Table 4. List of all alycaeid species-group names. Subgenera are treated at the same level as genera in the "previous classification". Previous classification does not include our papers (Páll-Gergely 2017, Páll-Gergely & Asami 2017, Páll-Gergely et al. 2017, Páll-Gergely & Auffenberg 2019). Abbreviations for species examined: DOD: detailed original description was sufficient for generic placement; NE: not examined; NT: non-type material; T: types.
Taxon Previous classification This study Remarks Rank (species or subspecies) Specimens examined
abdoui Dicharax Dicharax sp T
abei Awalycaeus Dicharax sp NT
aborensis Chamalycaeus Metalycaeus sp T
akhaensis Raptomphalus Dicharax sp T
akioi Cipangocharax Dicharax sp T
akiratadai Awalycaeus Dicharax sp T
akyabensis Alycaeus Dicharax ssp T
alticola Foon & Liew, 2017 Alycaeus Stomacosmethis ssp DOD
alticola Páll-Gergely & Hunyadi, 2017 Dicharax Dicharax sp T
altispirus Alycaeus Stomacosmethis sp T
amphora Dioryx Dioryx sp T
ananensis Cipangocharax Dicharax sp DOD
anapetes nomen nudum
anceyi Alycaeus Pincerna sp T
andamaniae Chamalycaeus Chamalycaeus sp T
anghamiensis Dioryx Dioryx ssp T
anonymus Alycaeus Dicharax sp T
anthostoma Dicharax Dicharax sp T
armillatus Dicharax Chamalycaeus sp T
asaluensis Dicharax Dicharax sp T
ataranensis Dicharax Dicharax sp T
avae Dicharax Dicharax sp T
awaensis Chamalycaeus Metalycaeus sp T
awalycaeoides Metalycaeus Metalycaeus sp T
bacca Dioryx Dioryx sp T
balingensis Alycaeus Stomacosmethis sp T
barowliensis Alycaeus Dicharax sp T
bawai Dicharax Dicharax sp T
beddomei Alycaeus Metalycaeus sp T
bembex Cycloryx Pincerna sp T
bhutanensis Chamalycaeus Metalycaeus sp T
bicrenatus Dicharax Dicharax sp T
biexcisus Cipangocharax Dicharax sp T
bifrons Dicharax Dicharax sp T
birugosus Dicharax Dicharax sp T
bison Dicharax Dicharax sp T
blanfordi Alycaeus Dicharax sp T
brahma Chamalycaeus Metalycaeus sp T
broti nomen nudum
burrailensis Cycloryx Pincerna sp T
burroiensis Cycloryx Dicharax sp T
burtii Alycaeus Metalycaeus sp T
busbyi Chamalycaeus Chamalycaeus sp T
calopoma Alycaeus Stomacosmethis sp DOD: detailed original description was sufficient for generic placement; NE
canaliculatus Chamalycaeus Chamalycaeus sp T
canaliculus Chamalycaeus Dicharax junior synonym of birugosus sp T
candrakirana Dicharax Dicharax sp DOD
carinatus Alycaeus Stomacosmethis sp NT
cariniger Dioryx Dioryx sp T
caroli Chamalycaeus Metalycaeus sp T
caudapiscis Dicharax Dicharax sp T
celebensis Chamalycaeus Chamalycaeus sp T
chanjukensis Alycaeus Metalycaeus junior synonym of brahma sp T
chaperi Alycaeus Alycaeus junior synonym of gibbosulus sp DOD: detailed original description was sufficient for generic placement; NE
charasensis Alycaeus Stomacosmethis ssp DOD
chennelli Dicharax Dicharax sp T
christae Alycaeus Stomacosmethis sp T
clementsi Alycaeus Stomacosmethis ssp DOD
cochinensis Dioryx Dioryx sp T
commutatus Raptomphalus Metalycaeus junior synonym of brahma sp T
compactus Dioryx Dioryx sp T
compressicosta Dicharax Metalycaeus junior synonym of heudei syn T
conformis Alycaeus Alycaeus sp T
congener Alycaeus Stomacosmethis sp T
conicus Alycaeus Dicharax sp T
constrictus Cycloryx Pincerna sp T
costacrassa Alycaeus Stomacosmethis ssp DOD
costata Cycloryx Pincerna sp T
costulosa Alycaeus Pincerna sp T
crassicollis Chamalycaeus Dicharax sp DOD: detailed original description was sufficient for generic placement; NE
crassus Dicharax Dicharax ssp T
crenatus Dicharax Dicharax sp T
crenilabris Alycaeus Pincerna sp T
crenulatus Dicharax Metalycaeus sp T
crispatus Dicharax Dicharax sp T
cristatus Dicharax Dicharax sp T
cucullatus Dicharax Dicharax sp T
cyclophoroides Chamalycaeus Dicharax sp T
cyphogyrus Chamalycaeus Metalycaeus sp T
daflaensis Godwin-Austen, 1876 Dicharax Dicharax sp T
daflaensis Godwin-Austen, 1914 Dioryx Dioryx replaced by Dioryx urnula niosiensis nom. nov. ssp T
dalingensis Dicharax Dicharax sp T
damsangensis Dicharax Dicharax sp T
dautzenbergi Dioryx Dioryx replacement name for major Bavay & Dautzenberg, 1900 sp T
davisi Chamalycaeus Dicharax sp T
degenerans Alycaeus Stomacosmethis ssp T
depressus Alycaeus Dicharax sp T
diagonius Dicharax Dicharax sp T
difficilis Cycloryx Pincerna sp T
digitatus Dicharax Dicharax sp T
dihingensis Cycloryx Pincerna ssp T
dikrangensis Alycaeus Metalycaeus sp T
diminutus Chamalycaeus Dicharax sp T
diplochilus Chamalycaeus Dicharax sp T
distinctus Chamalycaeus Metalycaeus sp T
distortus Dioryx Dioryx sp T
ditaceus Sigmacharax Dicharax ssp T
diyungensis Dicharax Dicharax ssp T
dohertyi Dicharax Dicharax sp DOD: detailed original description was sufficient for generic placement; NE
dohrni Alycaeus Stomacosmethis sp T
dolichodeiros Chamalycaeus Dicharax sp DOD: detailed original description was sufficient for generic placement; NE
dolomiticus Alycaeus Metalycaeus junior synonym of rathouisianus syn T
dongiensis Dioryx Dioryx sp T
draco Dicharax Dicharax sp T
duoculmen Raptomphalus Dicharax sp T
duorugosus Dicharax Dicharax sp T
duplicatus Chamalycaeus Dicharax sp T
edei Chamalycaeus Dicharax sp T
elegans Cycloryx Pincerna sp T
elevatus Alycaeus Dicharax sp T
ellipticus Dicharax Dicharax sp T
everetti Dicharax Chamalycaeus sp T
excisus Chamalycaeus Chamalycaeus sp T
expanstoma Chamalycaeus Dicharax sp T
expansus Foon & Liew, 2017 Alycaeus Stomacosmethis ssp DOD
expansus Heude, 1890 Alycaeus Metalycaeus junior synonym of muciferus syn T
expatriatus Chamalycaeus Dicharax sp T
eydouxi Alycaeus Alycaeus replacement name for C. gibbum Draparnaud, 1805 sp NT
fargesianus Chamalycaeus Dicharax sp T
feddenianus Dioryx Dioryx sp NT
fimbriatus Dicharax Dicharax sp T
footei Dicharax Dicharax sp T
fractus Dicharax Metalycaeus junior synonym of heudei syn T
fraterculus Dicharax Dicharax sp T
fruhstorferi Chamalycaeus Chamalycaeus sp T
fultoni Alycaeus Stomacosmethis sp T
galbanus Alycaeus Stomacosmethis sp T
gemma Dicharax Dicharax sp T
gemmula Dicharax Dicharax sp T
generosus Cycloryx Dicharax sp T
gibbosulus Alycaeus Alycaeus sp T
gibbus Alycaeus Alycaeus replaced by eydouxi sp NT
glaber Dicharax Dicharax sp T
globosus Godwin-Austen, 1914 Dioryx Dioryx replaced by rotundus nom. nov. ssp T
globosus H. Adams, 1870 Alycaeus Pincerna sp T
globuloides Dioryx Dioryx sp T
globulosus Dioryx Dioryx sp T
globulus Dicharax Dicharax sp T
godwinausteni nom. nov. Alycaeus Metalycaeus nom. nov. pro A. neglectus Godwin-Austen, 1914 sp T
granum Cycloryx Pincerna sp T
graphiaria Cycloryx Pincerna sp T
graphica Cycloryx Pincerna sp T
habiangensis Dicharax Dicharax sp T
harimensis Chamalycaeus Dicharax junior synonym of japonicus syn T
hebes Dicharax Dicharax sp NT
helicodes Alycaeus Metalycaeus junior synonym of muciferus syn T
heudei Dicharax Metalycaeus sp T
hirasei Chamalycaeus Metalycaeus sp T
hochstetteri Alycaeus Stomacosmethis junior synonym of jagori syn DOD: detailed original description was sufficient for generic placement; NE
hosei Alycaeus Stomacosmethis sp NT
huberi Alycaeus Stomacosmethis junior synonym of somnueki syn T
humilis Dicharax Dicharax sp NT
hungerfordianus Chamalycaeus Metalycaeus sp T
ibex Metalycaeus Metalycaeus sp T
ikanensis Alycaeus Stomacosmethis ssp DOD
imitator Dicharax Dicharax sp T
immaculatus Dicharax Dicharax sp T
inflatus Godwin-Austen, 1874 Chamalycaeus Metalycaeus sp T
inflatus Möllendorff, 1886 Chamalycaeus Dicharax replaced by moellendorffi sp T
ingrami Chamalycaeus Dicharax sp T
itonis Sigmacharax Dicharax sp NT
jagori Alycaeus Stomacosmethis sp T
japonicus Chamalycaeus Dicharax sp NT
jatingaensis nom. nov. Alycaeus Dicharax nom. nov. pro A. nanus Godwin-Austen, 1914 ssp T
jaintiacus Dicharax Dicharax sp T
jousseaumei Chamalycaeus Alycaeus sp T
juttingae nom. nov. Alycaeus Pincerna nom. nov. pro laevis van Benthem Jutting, 1959 ssp DOD: detailed original description was sufficient for generic placement; NE
kamakiaensis Alycaeus Metalycaeus sp T
kapayanensis Alycaeus Stomacosmethis sp T
kelantanensis Alycaeus Stomacosmethis sp T
kengtungensis Raptomphalus Metalycaeus sp T
kessneri Chamalycaeus Chamalycaeus sp T
kezamaensis Dicharax Dicharax sp T
khasiacus Dicharax Dicharax sp T
khunhoensis Cycloryx Pincerna sp T
kinabaluana Alycaeus Pincerna ssp T
kiuchii Cipangocharax Dicharax sp T
kobeltianus Dioryx Dioryx sp T
korintjiensis nom. nov. Alycaeus Pincerna nom. nov. pro latecostata van Benthem Jutting, 1959 ssp DOD: detailed original description was sufficient for generic placement; NE
koshuensis Chamalycaeus Dicharax ssp NT
kuekenthali Chamalycaeus Stomacosmethis sp T
kurauensis Alycaeus Stomacosmethis ssp DOD
kurodai Metalycaeus Dicharax junior synonym of spiracellum syn T
kurodatokubeii Chamalycaeus Dicharax sp T
kurzianus Dicharax Dicharax sp T
labrirubidum Dioryx Dioryx sp T
laevicervix Chamalycaeus Metalycaeus ssp T
laevis Pilsbry & Y. Hirase, 1909 Chamalycaeus Metalycaeus sp T
laevis van Benthem Jutting, 1959 Alycaeus Pincerna replaced by juttingae nom. nov. ssp DOD: detailed original description was sufficient for generic placement; NE
lahupaensis Raptomphalus Dicharax sp T
laosensis Metalycaeus Metalycaeus sp T
Pincerna latecostata van Benthem Jutting, 1959 Alycaeus Pincerna replaced by korintjiensis nom. nov. ssp DOD: detailed original description was sufficient for generic placement; NE
latecostatus Möllendorff, 1882 Chamalycaeus Metalycaeus sp T
latestriata nomen nudum
lectus Dicharax Dicharax sp T
lenticulus Dicharax Dicharax sp T
levis Alycaeus Dicharax sp T
libonensis Chamalycaeus Metalycaeus sp T
liratula Pincerna Pincerna sp T
logtakensis Alycaeus Dicharax sp T
lohitensis Alycaeus Metalycaeus sp T
longituba Alycaeus Dicharax sp NT
luyorensis Raptomphalus Metalycaeus sp T
macgregori Chamalycaeus Metalycaeus sp T
magnificus Raptomphalus Metalycaeus sp T
magnus Alycaeus Dicharax sp T
major Dioryx Dioryx replaced by dautzenbergi sp T
major (granum var.) Cycloryx Pincerna senior synonym of A. mangutensis Godwin-Austen, 1914 sp T
makarsae Dicharax Dicharax ssp T
mangutensis Cycloryx Pincerna junior synonym of A. granum var. major Godwin-Austen, 1893 sp T
maolanensis Dioryx Pincerna sp T
maosmaiensis Alycaeus Dicharax sp T
margarita Cycloryx Pincerna sp T
matchacheepiorum Alycaeus Stomacosmethis sp DOD
mediocris Alycaeus Dicharax ssp T
melanopoma Metalycaeus Metalycaeus junior synonym of nipponensis syn T
menglunensis Dioryx Dioryx sp T
messageri Dioryx Dioryx sp T
microconus Chamalycaeus Chamalycaeus sp T
microcostatus Dicharax Dicharax sp T
microdiscus Chamalycaeus Dicharax sp T
micropolitus Dicharax Dicharax sp T
microstoma Alycaeus Alycaeus junior synonyom of sculptilis syn DOD: detailed original description was sufficient for generic placement; NE
minatoi Metalycaeus Metalycaeus sp T
minimus Dicharax Dicharax ssp T
minor (birugosus var.) Dicharax Dicharax junior synonym of birugosus syn T
minor (constrictus var.) Cycloryx Pincerna junior synonym of constrictus syn DOD: detailed original description was sufficient for generic placement; NE
minor (graphicus var.) Cycloryx Pincerna junior synonym of graphicus syn DOD: detailed original description was sufficient for generic placement; NE
minor (jagori var.) Alycaeus Stomacosmethis junior synonym of jagori syn DOD: detailed original description was sufficient for generic placement; NE
minor (paviei var.) Alycaeus Metalycaeus junior synonym of heudei syn T
minor (pilula var.) Dioryx Dioryx junior synonym of pilula syn DOD: detailed original description was sufficient for generic placement; NE
minor (vestitus var.) Alycaeus Dicharax junior synonym of vestitus syn DOD: detailed original description was sufficient for generic placement; NE
mixtus Chamalycaeus Chamalycaeus sp T
miyazakii Chamalycaeus Dicharax sp NT
moellendorffi Chamalycaeus Dicharax replacement name for inflatus Möllendorff, 1886 sp T
monadicus Dioryx Dioryx sp T
montanus Chamalycaeus Dicharax sp T
mouhoti Alycaeus Pincerna sp T
muciferus Chamalycaeus Metalycaeus sp T
multicostulata Cycloryx Pincerna sp T
multidentatus junior synonym of fimbriatus syn T
multirugosus Dicharax Dicharax sp T
muluana Alycaeus Pincerna ssp T
mundulus Alycaeus Metalycaeus sp T
muroharai Chamalycaeus Dicharax ssp T
muspratti Raptomphalus Dicharax sp T
mutatus Dicharax Dicharax sp T
nagaensis Chamalycaeus Dicharax sp T
nakashimai Sigmacharax Dicharax sp T
nanus Godwin-Austen, 1914 Alycaeus Dicharax replaced by jatingaensis nom. nov. ssp T
nanus Möllendorff, 1886 Chamalycaeus Dicharax junior synonym of diminutus syn T
nattoungensis Alycaeus Dicharax ssp T
neglectus Godwin-Austen, 1914 Alycaeus Metalycaeus replaced by godwinausteni nom. nov. sp T
neglectus Heude, 1885 Chamalycaeus Metalycaeus junior synonym of rathouisianus syn T
nicobaricus Alycaeus Alycaeus junior synonym of reinhardti syn DOD: detailed original description was sufficient for generic placement; NE
niosiensis nom. nov. Dioryx Dioryx nom. nov. pro Alycaeus daflaensis Godwin-Austen, 1914 ssp T
nipponensis Chamalycaeus Metalycaeus sp T
nishii Chamalycaeus Dicharax sp T
nitidus Chamalycaeus Dicharax sp T
nongtungensis Dicharax Dicharax sp T
notatus Dicharax Dicharax sp T
notus Dicharax Dicharax sp T
nowgongensis Alycaeus Dicharax sp T
oakesi Raptomphalus Metalycaeus sp T
obscurus Dicharax Dicharax sp T
ochraceus Dicharax Dicharax sp T
oglei Alycaeus Chamalycaeus sp T
oharai Metalycaeus Metalycaeus sp T
okamurai Cipangocharax Dicharax sp NT
okinawaensis Chamalycaeus Dicharax sp NT
okuboi Metalycaeus Metalycaeus sp T
oligopleuris Chamalycaeus Dicharax sp T
omissus Chamalycaeus Dicharax sp T
oshimanus Chamalycaeus Dicharax sp T
otiphorus Cycloryx Pincerna sp NT
pachitaensis Dicharax Dicharax sp T
panggianus Alycaeus Metalycaeus sp T
panshiensis Chamalycaeus Dicharax sp DOD: detailed original description was sufficient for generic placement; NE
parvulus Chamalycaeus Dicharax sp T
paucicostata Cycloryx Pincerna sp T
paviei Alycaeus Metalycaeus junior synonym of heudei syn T
peilei Dicharax Dicharax sp T
pentagonus Alycaeus Dicharax junior synonym of anthostoma syn T
perakensis Alycaeus Stomacosmethis sp T
perplexus Alycaeus Chamalycaeus sp T
physis Alycaeus Metalycaeus sp NT
pilsbryi Chamalycaeus Chamalycaeus synonym of japonicus sp T
pilula Dioryx Dioryx sp NT
pingoungensis Dioryx Dioryx sp T
pisum Dioryx Dioryx ssp T
placenovitas Cipangocharax Dicharax sp T
planorbulus Chamalycaeus Dicharax sp T
plectocheilus Dicharax Dicharax sp T
plicilabris Chamalycaeus Dicharax sp T
pocsi Dioryx Dioryx sp T
politus Alycaeus Dicharax sp T
polygonoma Dicharax Metalycaeus sp T
porcilliferus Alycaeus Stomacosmethis sp T
praetextus Alycaeus Stomacosmethis sp T
pratatensis Alycaeus Dicharax sp T
principalis Chamalycaeus Dicharax ssp T
prosectus Dicharax Metalycaeus sp T
purus Chamalycaeus Dicharax sp T
pusillus Alycaeus Dicharax sp T
pygmaea Alycaeus Pincerna ssp T
pyramidalis Alycaeus Alycaeus sp T
quadrasi Chamalycaeus Metalycaeus sp T
rabongensis Alycaeus Pincerna ssp T
rarus Chamalycaeus Chamalycaeus sp T
rathouisianus Chamalycaeus Metalycaeus sp T
rechilaensis Dicharax Dicharax sp T
regalis Alycaeus Stomacosmethis sp DOD
reinhardti Mörch, 1872 Alycaeus Chamalycaeus sp T
reinhardti Pilsbry, 1900 Alycaeus Dicharax replaced by pilsbryi Kobelt 1902 sp T
requiescens Alycaeus Dioryx sp T
reticulatus Alycaeus Chamalycaeus sp T
richthofeni Dicharax Chamalycaeus sp T
rimatus Alycaeus Stomacosmethis sp T
robustus Dicharax Dicharax sp T
roebeleni Alycaeus Stomacosmethis sp T
rolfbrandti Alycaeus Alycaeus sp T
rosea Dioryx Dioryx sp T
rotundatus Alycaeus Metalycaeus sp T
rotundus Dioryx Dioryx nom. nov. pro globosus Godwin-Austen, 1914 ssp T
rubinus Alycaeus Metalycaeus sp T
rugosus Dicharax Metalycaeus sp T
ruyangensis Dioryx Dioryx sp DOD
rywukensis Dicharax Dicharax ssp T
sabangensis Alycaeus Chamalycaeus ssp T
sadoensis Chamalycaeus Dicharax ssp T
sadongensis Alycaeus Stomacosmethis sp T
sandowayensis Chamalycaeus Dicharax sp T
sarasinorum Stomacosmethis Stomacosmethis sp T
satsumanus Chamalycaeus Metalycaeus sp T
scepticus nomen nudum
sculptilis Chamalycaeus Chamalycaeus sp T
sculpturus Alycaeus Dicharax sp T
selangoriensis Alycaeus Stomacosmethis ssp DOD
semperi Metalycaeus Metalycaeus sp T
senyumensis Alycaeus Stomacosmethis sp DOD
serratus Alycaeus Dicharax sp T
setchuanensis Dioryx Dioryx sp T
shiibaensis Chamalycaeus Dicharax sp T
shiosakimasahiroi Awalycaeus Dicharax sp DOD
shiotai Sigmacharax Dicharax ssp T
sibbumensis Alycaeus Metalycaeus sp T
simplicilabris Dicharax Dicharax junior synonym of cristatus syn T
sinensis Chamalycaeus Metalycaeus sp T
smithi Chamalycaeus Dicharax junior synonym of cristatus syn T
solidus Dicharax Dicharax ssp T
somnueki Alycaeus Stomacosmethis sp DOD
somwangi Alycaeus Alycaeus sp DOD
sonlaensis Alycaeus Dicharax sp DOD
specus Alycaeus Chamalycaeus sp T
spiracellum Dicharax Dicharax sp T
spratti Cycloryx Stomacosmethis sp T
stoliczkii Chamalycaeus Dicharax sp T
strangulatus Dicharax Dicharax sp NT
strigatus Chamalycaeus Dicharax sp DOD: detailed original description was sufficient for generic placement; NE
stuparum Dicharax Dicharax sp T
stylifer Dicharax Metalycaeus sp T
subculmen Dicharax Dicharax sp T
subdigitata Dicharax Dicharax ssp T
subfossilis Chamalycaeus Chamalycaeus sp T
subhumilis Dicharax Dicharax sp T
subinflatus Chamalycaeus Metalycaeus sp T
sublimus Chamalycaeus Chamalycaeus ssp T
succineus Dicharax Dicharax sp T
suhajdai nom. nov. Dioryx Metalycaeus nom. nov. pro Alycaeus varius Godwin-Austen, 1914 sp T
sumatranus Alycaeus Chamalycaeus sp T
summus Cycloryx Pincerna sp T
swinhoei Dioryx Dioryx sp T
sylheticus Alycaeus Dicharax sp T
tadai Chamalycaeus Dicharax sp NT
takahashii Chamalycaeus Dicharax sp T
tanegashimae Chamalycaeus Dicharax sp T
tanghali Alycaeus Chamalycaeus sp T
tangmaiensis Chamalycaeus Dicharax sp T
tangmaiensis Dioryx Dioryx sp T
tenellus Cycloryx Pincerna sp T
teriaensis Dicharax Metalycaeus sp T
theobaldi Dicharax Dicharax sp T
thieroti Alycaeus Pincerna sp T
thompsoni Cycloryx Pincerna sp T
tokunoshimanus Chamalycaeus Dicharax sp T
tomotrema Chamalycaeus Metalycaeus sp T
toruputuensis Dicharax Metalycaeus sp T
trigonostoma nomen nudum
troglodytes Chamalycaeus Chamalycaeus sp T
tsushimanus Chamalycaeus Dicharax sp T
umbonalis Chamalycaeus Dicharax sp T
urceolus Dioryx Dioryx sp T
urnula Dioryx Dioryx sp T
vallis Pincerna Pincerna sp DOD
vanbuensis Dioryx Pincerna sp T
variabilis Cycloryx Pincerna ssp T
varius Godwin-Austen, 1914 Dioryx Metalycaeus replaced by suhajdai nom. nov. sp T
varius Pilsbry & Y. Hirase 1905 Chamalycaeus Metalycaeus sp T
vesica Alycaeus Metalycaeus sp T
vestitus Alycaeus Dicharax sp T
vinctus Chamalycaeus Metalycaeus sp T
virgogravida Alycaeus Stomacosmethis ssp DOD
vulcani Alycaeus Chamalycaeus sp T
wilhelminae Alycaeus Stomacosmethis sp DOD
woodthorpi Dicharax Dicharax sp T
yamneyensis Alycaeus Metalycaeus sp T
yanoshigehumii Chamalycaeus Dicharax sp T
yanoshokoae Awalycaeus Dicharax sp DOD
yanseni Pincerna Pincerna sp T
yetayensis Alycaeus Metalycaeus ssp T
zayuensis Chamalycaeus Metalycaeus sp see remarks
zhuangiyucuii Alycaeus Metalycaeus junior synonym of heudei syn T