthews (1974) and subsequent workers is the most heterogeneous of the Australian Canthonini. It comprises 24 described and at least 60 undescribed species (Matthews 1974, T. Weir pers. comm, G.B. Monteith pers. comm. and Gunter pers. comm. 2012). Lepanus gelasinus as defined by Matthews (1974) is a complex of species distributed in central coastal Queensland. Lepanus gelasinus itself appears to be confined to an area around Bulburin State Forest and this assessment is confined to this." 137718,Onthophagus reyesi,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,reyesi,,,,,"Zunino & Halffter, 1988", 137720,Deltochilum carinatum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Deltochilum,carinatum,,,,,"(Westwood, 1837)","Deltochilum was described in 1822 by Eschscholtz. The main diagnostic characteristic is the presence of short carinae or tubercles at the apex of some elitral interstriae. The genus is composed by eight subgenera: Aganhyboma Kolbe, 1893; Calhyboma Kolbe, 1893; Deltochilum s. str.; Deltohyboma Lane, 1946; Euhyboma Kolbe, 1983; Hybomidium Shipp, 1897; Parahyboma Paulian, 1938; Rubrohyboma Paulian, 1938." 137725,Copris ritsemae,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Copris,ritsemae,,,,,"Harold, 1875","The genus Copris shows widespread biogeographical occurrence across the Afrotropical, Palaearctic, Oriental, Nearctic and northern part of the Neotropical regions, with a current total of 105 species known from the Afrotropical region (ScarabNet 2009). This species belongs to a group that is restricted to the southeast coastline and adjoining highlands of South Africa and is currently represented by six valid species between which there has been much taxonomic confusion (see also assessment of Copris victorini Boheman, 1857). Copris ritsemae has been described as a taxon that is distinct from its close relative, Copris caelatus (Fabricius, 1794), and it appears that most published locality data for Copris ritsemae is due to confusion with that species." 137729,Pedaria puncticollis,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Pedaria,puncticollis,,,,,"Waterhouse, 1890", 137734,Gymnopleurus bicolor,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Gymnopleurus,bicolor,,,,,"Latreille, 1823","The genus Gymnopleurus Illiger, 1803 shows a range in the Afrotropical, Oriental and Palaearctic regions. It is currently represented by 33 species in the Afrotropical region (ScarabNet 2009)." 137742,Cephalodesmius armiger,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Cephalodesmius,armiger,,,,,"Westwood, 1842","The flightless endemic Australian genus Cephalodesmius Westwood, 1841 comprises three described and one undescribed species (Matthews 1974, Monteith and Storey 1981, T. Weir pers. comm. and G.B. Monteith pers. comm. 2012)." 137751,Canthon columbianus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthon,columbianus,,,,,"Schmidt, 1920","Canthon is a diverse and heterogeneous genus of dung beetles. Currently, the genus is composed of nine subgenera: Boreocanthon, Canthon s.str., Francmonrosia, Glaphyrocanthon, Goniocanthon, Nesocanthon, Peltecanthon, Pseudepilissus, Trichocanthon. Of these, only Boreocanthon and Nesocanthon do not occur in South America. The genus is taxonomically well known.
" 137756,Onthophagus caprai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,caprai,,,,,"Frey, 1956","The mega-diverse subcosmopolitan genus Onthophagus is currently represented by approximately 2,000 species worldwide, of which around 500 species occur in the Oriental region (ScarabNet 2009). The Oriental subgenus Colobonthophagus includes about 30 species (ScarabNet 2009)." 137760,Canthidium aurichalceum,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Canthidium,aurichalceum,,,,,"Preudhomme de Borre, 1886","Canthidium is a diverse, species rich genus, with many species unknown. They are distributed from the United States to Argentina. The group also has various taxonomic and nomenclatural problems. In addition to this, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task. There are still a considerable number of species to describe in this genus.
" 137761,Scatonomus lauropalui,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Scatonomus,lauropalui,,,,,"Vulcano & Pereira, 1973","Scatonomus is a small genus, containing only eight described species, of which little taxonomic and biological information is available. The genus is known from Brazil, Argentina, Uruguay and Paraguay, and its species are distributed throughout “Cerrado”, “Pampa” and Atlantic forest." 137764,Epirinus relictus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Epirinus,relictus,,,,,"Scholtz & Howden, 1987",The genus Epirinus currently comprises 29 species with a combined distribution restricted primarily to the coolest climates of South Africa (ScarabNet 2009). 137765,Dichotomius nutans,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Dichotomius,nutans,,,,,"(Harold, 1867)","The genus consists of approximately 160 valid species that are distributed throughout Americas. However, the greater diversity of Dichotomius occurs in Brazil. The species that compose the genus are divided into four subgenera: (Dichotomius, Homocanthonides, Selenocopris and Luederwaldtinia). Currently, Dichotomius is being revised by F. Vaz-de-Mello. Thus, the number of valid species and the internal structure of the genus can be changed." 137768,Uroxys elongatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Uroxys,elongatus,,,,,"Harold, 1868","Uroxys presents about 60 valid species, distributed from Mexico to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task." 137770,Ontherus aequatorius,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ontherus,aequatorius,,,,,"Bates, 1891","The genus Ontherus was revised by Génier (1996). Its consists approximately of 60 species that are distributed throughout continental tropical and subtropical Latin America. However, the greater diversity of the group occurs in southeastem Brazil and the Andes (Génier 1996). The species that compose the genus are divided into three subgenera: Caelontherus, Planontherus and Ontherus.
" 137771,Uroxys coarctatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Uroxys,coarctatus,,,,,"Harold, 1867","
Uroxys presents about 60 valid species, distributed from Mexico to Argentina. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy in identification of most South American species an extremely difficult task.
" 137772,Ateuchus carbonarius,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Ateuchus,carbonarius,,,,,"(Harold, 1868)","Ateuchus is a diverse genus, with approximately 70 species, distributed from the United States to Argentina and Lesser Antilles. The diversity of the group is also accompanied by various taxonomic and nomenclatural problems. In addition, the need for taxonomic revisions makes the accuracy of identification of most South American species an extremely difficult task. The genus still presents a considerable number of species to describe." 137774,Amphistomus speculifer,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Amphistomus,speculifer,,,,,"Matthews, 1974","The genus Amphistomus Lansberge, 1874 is found in Australia, New Guinea and Maluku (Moluccas) Archipelago. In Australia it comprises 18 described species and 8 undescribed species. Amphistomus speculifer, as originally described, is now known to consist of two allopatric species. Amphistomus speculifer is confined to New South Wales only." 137775,Onthophagus lugubris,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Onthophagus,lugubris,,,,,"Fahraeus, 1857","Onthophagus Latreille 1802 is the most species-rich genus in the subfamily Scarabaeinae, with a distribution extending from tropical to warm temperate areas in the Palaearctic, Afrotropical, Oriental, Nearctic, Neotropical and Australasian regions. There are some 791 species in the Afrotropical region that have been divided into 32 species groups, of which 29 currently remain classified within the genus (ScarabNet 2009). Some of these groups extend into the Palaearctic and Oriental regions, but they have not yet been formally integrated into the subgeneric classification system erected for these regions. This species is classified in Onthophagus Group 16 (d’Orbigny 1913), which is widespread in forest, savanna and grassland of the Afrotropical region." 137778,Temnoplectron bornemisszai,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Temnoplectron,bornemisszai,,,,,"Matthews, 1974","The genus Temnoplectron Westwood, 1841 occurs in Australia and New Guinea. The Australian fauna comprises 16 species." 137779,Clypeodrepanus striatus,ANIMALIA,ARTHROPODA,INSECTA,COLEOPTERA,SCARABAEIDAE,Clypeodrepanus,striatus,,,,,"(Boucomont, 1921)","
This species has been recently transferred from