This document describes the structure of the AusTraits compilation, corresponding to Version 6.0.0 of the dataset. Note that the information provided below is based on the information provided within the file system.file("data", "schema.yml", package = "traits.build").

For details on access, structure and usage please visit https://doi.org/10.5281/zenodo.3568417

Elements of AusTraits

The compiled AusTraits database has the following main components:

austraits
├── traits
├── locations
├── contexts
├── methods
├── excluded_data
├── taxonomic_updates
├── taxa
├── contributors
├── sources
├── definitions
├── schema
├── metadata
└── build_info

These elements include all the data and contextual information submitted with each contributed datasets. Each component is defined as follows:

traits

Description: A table containing measurements of traits.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
taxon_name Scientific name of the taxon on which traits were sampled, without authorship. When possible, this is the currently accepted (botanical) or valid (zoological) scientific name, but might also be a higher taxonomic level.
observation_id A unique integral identifier for the observation, where an observation is all measurements made on an individual at a single point in time. It is important for joining traits coming from the same observation_id. Within each dataset, observation_id’s are unique combinations of taxon_name, population_id, individual_id, and temporal_context_id.
trait_name Name of the trait sampled. Allowable values specified in the table definitions.
value The measured value of a trait, location property or context property.
unit Units of the sampled trait value after aligning with AusTraits standards.
entity_type A categorical variable specifying the entity corresponding to the trait values recorded.
value_type A categorical variable describing the statistical nature of the trait value recorded.
basis_of_value A categorical variable describing how the trait value was obtained.
replicates Number of replicate measurements that comprise a recorded trait measurement. A numeric value (or range) is ideal and appropriate if the value type is a mean, median, min or max. For these value types, if replication is unknown the entry should be unknown. If the value type is raw_value the replicate value should be 1. If the trait is categorical or the value indicates a measurement for an entire species (or other taxon) replicate value should be .na.
basis_of_record A categorical variable specifying from which kind of specimen traits were recorded.
life_stage A field to indicate the life stage or age class of the entity measured. Standard values are adult, sapling, seedling and juvenile.
population_id A unique integer identifier for a population, where a population is defined as individuals growing in the same location (location_id /location_name) and plot (plot_context_id, a context category) and being subjected to the same treatment (treatment_context_id, a context category).
individual_id A unique integer identifier for an individual, with individuals numbered sequentially within each dataset by taxon by population grouping. Most often each row of data represents an individual, but in some datasets trait data collected on a single individual is presented across multiple rows of data, such as if the same trait is measured using different methods or the same individual is measured repeatedly across time.
repeat_measurements_id A unique integer identifier for repeat measurements of a trait that comprise a single observation, such as a response curve.
temporal_context_id A unique integer identifier assigned where repeat observations are made on the same individual (or population, or taxon) across time. The identifier links to specific information in the context table.
source_id For datasets that are compilations, an identifier for the original data source.
location_id A unique integer identifier for a location, with locations numbered sequentially within a dataset. The identifier links to specific information in the location table.
entity_context_id A unique integer identifier indicating specific contextual properties of an individual, possibly including the individual’s sex or caste (for social insects).
plot_context_id A unique integer identifier for a plot, where a plot is a distinct collection of organisms within a single geographic location, such as plants growing on different aspects or blocks in an experiment. The identifier links to specific information in the context table.
treatment_context_id A unique integer identifier for a treatment, where a treatment is any experimental manipulation to an organism’s growing/living conditions. The identifier links to specific information in the context table.
collection_date Date sample was taken, in the format yyyy-mm-dd, yyyy-mm or yyyy, depending on the resoluton specified. Alternatively an overall range for the study can be indicating, with the starting and ending sample date sepatated by a /, as in 2010-10/2011-03.
measurement_remarks Brief comments or notes accompanying the trait measurement.
method_id A unique integer identifier to distinguish between multiple sets of methods used to measure a single trait within the same dataset. The identifier links to specific information in the methods table.
method_context_id A unique integer identifier indicating a trait is measured multiple times on the same entity, with different methods used for each entry. This field is only used if a single trait is measured using multiple methods within the same dataset. The identifier links to specific information in the context table.
original_name Name given to taxon in the original data supplied by the authors.

locations

Description: A table containing observations of location/site characteristics associated with information in traits. Cross referencing between the two dataframes is possible using combinations of the variables dataset_id, location_name.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
location_id A unique integer identifier for a location, with locations numbered sequentially within a dataset. The identifier links to specific information in the location table.
location_name The location name.
location_property The location characteristic being recorded. The name should include units of measurement, e.g. MAT (C). Ideally we have at least the following variables for each location, longitude (deg), latitude (deg), description.
value The measured value of a location property.

contexts

Description: A table containing observations of contextual characteristics associated with information in traits. Cross referencing between the two dataframes is possible using combinations of the variables dataset_id, link_id, and link_vals.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
context_property The contextual characteristic being recorded. If applicable, name should include units of measurement, e.g. CO2 concentration (ppm).
category The category of context property, with options being plot, treatment, individual_context, temporal and method.
value The measured value of a context property.
description Description of a specific context property value.
link_id Variable indicating which identifier column in the traits table contains the specified link_vals.
link_vals Unique integer identifiers that link between identifier columns in the traits table and the contextual properties/values in the contexts table.

methods

Description: A table containing details on methods with which data were collected, including time frame and source. Cross referencing with the traits table is possible using combinations of the variables dataset_id, trait_name.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
trait_name Name of the trait sampled. Allowable values specified in the table definitions.
methods A textual description of the methods used to collect the trait data. Whenever available, methods are taken near-verbatim from the referenced source. Methods can include descriptions such as ‘measured on botanical collections’, ‘data from the literature’, or a detailed description of the field or lab methods used to collect the data.
method_id A unique integer identifier to distinguish between multiple sets of methods used to measure a single trait within the same dataset. The identifier links to specific information in the methods table.
description A 1-2 sentence description of the purpose of the study.
sampling_strategy A written description of how study locations were selected and how study individuals were selected. When available, this information is lifted verbatim from a published manuscript. For preserved specimens, this field ideally indicates which records were ‘sampled’ to measure a specific trait.
source_primary_key Citation key for the primary source in sources. The key is typically formatted as Surname_year.
source_primary_citation Citation for the primary source. This detail is generated from the primary source in the metadata.
source_secondary_key Citation key for the secondary source in sources. The key is typically formatted as Surname_year.
source_secondary_citation Citations for the secondary source. This detail is generated from the secondary source in the metadata.
source_original_dataset_key Citation key for the original dataset_id in sources; for compilations. The key is typically formatted as Surname_year.
source_original_dataset_citation Citations for the original dataset_id in sources; for compilationse. This detail is generated from the original source in the metadata.
data_collectors The person (people) leading data collection for this study.
assistants Names of additional people who played a more minor role in data collection for the study.
dataset_curators Names of AusTraits team member(s) who contacted the data collectors and added the study to the AusTraits repository.

excluded_data

Description: A table of data that did not pass quality tests and so were excluded from the master dataset. The structure is identical to that presented in the traits table, only with an extra column called error indicating why the record was excluded. Common reasons are missing_unit_conversions, missing_value, and unsupported_trait_value.

Content:

key value
error Indicating why the record was excluded. Common reasons are missing_unit_conversions, missing_value, and unsupported_trait_value.
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
taxon_name Scientific name of the taxon on which traits were sampled, without authorship. When possible, this is the currently accepted (botanical) or valid (zoological) scientific name, but might also be a higher taxonomic level.
observation_id A unique integral identifier for the observation, where an observation is all measurements made on an individual at a single point in time. It is important for joining traits coming from the same observation_id. Within each dataset, observation_id’s are unique combinations of taxon_name, population_id, individual_id, and temporal_context_id.
trait_name Name of the trait sampled. Allowable values specified in the table definitions.
value The measured value of a trait.
unit Units of the sampled trait value after aligning with AusTraits standards.
entity_type A categorical variable specifying the entity corresponding to the trait values recorded.
value_type A categorical variable describing the statistical nature of the trait value recorded.
basis_of_value A categorical variable describing how the trait value was obtained.
replicates Number of replicate measurements that comprise a recorded trait measurement. A numeric value (or range) is ideal and appropriate if the value type is a mean, median, min or max. For these value types, if replication is unknown the entry should be unknown. If the value type is raw_value the replicate value should be 1. If the trait is categorical or the value indicates a measurement for an entire species (or other taxon) replicate value should be .na.
basis_of_record A categorical variable specifying from which kind of specimen traits were recorded.
life_stage A field to indicate the life stage or age class of the entity measured. Standard values are adult, sapling, seedling and juvenile.
population_id A unique integer identifier for a population, where a population is defined as individuals growing in the same location (location_id /location_name) and plot (plot_context_id, a context category) and being subjected to the same treatment (treatment_context_id, a context category).
individual_id A unique integer identifier for an individual, with individuals numbered sequentially within each dataset by taxon by population grouping. Most often each row of data represents an individual, but in some datasets trait data collected on a single individual is presented across multiple rows of data, such as if the same trait is measured using different methods or the same individual is measured repeatedly across time.
repeat_measurements_id A unique integer identifier for repeat measurements of a trait that comprise a single observation, such as a response curve.
temporal_context_id A unique integer identifier assigned where repeat observations are made on the same individual (or population, or taxon) across time. The identifier links to specific information in the context table.
source_id For datasets that are compilations, an identifier for the original data source.
location_id A unique integer identifier for a location, with locations numbered sequentially within a dataset. The identifier links to specific information in the location table.
entity_context_id A unique integer identifier indicating specific contextual properties of an individual, possibly including the individual’s sex or caste (for social insects).
plot_context_id A unique integer identifier for a plot, where a plot is a distinct collection of organisms within a single geographic location, such as plants growing on different aspects or blocks in an experiment. The identifier links to specific information in the context table.
treatment_context_id A unique integer identifier for a treatment, where a treatment is any experimental manipulation to an organism’s growing/living conditions. The identifier links to specific information in the context table.
collection_date Date sample was taken, in the format yyyy-mm-dd, yyyy-mm or yyyy, depending on the resoluton specified. Alternatively an overall range for the study can be indicating, with the starting and ending sample date sepatated by a /, as in 2010-10/2011-03.
measurement_remarks Brief comments or notes accompanying the trait measurement.
method_id A unique integer identifier to distinguish between multiple sets of methods used to measure a single trait within the same dataset. The identifier links to specific information in the methods table.
method_context_id A unique integer identifier indicating a trait is measured multiple times on the same entity, with different methods used for each entry. This field is only used if a single trait is measured using multiple methods within the same dataset. The identifier links to specific information in the context table.
original_name Name given to taxon in the original data supplied by the authors.

taxonomic_updates

Description: A table of all taxonomic changes implemented in the construction of AusTraits. Changes are determined by comparing the originally submitted taxon name against the taxonomic names listed in the taxonomic reference files, best placed in a subfolder in the config folder . Cross referencing with the traits table is possible using combinations of the variables dataset_id and taxon_name.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
original_name Name given to taxon in the original data supplied by the authors.
aligned_name The taxon name without authorship after implementing automated syntax standardisation and spelling changes as well as manually encoded syntax alignments for this taxon in the metadata file for the corresponding dataset_id. This name has not yet been matched to the currently accepted (botanical) or valid (zoological) taxon name in cases where there are taxonomic synonyms, isonyms, orthographic variants, etc.
taxonomic_resolution The rank of the most specific taxon name (or scientific name) to which a submitted orignal name resolves.
taxon_name Scientific name of the taxon on which traits were sampled, without authorship. When possible, this is the currently accepted (botanical) or valid (zoological) scientific name, but might also be a higher taxonomic level.
aligned_name_taxon_id An identifier for the aligned name before it is updated to the currently accepted name usage. This may be a global unique identifier or an identifier specific to the data set. Must be resolvable within this dataset.
aligned_name_taxonomic_status The status of the use of the aligned_name as a label for a taxon. Requires taxonomic opinion to define the scope of a taxon. Rules of priority then are used to define the taxonomic status of the nomenclature contained in that scope, combined with the experts opinion. It must be linked to a specific taxonomic reference that defines the concept.

taxa

Description: A table containing details on taxa associated with information in traits. Whenever possible, this information is sourced from curated taxon lists that include identifiers for each taxon. The information compiled in this table is released under a CC-BY3 license. Cross referencing between the two dataframes is possible using combinations of the variable taxon_name.

Content:

key value
taxon_name Scientific name of the taxon on which traits were sampled, without authorship. When possible, this is the currently accepted (botanical) or valid (zoological) scientific name, but might also be a higher taxonomic level.
taxonomic_dataset Name of the taxonomy (tree) that contains this concept. ie. APC, AusMoss etc.
taxon_rank The taxonomic rank of the most specific name in the scientific name.
trinomial The infraspecific taxon name match for an original name. This column is assigned na for taxon name that are at a broader taxonomic_resolution.
binomial The species-level taxon name match for an original name. This column is assigned na for taxon name that are at a broader taxonomic_resolution.
genus Genus of the taxon without authorship.
family Family of the taxon.
taxon_distribution Known distribution of the taxon, by Australian state.
establishment_means Statement about whether an organism or organisms have been introduced to a given place and time through the direct or indirect activity of modern humans.
taxonomic_status The status of the use of the scientificName as a label for the taxon in regard to the ‘accepted (or valid) taxonomy’. The assigned taxonomic status must be linked to a specific taxonomic reference that defines the concept.
taxon_id An identifier for the set of taxon information (data associated with the taxon class). May be a global unique identifier or an identifier specific to the data set. Must be resolvable within this dataset.
taxon_id_genus An identifier for the set of taxon information (data associated with the taxon class) for the genus associated with a taxon name. May be a global unique identifier or an identifier specific to the data set. Must be resolvable within this dataset.
taxon_id_family An identifier for the set of taxon information (data associated with the taxon class) for the family associated with a taxon name. May be a global unique identifier or an identifier specific to the data set. Must be resolvable within this dataset.
scientific_name The full scientific name, with authorship and date information if known.
scientific_name_id An identifier for the set of taxon information (data associated with the taxon class). May be a global unique identifier or an identifier specific to the data set. Must be resolvable within this dataset.

contributors

Description: A table of people contributing to each study.

Content:

key value
dataset_id Primary identifier for each study contributed to AusTraits; most often these are scientific papers, books, or online resources. By default this should be the name of the first author and year of publication, e.g. Falster_2005.
last_name Last name of the data collector.
given_name Given names of the data collector.
ORCID ORCID of the data collector.
affiliation Last known institution or affiliation.
additional_role Additional roles of data collector, mostly contact person.

sources

Description: Bibtex entries for all primary and secondary sources in the compilation.

definitions

Description: A copy of the definitions for all tables and terms. Information included here was used to process data and generate any documentation for the study.

schema

Description: A copy of the schema for all tables and terms. Information included here was used to process data and generate any documentation for the study.

metadata

Description: Metadata associated with the dataset, including title, creators, license, subject, funding sources.

build_info

Description: A description of the computing environment used to create this version of the dataset, including version number, git commit and R session_info.

Methological details

Dataset IDs

The core organising unit behind AusTraits is the dataset_id. Records are organisation as coming from a particular study, defined by the dataset_id. Our preferred format for dataset_id is surname of the first author of any corresponding publication, followed by the year, as surname_year. E.g. Falster_2005. Wherever there are multiple studies with the same id, we add a suffix _2, _3 etc. E.g.Falster_2005, Falster_2005_2.

Other IDs

As well as a dataset_id, each trait measurement has 10 additional identifiers, observation_id, population_id, individual_id, temporal_id, source_id, location_id, entity_context_id, plot_id, treatment_id, and method_id.

All except source_id are simply integral identifiers that link groups of measurements and are automatically generated through the AusTraits workflow (individual_id can be assigned in the metadata file or automatically generated.)

To expand on the definitions provided above,

  • observation_id links measurements made on the same entity (individual, population, or species) at a single point in time.

  • population_id indicates entites that share a common location_id, plot_id, and treatment_id. It is used to align measurements and observation_id’s for individuals versus populations (i.e. distinct entity_types) that share a common population_id. It is numbered sequentially within a dataset.

  • individual_id indicates a unique organisms. It is numbered sequentially within a dataset by population. Multiple observations on the same organism across time (with distinct observation_id values), share a common individual_id.

  • temporal_id indicates a distinct point in time and is used only if there are repeat measurements on a population or individual across time. The identifier links to context properties (& their associated information) in the contexts table for context properties of type temporal.

  • source_id is applied if not all data within a single dataset (dataset_id) is from the same source, such as when a dataset represents a compilation for a meta-analysis.

  • location_id links to a distinct location_name and associated location_properties in the location table.

  • entity_context_id links to information in the contexts table for context properties (& associated values/descriptions) with category entity_context. Entity_contexts include organism sex, organism caste and any other features of an entity that needs to be documented.

  • plot_id links to information in the contexts table for context properties (& associated values/descriptions) with category plot. Plot contexts include both blocks/plots within an experimental design as well as any stratified variation within a location that needs to be documented (e.g. slope position).

  • treatment_idlinks to information in the contexts table for context properties (& associated values/descriptions) with category treatment. Treatment contexts are experimental manipulations applied to groups of individuals.

  • method_idlinks to information in the contexts table for context properties (& associated values/descriptions) with category method. A method context indicates that the same trait was measured on or across individuals using different methods.

Values, Value types, basis of value

Each record in the table of trait data has an associated value, value_type, and basis_of_value.

Value type: A trait’s value_type is either numeric or categorical. - For traits with numerical values, the recorded value has been converted into standardised units and the AusTraits workflow has confirmed the value can be converted into a number and lies within the allowable range. - For categorical variables, records have been aligned through substitutions to values listed as allowable values (terms) in a trait’s definition. * we use _ for multi-word terms, e.g. semi_deciduous
* we use a space for situations where two values co-occur for the same entity. For instance, a flora might indicate that a plant species can be either annual or biennial, in which case the trait is scored as annual biennial.

Each trait measurement has an associated value_type, which is a categorical variable describing the statistical nature of the trait value recorded. Possible values are:

key value
raw Value recorded for an entity.
minimum Value is the minimum of values recorded for an entity.
mean Value is the mean of values recorded for an entity.
median Value is the median of values recorded for an entity.
maximum Value is the maximum of values recorded for an entity.
mode Value is the mode of values recorded for an entity. This is the appropriate value type for a categorical trait value.
range Value is a range of values recorded for an entity.
bin Value for an entity falls within specified limits.
unknown Not currently known.

Each trait measurement also has an associated basis_of_value, which is a categorical variable describing how the trait value was obtained. Possible values are:

key value
measurement Value is the result of a measurement(s) made on a specimen(s).
expert_score Value has been estimated by an expert based on their knowledge of the entity.
model_derived Value is derived from a statistical model, for example via gap-filling.
unknown Not currently known.

AusTraits does not include intra-individual observations made at a single point in time. When multiple measurements per individual are submitted to AusTraits, we take the mean of the values and record the value_type as mean and indicate under replicates the number of measurements made.

Taxonomy

Version 6.0.0 of AusTraits contains records for 33494 different taxa. We have aligned taxa with known taxonomic units in the Australian Plant Census (APC) and/or the Australian Plant Names Index (APNI). Of the 33494 taxa included, 33319 are aligned with known taxa.

The traits table reports both the original and the updated taxon name alongside each trait record.

The table taxa lists all taxa in the database, including additional information about the taxa.

The table taxonomic_updates provides details on all taxonomic names changes implemented in aligning with APC and APNI.

Sources

For each dataset in the compilation there is the option to list primary and secondary citations as well as, for compilations, original citations. The primary citation The original study in which data were collected. while the secondary citation is A subsequent study where data were compiled or re-analysed.. These references are included in two places:

  1. Within the table methods, where we provide a formatted version of each.
  2. In the element sources, where we provided bibtex versions of all sources which can be imported into your reference library. The keys for these references are listed within the methods.

definitions

Following is a list of traits included in this version.

leaf_compoundness

  • label: Leaf compoundness
  • description: A leaf shape trait [TO:0000492] which is whether a leaf [PO:0025034] is a simple leaf [PO:0020042] or is divided into leaflets [PO:0020049] making it a compound leaf [PO:0020043].;A binary trait that indicates whether a leaf lamina is simple or divided into discontinuous leaflets (compound).
  • type: categorical
  • allowable values:
    • compound: A leaf that is divided into multiple leaflets. [PO:0020043]
    • simple: A leaf with a single undivided blade. [PO:0020042]

seed_length

  • label: Seed length
  • description: A seed morphology trait [TO:0000184] which is the greatest linear dimension [PATO:0001708] (axis) of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed width [PATO:0000921] and seed height [PATO:0000119] axes [NCIT:C25154].;Longest linear seed dimension.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1000 mm

leaf_phenology

  • label: Leaf phenology
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait characterising whether there is a part of the year when a plant sheds all leaves [PO:0025034] (shedability [PATO:0001729]) and the environmental conditions [ENVO:01000203] that cause a plant to shed its leaves.;Trait indicating whether a plant retains a leaf canopy throughout the year and what environmental cues are associated with canopy shedding.
  • type: categorical
  • allowable values:
    • brevi_deciduous: Plant briefly loses its entire leaf canopy but flushes a new leaf canopy almost simultaneously to leaf loss.
    • cold_deciduous: Plant loses its entire leaf canopy for part of the year, in response to cold.
    • deciduous: Plant loses its entire leaf canopy for part of the year, in response to either drought or cold.
    • drought_deciduous: Plant loses its entire leaf canopy for part of the year, in response to drought.
    • evergreen: Plant retains a leaf canopy throughout the year.
    • facultative_drought_deciduous: Plant sometimes loses its leaf canopy for part of the year, in response to drought.
    • semi_deciduous: Plant thins, but does not entirely lose, its leaf canopy in response to an environmental cue, usually drought.

leaf_length

  • label: Leaf length
  • description: Length [PATO:0000122] of the leaf [PO:0025034] (including the petiole [PO:0020038]), from the leaf apex [PO:0020137] to the leaf axil [PO:0009023] (leaf base [PO:0020040]).;Length of a leaf or a leaf analogue, from the leaf apex to the leaf axil (base), including the petiole.
  • type: numeric
  • units: mm
  • allowable range: 0.1 - 1e+05 mm

leaf_width

  • label: Leaf width
  • description: The maximum [NCIT:C25564] width [PATO:0000921] axis of a leaf [PO:0025034] measured perpendicular to [PATO:0002434] the leaf length [PATO:0000122].;Maximum width axis of a leaf or a leaf analogue (a leaflet, cladode, or phyllode), measured perpendicular to the leaf length.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1e+05 mm

plant_height

  • label: Plant vegetative height
  • description: A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the vegetative shoot system [PO:0025607] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341].;Maximum vertical height of the vegetative shoot system for a taxon in a given location.
  • type: numeric
  • units: m
  • allowable range: 0.001 - 130 m

flowering_time

  • label: Flowering time, by month
  • description: The months [UO:0000035] during which a plant [PO:0000003] has open (anthetic) flowers [PO:0009046] and is therefore at the whole plant flowering stage [PO:0007016].;Months during which taxon has open (anthetic) flowers.
  • type: categorical
  • allowable values:
    • **:

seed_shape

  • label: Seed shape
  • description: A seed morphology trait [TO:0000184] which is the 3-D [PATO:0001710] shape [PATO:0000052] of a seed [PO:0009010] considering the relative seed length [PATO:0000122], seed width [PATO:0000921], and seed height PATO:0000119] dimensions and the curvature [PATO:0001591] of the seed’s surface [NCIT:C25245].;Three-dimensional shape of the seed.
  • type: categorical
  • allowable values:
    • comma-shaped: A seed displaying a single arc of curvature, such that it bilaterally symmetrical. (Synonyms, comma-shaped, falcate, sickle-shaped scimitar-shaped)
    • conical: Cone-shaped, with a broad flat base and narrow apex; turbinate refers to the inverse shape. (Synonyms, obconical, cone-shaped, turbinate, top-shaped)
    • cuneate: A 3-dimensional shape that is wedge-shaped or boat-shaped.
    • cylindrical: A cylinder, a shape with a flat top and bottom that are circular or elliptical in outline and are linked by a curved surface. (Synonyms, terete)
    • discoid: A subset of cylinder, a shape with a flat, circular or elliptical top and bottom, linked by a curved surface, where the length of the curved surface is much shorter than the cross-section of the flat surfaces. (Synonyms, flat)
    • ellipsoid: A 3-dimensional shape that is elliptic in long section and circular in cross section; length:width ratio between 2:1 and 3:2. (Synonyms, obloid, oblong)
    • ellipsoid_flattened: A 3-dimensional shape that is elliptic in long section and and laterally compressed/flattened such that it is non-circular in cross section; length:width ratio between 2:1 and 3:2.
    • fusiform: Spindle-shaped; A 3-dimension shape that is narrowly ellipsoid, circular in cross-section and tapering sharply toward both ends. (Synonyms, spindle-shaped)
    • globoid: A 3-dimensional shape that is circular in both long section and cross section. (Synonyms, globose, globular, spherical)
    • globoid_flattened: A 3-dimensional shape that is circular in both long section and somewhat flattened such that it is non-circular in cross section.
    • hemispheric: Half a sphere or globe, with a flat base, domed top, and circular in cross-section. (Synonyms, convex)
    • hemispheric_flattened: A 3-dimensional shape that is a laterally compressed/flattened half-sphere or globe; shape has a flat base, a domed top, and is elliptical in cross-section.
    • lenticular: Shaped like a biconvex lens and circular in cross section; shaped like a lentil.
    • orbicular: A poorly defined term that remains in AusTraits only as a placeholder; it appears to be used as both globoid (spherical) and lenticular by different authors.
    • ovoid: A 3-dimensional shape that is egg-shaped in long section and circular in cross section. Term as used in AusTraits includes variants on egg-shaped including pyriform (pear-shaped) and obovoid (inversely egg-shaped) (Synonyms, pyriform, obovoid)
    • ovoid_elongated: A 3-dimensional shape that is egg-shaped in long section and circular in cross section, where the length dimension is much longer than the width/height dimensions. Term as used in AusTraits includes specific shapes club-shaped (clavate/obclavate) and tear-shaped (lachrimiform). (Synonyms, lachrimiform, tear-shaped, lachrymiform, clavate; obclavate, club-shaped)
    • ovoid_flattened: A 3-dimensional shape that is egg-shaped in long section and laterally compressed/flattened such that is non-circular in cross section.
    • polyhedral: A 3-dimensional shape with distinct, fairly flat faces and edges; includes prisms and tetrahedral shaped seeds. (Synonyms, rhomboid, prismatic, pyramidal, rectangular, square)
    • polyhedral_elongated: A 3-dimensional shape with distinct, fairly flat faces and edges, where the length dimension is much longer than the width/height dimensions; includes elongate prisms and elongate tetrahedral shaped seeds.
    • polyhedral_inflated: A 3-dimensional shape with distinct faces and edges but the faces tend to bulge outwards.
    • reniform: Having a cross-section that is roughly circular with a notch; strictly a kidney-shape but the definition applied here includes heart-shaped seeds and other seeds with a distinct notch. (Synonyms, cordiform, kidney-shaped, heart-shaped)
    • sectoroid: Shaped like an orange segment or wedge of a sphere. (Synonyms, wedge-shaped)
    • sinusoidal: A seed with two arcs of curvature or a single arc, such that it lacks an axis of symmetry. (Synonyms, sigmoidal)
    • winged: A seed with wings. (Synonyms, samara)

seed_width

  • label: Seed width
  • description: A seed morphology trait [TO:0000184] which is the longest [NCIT:C25558] width [PATO:0000921] dimension of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed length [PATO:0000122] and seed height [PATO:0000119] axes [NCIT:C25154].;Longest width dimension of a seed; orthogonal to the length and height axes.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1000 mm

seed_height

  • label: Seed height
  • description: A seed morphology trait [TO:0000184] which is the shortest diameter [NCT:C120715] axis [NCIT:C25154] of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed length [PATO:0000122] and seed width [PATO:0000921] axes.;Shorter width dimension of a seed; orthogonal to its length and width axes.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1000 mm

fruit_length

  • label: Fruit length
  • description: A fruit morphology trait [TO:0002629] which is the length [PATO:0000122] of a fresh [EnvThes:21976] fruit [PO:0009001] from the fruit proximal end [PO:0008002] (base) to the fruit distal end [PO:0008001] (apex).;Linear dimension from the base to the apex of a fresh fruit, even if this is not the longest dimension.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 2000 mm

fruit_width

  • label: Fruit width
  • description: A fruit morphology trait [TO:0002629] which is the width [PATO:0000921] of a fresh [EnvThes:21976] fruit [PO:0009001], where width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the fruit length [PATO:0000122] and fruit height [PATO:0000119].;Greatest linear width dimension of a fresh fruit, sometimes referred to as ‘diameter’ in fruits where the width and height are identical; orthogonal to the fruit length, the base-to-apex dimension.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 2000 mm

dispersal_appendage

  • label: Dispersal appendage
  • description: A seed morphology trait [TO:0002629] which indicates external [NCIT:C44280] appendages [NCIT:C61460] on the seed [PO:0009010] or the seed’s dispersal [OMIT: 0027297] structures [PATO:0000141] whose presumed function [NCIT:C25518] is to facilitate seed dispersal [EnvThes:21042].;External appendages of the dispersule presumed to facilitate seed dispersal away from the parent plant.
  • type: categorical
  • allowable values:
    • aril: Fleshy outgrowth of a seed, that often attracts animals like birds or ants.
    • bracts_and_glumes: When the bracts below the inflorescence are persistent and functional as dispersal appendages. This includes persistent glumes, the modified membranous bracts in the family Poaceae that surround the spikelet of a grass.
    • bristles: Bristle-like projections. Trait value includes awns, which are slender, bristle-like projections in some grasses.
    • elaiosome: Fleshy (often fatty) appendage on seeds that attracts ants.
    • fleshy_reward: Broad term that includes a suite of appendages that provides a fleshy reward to attract dispersers; highest level of resolution available for some studies.
    • floating_seed: Seed that is dispersed by floating on water.
    • floral_parts: When some plant floral parts (the petals, sepals, or style) are persistent and aid in seed and fruit dispersal. This is a broad term not explicitly suggesting a dispersal mechanism.
    • flotation_scales: Scales enhancing flotation, thereby assisting with dispersal.
    • hairs: Modified hairs that aid in seed dispersal.
    • hooks: A rear-facing point, as in a fish hook that aids in seed and fruit dispersal.
    • inflated_parts: When some part of the seed, fruit, or associated tissues is inflated, aiding in seed or fruit dispersal.
    • none: When a fruit and associated tissues lack any dispersal appendages. Includes exarillate and taxa with explicitly deciduous pappus.
    • pappus: The calyx in Asteraceae that has been modified into bristles, hairs, scales or awns that are attached to the apex of the fruit. Taxa assigned this trait value have a persistent pappus that aids in dispersal, often, but not exclusively through wind dispersal. The pappus can also assist with flotation.
    • plumose: Fruit, seed, style, or other floral component that is plumose and assists with dispersal.
    • receptacle: Fleshy receptacle that aids in dispersal.
    • sarcotesta: Fleshy seed coat that aids in dispersal.
    • spines: Spines that aid in dispersal.
    • tumbleweed: Plant growth form, whereby the aboveground plant component detaches from the roots and is readily rolled by wind across the ground, aiding in dispersal.
    • wing_or_plume: Broad term that includes a suite of appendages that aid in wind dispersal; highest level of resolution available for some studies.
    • wings: Referring to wing-like seed extensions that aid in wind dispersal.

plant_width

  • label: Plant canopy width
  • description: A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] width [PATO:0000600] dimension of a plant canopy [ENVO:01001242] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341]. This is a plant’s larger [NCIT:C49508] width dimension and is orthogonal to its height and breadth.;Maximum width dimension of a plant’s vegetative canopy for a taxon in a given location; this is the larger width dimension and is orthogonal to its height and breadth.
  • type: numeric
  • units: m
  • allowable range: 0.001 - 20 m

plant_growth_substrate

  • label: Plant growth substrate
  • description: A whole plant [PO:0000003] life history [ECOCORE:00000018] trait which indicates either the substrate [EFO:0005061] on which a plant grows [GO:0040007] or, broadly, whether the plant grows in an aquatic [EnvThes:21785] or terrestrial [EnvThes:21880] habitat [ENVO:01000739]. Substrate is defined as the surface to which an organism is attached.;Trait broadly indicating where a plant is usually found, referencing the habitat in which a plant grows, the substrate to which a plant is attached, or a plant’s growth medium.
  • type: categorical
  • allowable values:
    • aquatic: A plant that grows in water.
    • epiphyte: A plant that grows upon another plant.
    • hemiepiphyte: A plant that spends part of its life cycle as an epiphyte, growing upon another plant.
    • lithophyte: A plant that grows upon a rock.
    • marine: A plant that grows in the sea.
    • semiaquatic: A plant that grows partially submerged in water or spends part of its life growing in water.
    • terrestrial: A plant that grows on land, and generally referencing a plant that grows in soil.

fruit_height

  • label: Fruit breadth
  • description: A fruit morphology trait [TO:0002629] which is the height [PATO:0000119] of a fresh [EnvThes:21976] fruit [PO:0009001], where height is the shorter linear width dimension [PATO:0001708] perpendicular to [PATO:0002434] both the fruit length [PATO:0000122] and fruit width [PATO:0000921].;Shorter linear width dimension of a fresh fruit and generally not reported in fruits where the width and height are identical; orthogonal to the length, the base-to-apex dimension.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 2000 mm

petiole_length

  • label: Petiole length
  • description: Length [PATO:0000122] of the petiole [PO:0020038]), from the leaf base [PO:0020040] to the leaf axil [PO:0009023].;Length of the petiole from the leaf base to the leaf axil.
  • type: numeric
  • units: cm
  • allowable range: 0 - 1000 cm

life_history

  • label: Life history
  • description: Categorical description of the duration [PATO:0001309] of a plant’s lifespan (longevity [NCIT:C153298]), from seed germination [GO:0009845] to death [GO:0016265].;Categorical description of the duration of a plant’s lifespan, from germination to death.
  • type: categorical
  • allowable values:
    • annual: A plant that lives for up to one year after germinating, completing its life cycle in a single growing season.
    • biennial: A plant that lives for up to two years after germinating, requiring two growing seasons to complete its life cycle and dying following the second growing season.
    • ephemeral: A very short-lived plant, generally with a lifespan of only a few months and the exact length of lifespan quite variable and determined by environmental conditions.
    • perennial: A plant that lives for three or more growing seasons, with an exact lifespan that is indeterminate.
    • short_lived_perennial: A perennial whose lifespan is less than approximately five years, with the exact lifespan generally dependent on environmental conditions.

plant_growth_form

  • label: Plant growth form
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait that summarises different components of a whole plant’s shape [PATO:0000052] and structure [PATO:0000141] into general groups [NCIT:C43359]. Characters that contribute to the plant form categories include the height [PATO:0000119] of the vegetative shoot system [PO:0025607], the distribution of leaves [PO:0025034] along the plant stem [PO:0009047], and the position [PATO:0000140] and count [NCIT:C25463] of stems and/or leaves relative to the ground surface.;The form of a plant, capturing a general sense of plant height and shape, which parts of a plant are woody versus herbaceous, the number of stems arising at ground level, and the distribution of leaves relative to the main stem.
  • type: categorical
  • allowable values:
    • basal_large: A plant whose leaves form a distinctly large basal tuft or rosette, including large acaulescent monocots and cycads.
    • climber: Plant that climbs up another plant’s stem or branches, rather than being able to support itself. Term used only if datasets do not specify if a climbing plant is herbaceous (a vine) or woody (a liana).
    • climber_herbaceous: Herbaceous plant that climbs up another plant’s stem or branches, rather than being able to support itself. This is usually synonymous with vine. Types of climbers (scrambling, twining) are captured under the trait plant_climbing_mechanism. (Synonyms, vine)
    • climber_woody: Woody or thick-stemmed (>2 cm) plant that climbs up another plant’s stem or branches, rather than being able to support itself. This is usually synonymous with liana. Woody climbers generally use hooks, tendrils, and/or adventitious roots to climb; the climbing mechanisms used by a taxon are captured under the trait plant_climbing_mechanism. (Synonyms, liana)
    • fern: A taxonomic grouping that includes acaulescent ferns; ferns are vascular plants that reproduce via spores and have neither seeds nor flowers.
    • geophyte: Subcategory of herb, specifically referring to plants that survive the winter/dry season as an organ (rhizome, bulb, corm, or tuber) buried in dry soil, with all leaves dying back each year.
    • graminoid: Herbaceous plant with a grass-like morphology that is within the order Poales. Taxa both with and without a tussock form are mapped to this term. This term is most frequently applied to three large monocot families, Poaceae, Cyperaceae, and Juncaceae, but the term could apply to other taxa within Poales with a grass-like morphology, including narrow, linear leaves and non-showy flowers, including Restionaceae, Centrolepidaceae, Anarthriaceae, and Ecdeiocoleaceae. Taxa outside Poales with a grass-like morphology should be designated as herbs and also have the term tufted mapped to the trait stem_growth_habit.
    • graminoid_not_tussock: A subcategory of graminoid, indicating a plant which explicitly lacks a tussock morphology, with the leaves not growing in tufts.
    • herb: A seed-bearing plant which does not have a woody above-ground stem; when available this term has been divided into more detailed terms. (Synonyms, forb)
    • hummock: Plant that grows as a dense mound up to 1 metre in height. The term generally applies only to species within the genera Triodia that form such mounds and these are considered a distinctive and uniquely Australian growth form.
    • lycophyte: A taxonomic grouping that includes quillworts, clubmosses, and spike mosses.
    • mallee: Refers to Eucalyptus species which are small trees that form a large lignotuber. They have multiple stems arising at or near ground level and regenerate from the lignotuber following fire.
    • palmoid: Plant that bears a rosette-like canopy of typically large, often compound leaves atop a usually thick (pachycaulous), columnar, unbranched or little-branched stem. Palms, tree ferns, trunked cycads, trunked grass-trees, and Pandanus are mapped to this term. (Synonyms, rosette-tree, palm, grass-tree)
    • shrub: A woody plant that is distinguished by NOT being a tree, due to one of more of the following characteristics, 1) they are less than 8 metres in height; 2) if a plant community includes trees, the shrubs are shorter than the surrounding trees and not part of the canopy; 3) in comparison to trees, the leaves are more distributed along the entire trunk; 4) the plant has multiple relatively narrow stems arising at or near ground level. The definition of shrub is complex, as there are many single-stemmed shrubs within Australia and many taxa that are described in the taxonomic literature as a shrub or small tree.
    • subshrub: A plant that is only woody at the base, with the remainder of the stems regenerating yearly. This term is not meant to encompass short-lived plants that are only borderline woody. (Synonyms, suffrutescent)
    • tree: A tall, woody, perennial plant, usually with a single main trunk, and its leaves predominantly elevated above the ground surface.
    • tussock: A subcategory of graminoid, which is a plant where many shoots from the basal meristem form prominent tufts.

stem_growth_habit

  • label: Stem growth habit
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait that indicates the structure [PATO:0000141] and 3-dimensional extent [PATO:0001710] of a plant’s vegetative shoot system [PO:0025607], including the position [PATO:0000140] of stems [PO:0009047], degree of branchiness [PATO:0002009], density [PATO:0001019] of branches [PO:0025073], and branch angles [PATO:0002326] relative to each other and the ground surface [sweetontology.net/realmSoil/Ground].;Trait characterising a stem’s structure and 3-dimensional extent, including the position of stems, degree of branchiness, density of branches, and branch angles relative to other branches and the ground surface.
  • type: categorical
  • allowable values:
    • arborescent: Tree-like growth habit.
    • bushy: Plant with much-branched branches. (Synonyms, shrubby)
    • caespitose: Multiple short-shoots arise from the base of the plant forming a dense tuft, as in a bunch-grass. (Synonyms, cespitose, bunch-grass, tussock)
    • climbing: Stem that does not have a self-supporting structure and ascends by leaning on or attaching itself to surrounding vegetation or another substrate. Climbing mechanisms include tendrils, petioles, adventitious roots, scrambling, and twining and are distinguished between in the trait plant_climbing_mechanism.
    • compact: Plant with closely packed branches.
    • creeping: Stem that creeps along the ground, rooting at nodes. (Synonyms, repent)
    • cushion-forming: Multiple erect, short stems arise from the soil and are so tightly packed that they form dense, even, rounded canopy with no gaps. (Synonyms, cushion)
    • decumbent: Stems whose bases lie flat on the ground but whose tips grow upward. (Synonyms, ascending)
    • dense: Plant with a high density of branches.
    • erect: Stems that grow upright.
    • floating: Plant where most of the leaves are floating on water.
    • mat-forming: Multiple stems that are mostly horizontal and are densely packed but open enough that there are still a few gaps between them.
    • open: Plant with an open architecture of branches.
    • prostrate: Stems lie flat on the ground but do not root at nodes. (Synonyms, procumbent, reclining, trailing)
    • rhizomatous: Plants with short horizontal underground stems from which shoots emerge.
    • rosette: Plants whose leaves radiate from a central point at or near ground level. (Synonyms, acaulescent, rosulate)
    • spindly: Plants with weak, skinny branches.
    • sprawling: Stem that sprawls across other plants or objects but lacks specialised climbing structures or mechanisms. (Synonyms, clambering, scandent)
    • spreading: Plants with a notable horizontal extent, referring either to a broad tree canopy or a shrub or herb that spreads across the ground.
    • stoloniferous: Plants with long horizontal branches either slightly belowground or aboveground, that root at their nodes or tips, producing new plants. (Synonyms, sarmentose)
    • submerged: Plant where most of the leaves are submerged beneath the surface of a waterbody.
    • tufted: Plant has a dense clump of leaves, either at the base of the plant or clusters of leaves growing close together higher on the plant. This term can refer to grasses, shrubs, or trees.

woodiness_detailed

  • label: Detailed woodiness categories
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait which indicates the vertical [PATO:0001854] extent, distribution [PATO:0000060], and type [NCIT:C25284] of wood present in stems [PO:0009047], roots [PO:0009005], and reproductive shoot systems [PO:0025082], distinguishing between plants with secondary xylem (true wood) [PO:0005848] versus wood-like lignification [CHEBI:6457] (woodiness in monocots and ferns).;Trait indicating the extent, distribution, and type of secondary xylem (true wood) and wood-like lignification (woodiness in monocots and ferns) in stems, roots, and reproductive shoots.
  • type: categorical
  • allowable values:
    • herbaceous: Plant with non-lignified stems.
    • semi_woody: Plant that forms stems considered only “partially woody”, generally having lower lignin and cellulose contents and possibly not forming secondary xylem. These are generally short-lived, fast-growing stems. (Synonyms, green-stem shrub, soft shrub, soft wood)
    • woody: Plant whose stems produce secondary xylem (true wood).
    • woody_base: Plant where secondary xylem (true wood) is present only at the base of stems. (Synonyms, suffrutescent)
    • woody_like_inflorescence: Taxa, including monocots, that produce thick, stiff, robust lignified inflorescence shoots but are otherwise non-woody.
    • woody_like_stem: Taxa, including monocots and ferns, that produce thick, stiff, robust lignified stems but do not produce secondary xylem.
    • woody_root: Plant that has a woody rootstock but non-woody above-ground stems.

sex_type

  • label: Plant sex type
  • description: A plant structure morphology trait [TO:0000839] which is the distribution [PATO:0000060] of male [PATO:0000384], female [PATO:0000383], and hermaphroditic [PATO:0001340] flowers [PO:0009046] on one individual [EFO:0000542] and across multiple individuals in a population [PCO:0000001].;Plant sex type, indicating, for a taxon the type of flowers (male, female, and hermaphroditic) present on individual plants versus separate plants within a population.
  • type: categorical
  • allowable values:
    • androdioecious: Taxa in which hermaphroditic and staminate (male) plants coexist within a population.
    • andromonoecious: Taxa with hermaphrodite flowers and staminate flowers (producing just male gametes) on the same plant but no pistillate (female) flowers.
    • dioecious: Taxa that produce male and female flowers on separate plants.
    • gynodioecious: Taxa in which hermaphroditic and pistillate (female) plants coexist within a population.
    • gynomonoecious: Taxa with hermaphrodite flowers and pistillate flowers (producing just female gametes) on the same plant but no staminate (male) flowers.
    • hermaphrodite: Taxa with flowers that produce both male and female gametes; bisexual.
    • monoecious: Taxa that have both male and female flowers on the same plant but an individual flower produces only male or female gametes.
    • polygamodioecious: Taxa where some flowers on a plant are hermaphrodite and other are pistillate (female) or staminate (male).
    • polygamonoecious: Taxa where some plants in a population have hermaphrodite and pistillate (female) flowers and others have hermaphrodite and staminate (male) flowers.
    • polygamous: Taxa with both unisexual and bisexual flowers on the same plant.
    • trioecious: Taxa where plants in a population have either hermaphroditic (bisexual) flowers, pistillate (female) flowers, or staminate (male) flowers.
    • unknown: Inadequate information to assign a sex type.

parasitic

  • label: Plant parasitism status
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait that indicates if a plant is involved in a parasitic [ECOCORE:00000146] symbiosis [ECOCORE:00000022] where it forms haustorium [GO:0085035] on a host [GO:0018995] plant and the host plant provides all or some of its high energy [PATO:0001021] carbon [CHEBI:27594] compounds [NCIT:C43366], mineral nutrition (nutrients [CHEBI:33284]) and/or water [CHEBI:15377].;Trait indicating whether a plant forms haustoria and obtains part or all of its energy-rich carbon compounds (photosynthetic sugars), mineral nutrition and/or water directly from another plant (its host) in an interaction that does not benefit the host plant.
  • type: categorical
  • allowable values:
    • hemiparasitic: Plant that is photosynthetic but augments its energy, mineral nutrition, and /or water supply through parasitism. This trait value does not distinguish between obligate hemiparasites, plants which require a host for at least part of their life cycle, and facultative hemiparasites, plants which can grow and reproduce without a host.
    • hemiparasitic_facultative: Plant that is photosynthetic and can grow and reproduce without a host but augments its energy, mineral nutrition, and /or water supply through parasitism.
    • hemiparasitic_obligate: Plant that is photosynthetic but requires a host for at least part of its life cycle.
    • holoparasitic: Plant that obtains all (or nearly all) of its energy through parasitism.
    • not_parasitic: In the context of this trait, a plant that does not acquire energy, mineral nutrition, and/or water via haustoria connected to another plant. It can include species that form a symbiosis, including a parasitic interaction, with a fungal partner.
    • parasitic: Plant that obtains some energy, mineral nutrition and/or water through parasitism. This term is used if a dataset does not specify if a plant is a hemiparasite or a holoparasite.
    • root_parasitic: Parasitic plant whose haustoria penetrate the root of a host plant.
    • stem_parasitic: Parasitic plant whose haustoria penetrate the stem of a host plant.

fruit_colour

  • label: Fruit colour
  • description: A fruit morphology trait [TO:0002629] which is the primary [NCIT:C25251] colour [PATO:0000014] of the exocarp [PO:0009085] of a mature [PATO:0001701] fruit [PO:0009001].;Human visible dominant colour of a fresh fruit’s exocarp (outermost layer) at maturity.
  • type: categorical
  • allowable values:
    • black: Fruit’s exocarp is black.
    • blue: Fruit’s exocarp is blue.
    • brown: Fruit’s exocarp is brown.
    • cream: Fruit’s exocarp is cream.
    • green: Fruit’s exocarp is green.
    • grey: Fruit’s exocarp is grey.
    • orange: Fruit’s exocarp is orange.
    • pink: Fruit’s exocarp is pink.
    • purple: Fruit’s exocarp is purple.
    • red: Fruit’s exocarp is red.
    • white: Fruit’s exocarp is white.
    • yellow: Fruit’s exocarp is yellow.

fruit_dehiscence

  • label: Fruit dehiscence
  • description: A fruit morphology trait [TO:0002629] which indicates whether mature [PATO:0001701] fruits [PO:0009001] are dehiscent [GO:0009900].;Binary variable, dividing fruits into ‘dehiscent’ versus ‘indehiscent’ based on whether the individual fruit units are dehiscent when mature.
  • type: categorical
  • allowable values:
    • dehiscent: Fruits are dehiscent when mature.
    • indehiscent: Fruits are indehiscent when mature.

fruit_type

  • label: Fruit type
  • description: A fruit morphology trait [TO:0002629] which is the botanical classification [NCIT:C25161] or type [NCIT:C25284] of a fruit [PO:0009001].;Botanical fruit type classification.
  • type: categorical
  • allowable values:
    • achene: A simple, dry, indehiscent fruit with a single seed that is attached to the pericarp at one point only. (Synonyms, cypsela)
    • anthocarp: A ripened ovary surrounded by attached accessory floral tissues; the floral tissues may be fleshy or dry. (Synonyms, accessory_fruit, pseudocarp, diclesium)
    • berry: A simple, indehiscent fruit with one to many seeds embedded in a fleshy pericarp.
    • capsule: A dry, dehiscent fruit composed of two or more united carpels; different types of capsules distinguished by type of dehiscence (e.g. valves, slits, pores).
    • caryopsis: A simple, dry, indehiscent fruit with a single seed which is fused to the pericarp at one point only; the fruit type of all Poaceae (Synonyms, grain)
    • compound_fruit_other: A fruit derived from more than one flower. (Synonyms, aggregate_fruit)
    • drupe: A simple indehiscent fruit with a hard, stony endocarp and a fleshy mesocarp.
    • follicle: A dry, dehiscent fruit derived from a single carpel that splits along one suture.
    • legume: A dry, dehiscent fruit derived from a single carpel that splits along two sutures; applies to Fabaceae. (Synonyms, pod)
    • legume_indehiscent: A dry, indehiscent fruit derived from a single carpel; applies to Fabaceae.
    • mericarp: A fruitlet (seed-containing portion of a fruit) that is a half-carpel; the individual fruitlets separate from one another at maturity; a specific type of schizocarp. (In some datasets, term likely refers to any fruitlet derived from a portion of a schizocarp that separates at maturity, not explicitly just a half-carpel.)
    • multiple_fruit: A fruit composed of several fruitlets developing from a single flower (e.g. raspberries, magnolias); fruits developing from more than one flower are included under compound (aggregate) fruits. (Synonyms, apocarp)
    • nut: A simple, dry, indehiscent fruit with a single seed and a hard pericarp.
    • nutlet: A diminutive of nut, referring to a simple, dry, indehiscent structure with a single seed that might be an entire fruit (nut; monocarp) or a portion of a fruit, such as the individual components derived from an apocarp (fruit with multiple unfused ovaries) or schizocarp (fruit with multiple ovaries that separate at maturity).
    • pepo: A fleshy fruit derived from a single ovary that is divided into several carpels, has multiple seeds, and has a tough rind when mature.
    • pome: A fleshy fruit with the bulk of the fleshy tissue derived from flora tissue that surrounds the inferior ovary.
    • pyrene: A fleshy fruit in which the seed(s) are covered by a hard layer of endocarp.
    • samara: A winged, dry fruit.
    • schizocarp: Category that includes all fruits derived from a compound pistil that separate into individual carpellary constituents at maturity; broad terms that includes dry and fleshy fruits, some with accessory tissues, others without. The individual seed-containing units that a schizocarp breaks apart into include mericarps, nutlets, and fruitlets.
    • silique: A type of capsule (dry, dehiscent fruit) where the fruit is divided into two parts by a thin partition; more than three times as long as wide; applies to Brassicaceae (Synonyms, silicle, siliqua)
    • strobilus: A modified reproductive shoot system found in many nonflowering plants, especially gymnosperms, comprised of overlapping bracts (scales). (Synonyms, cone)
    • syconium: A fleshy compound fruit, in which the fruitlets are enclosed in accessory material, such as the hollow-centered fruits of Moraceae. (Synonyms, synconium, fig)
    • syncarp: A compound fruit produced by the adhesion of the fruits from several flowers.
    • utricle: A bladdery or inflated dry fruit with a single seed; essential an achene in which the pericarp is significantly larger than the mature seed.

leaf_margin

  • label: Leaf margin
  • description: A leaf shape trait [TO:0000492] which is the edge shape [PATO:0002267] of the leaf margin [PO:0020128], in particular the presence or absence of leaf lamina teeth [PO:0025518] and the shape [PATO:0000052] of the leaf lamina teeth (serrate, dentate, or crenate).;Shape of the leaf margin, referencing incisions or projections in the outermost 1/8 of the leaf lamina.
  • type: categorical
  • allowable values:
    • entire: Leaf margin lacks projections or indentations that extend less than 1/8 of the distance to the midrib.
    • toothed: Leaf margin has small projections or lobes along the edge of the leaf lamina. This is a broad term that includes serrate, dentate, and crenulate.
    • toothed_crenate: Type of toothed leaf margin with teeth that are rounded or obtuse, rather than having a sharp apex. [PO:0025519]
    • toothed_dentate: Type of toothed leaf margin with sharp teeth that point outwards at right angles to the leaf margin.
    • toothed_serrate: Type of toothed leaf margin with sharp teeth that are angled toward the tip of the leaf. (Synonyms, saw-toothed)

leaf_shape

  • label: Leaf shape
  • description: A leaf shape trait [TO:0000492] which is the shape [PATO:0000052] and form of a leaf [PO:0025034], considering the ratio [PATO:0001470] of leaf length [PATO:0000122] to leaf width [PATO:0000921], the position [PATO:0000140] of the maximum [NCIT:C25564] width dimension relative to the length axis [NCIT:C25154], and the curvature [PATO:0001591] of the leaf margin [PO:0020128].;Shapes of a leaf or leaflet.
  • type: categorical
  • allowable values:
    • acicular: A 2-dimensional or 3-dimensional shape with a needlelike tip, often round in cross-section, with margins straight and parallel, and a length:width ratio greater than 12:1. (Synonyms, needle-shaped)
    • cordate: A 2-dimensional heart shape, approximately ovate with a cordate base, with the petiole attaching at the notch between the lobes. (Synonyms, cordiform)
    • deltate: A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio of 1:1 and the petiole attaching along one of the broad sides.
    • elliptical: A 2-dimensional shape that is an oval, with curved margins, lacking a distinct point, and a length:width ratio between 2:1 and 3:2. (Synonyms, oval)
    • ensiform: A 2-dimensional shape that is sword-shaped with a pointed tip and a length:width ratio greater than 12:1. (Synonyms, sword-shaped)
    • falcate: A 2-dimensional sickle-shape, approximately lanceolate to linear and curved on one side. (Synonyms, scimitar-shaped, sickle-shaped, falciform, crescent-shaped)
    • filiform: A 2-dimensional shape that is long, thin, and typically flexuous, threadlike, filamentous.
    • lanceolate: A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The petiole attaches near the wide, curved portion of the leaf lamina.
    • linear: A 2-dimensional shape that is very long and narrow, has parallel sides and a length:width ratio between 12:1 and 6:1.
    • linear_elliptical: A 2-dimensional shape that is a very narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 12:1 and 6:1.
    • narrowly_elliptical: A 2-dimensional shape that is a narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 6:1 and 3:1.
    • narrowly_lanceolate: A 2-dimensional shape that is very narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The petiole attaches near the wide, curved portion of the leaf lamina.
    • narrowly_oblanceolate: A 2-dimensional shape that is very narrowly teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, linear_oblanceolate)
    • narrowly_oblong: A 2-dimensional shape that is long and narrow, has parallel sides and a length:width ratio between 6:1 and 3:1. (Synonyms, lorate)
    • narrowly_obovate: A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped with a length:width ratio between 3:1 and 2:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, oblance-obovate)
    • narrowly_obtrullate: A 2-dimensional shape that is narrowly parallelogram-shaped, widest near the tip, and with a length:width ratio 3:1 to 6:1.
    • narrowly_ovate: A 2-dimensional shape that is narrowly egg-shaped, with a length:width ratio between 3:1 and 2:1. The petiole attaches near the wide, curved portion of the leaf lamina. (Synonyms, lance_ovate, widely_lanceolate)
    • narrowly_rhomboidal: A 2-dimensional shape that is like a narrow diamond, with approximately straight edges, widest near the middle, and a length:width ratio 6:1 to 3:1. The petiole attaches near one of the points.
    • narrowly_triangular: A 2-dimensional shape that is narrowly triangular to wedge-shaped, with a length:width ratio 6:1 to 3:1 and the petiole attaching near the widest portion of the leaf. (Synonyms, triangular)
    • obcordate: A 2-dimensional heart shape, approximately ovate with a cordate apex, and the petiole attaching along the broad, curve side.
    • oblanceolate: A 2-dimensional shape that is narrowly teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
    • oblate: A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 5:6. The petiole attaches along the broader side.
    • oblong: A 2-dimensional shape that is somewhat elongated with parallel sides and a length:width ratio between 2:1 and 3:2.
    • obovate: A 2-dimensional shape that is egg-shaped with a length:width ratio between 2:1 and 3:2. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
    • obtriangular: A 2-dimensional shape that is triangular to fan-shaped or wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the narrowest portion of the leaf. (Synonyms, flabelliform, fan-shaped, wedge-shaped, cuneate)
    • obtrullate: A 2-dimensional shape that is approximately parallelogram-shaped, widest near the tip, and with a length:width ratio 2:1 to 3:2.
    • orbicular: A 2-dimensional shape that is approximately circular. (Synonyms, circular)
    • ovate: A 2-dimensional shape that is egg-shaped, with a length:width ratio between 2:1 and 3:2. The petiole attaches near the wide, curved portion of the leaf lamina.
    • peltate: A 2-dimensional orbicular shape, distinguished by the petiole attaching near the center of the leaf lamina.
    • reniform: A 2-dimensional shape that is roughly kidney-shaped, wider than long with a rounded apex and two rounded, basal lobes that are smoothly concave at their intersection. (Synonyms, kidney-shaped)
    • rhomboidal: A 2-dimensional shape that is approximately diamond or parallelogram-shaped, with approximately straight edges, widest near the middle, and a length:width ratio 2:1 to 3:2. The petiole attaches near one of the points. (Synonyms, rhombic, diamond)
    • spathulate: A 2-dimensional shape that can be approximately oblong, obovate, or oblanceolate in outline with a long attenuate base, like a spoon. (Synonyms, spoon-shaped, clavate)
    • strap-shaped: A 2-dimensional shape that is strap-shaped with a rounded tip and a length:width ratio greater than 12:1.
    • subulate: A 2-dimensional shape that is awl-shaped, approximately narrowly oblong to narrowly triangular with a broad base. (Synonyms, awl-shaped)
    • terete: A 3-dimensional shape that is a long, narrow cylinder approximately circular in cross-section. Terete leaves often taper toward the tip. (Synonyms, cylindrical)
    • triangular: A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the widest portion of the leaf.
    • trullate: A 2-dimensional shape that is approximately parallelogram-shaped, widest near where the petiole attaches (base), and with a length:width ratio 2:1 to 3:2.
    • widely_elliptical: A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 6:5.
    • widely_obovate: A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
    • widely_obtrullate: A 2-dimensional shape that is broadly parallelogram-shaped, widest near the tip, and with a length:width ratio of approximately 6:5.
    • widely_ovate: A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The petiole attaches near the wide, curved portion of the leaf lamina.

plant_photosynthetic_organ

  • label: Plant photosynthetic organ
  • description: A plant structure morphology trait [TO:0000839] which is the primary [NCIT:C25251] organ [PO:0009008] or tissue [CARO:0000043] a plant [PO:0000003] uses for photosynthesis [GO:0015979], most commonly a leaf [PO:0025034], cladode [PO:0025354], or phyllode [PO:0025335].;Trait indicating a plant’s primary photosynthetic organ or tissue.
  • type: categorical
  • allowable values:
    • cladode: A photosynthetic shoot, regardless of form, that functions as a leaf. For example, cacti pads and Casuarina articles. (Synonyms, article, phylloclade, cladophyll)
    • leaf: A true leaf, including both laminar and needle-shaped leaves.
    • non-photosynthetic_plant: A plant that lacks any photosynthetic organs.
    • phyllode: A flattened bladelike petiole. (Synonyms, switch plant)

leaf_lamina_posture

  • label: Leaf lamina posture (leaf 3-dimensional shape)
  • description: A leaf shape trait [TO:0000492] which is the 3-D shape [PATO:0002266] of the surface [NCIT:C25245] of the leaf lamina [PO:0020039], including its degree of curvature [PATO:0001591] and whether it is bent [PATO:0000617].;The 3-dimension shape of the surface of the leaf lamina, including its degree of curvature and bending.
  • type: categorical
  • allowable values:
    • concave: A longitudinal posture term indicating the leaf lamina is curved upward along the central axis, such that the upper leaf lamina surface is concave.
    • conduplicate: A longitudinal posture term indicating the leaf lamina is folded upward along the central axis. (Synonyms, folded)
    • convex: A longitudinal posture term indicating the leaf lamina is curved downward along the central axis, such that the lower leaf lamina surface is concave. (Synonyms, concavoconvex, planoconvex)
    • convolute: Leaf lamina is contorted. (Synonyms, twisted)
    • flat: The leaf lamina is a 2-dimensional plane.
    • incurved: Leaf tip is curved upward.
    • involute: A longitudinal posture term indicating the outer portions of the leaf lamina are rolled inward or upward.
    • plicate: Leaf margin is pleated, with a series of longitudinal folds.
    • recurved: Leaf tip is curved outward or downward. [PATO:0002211]
    • revolute: A longitudinal posture term indicating the outer portions of the leaf lamina are rolled outward or downward.
    • undulate: The leaf lamina surface is rippled or wavy in its 2-dimensional plane. [PATO:0000967] (Synonyms, wavy, rippled, sinuate)

leaf_base_shape

  • label: Leaf base shape
  • description: A leaf shape trait [TO:0000492] which is the variation in shapes and forms of a leaf base [PO:0020040], especially considering the curvature [PATO:0001591] of the leaf margin [PO:0020128] near where it attaches to the petiole [PO:0020038].;The shape of the basal margin of a leaf.
  • type: categorical
  • allowable values:
    • acute: Two sides of the leaf meet at an angle that is less than 90°.
    • attenuate: Basal leaf margin abruptly incurved (concave), the two sides intersecting at an angle of less than 45°.
    • auriculate: Basal leaf margin with two rounded lobes. The leaf margin above the lobes is concave. The petiole attaches in the incised region between the lobes.
    • cordate: Basal leaf margin with two rounded lobes and the petiole attaches in the incised region between the lobes. The leaf margin above the lobes is rounded.
    • cuneate: Basal leaf margin approximately straight, the two sides intersecting at an angle between 45° to 90°.
    • hastate: Basal leaf margin with two lobes oriented outward, such that they are at approximately 90° relative to the central axis and the leaf margin is fairly flat at the base.
    • oblique: The leaf base is asymmetrical, with the midrib not lying in the middle of the leaf where it attaches to the petiole.
    • obtuse: Two sides of the leaf meet at an angle that is greater than 90°.
    • rounded: Basal leaf margin is rounded.
    • sagittate: Basal leaf margin with two lobes oriented downward, away from apex. The petiole attaches in the concave region between the lobes.
    • sheathing: The leaf base is broad and envelops the stem.
    • truncate: The basal leaf margin is cut straight across with the two sides intersecting at approximately 180°.

leaf_lobation

  • label: Leaf lobation
  • description: A leaf shape trait [TO:0000492] which is the presence or absence of leaf lamina lobes [PO:0025517] along the leaf margin [PO:0020128] and how far the lobes extend toward the midrib [PO:0020139].;A trait that indicates whether a leaf lamina or leaflets display any divisions or dissections on a 2-dimensional plane that extend greater than 1/8 of the distance to the midrib.
  • type: categorical
  • allowable values:
    • lobed: Leaf margin has incisions, either rounded or sharp-tipped, that extend more than 1/8 of the distance to the midrib.
    • lobed_deep: Leaf margin with lobes that extend more than halfway to the midrib. (Synonyms, parted, divided)
    • lobed_shallow: Leaf margin with lobes that extend less than halfway to the midrib. (Synonyms, cleft, sinuate)
    • unlobed: Leaf margin lacks projections or indentations that extend more than 1/8 of the distance to the midrib.

leaf_phyllotaxis

  • label: Leaf phyllotaxis
  • description: A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the phyllotaxy [TO:0006014] of leaves, indicating a leaf’s position [TO:0000541] relative to other leaves at a stem node [PO:0020141].;Leaf arrangement at a stem node, indicating the relative number and position of leaves at a single stem note.
  • type: categorical
  • allowable values:
    • alternate: A single leaf is attached at each stem node.
    • opposite: A pair of leaves are attached opposite to one another at each stem node.
    • whorled: Three or more leaves are attached at each stem node, arranged in a symmetrical pattern around the stem. (Synonyms, verticillate)

leaf_glaucousness

  • label: Leaf glaucousness
  • description: A leaf morphology trait [TO:0000748] which is the presence or absence of epicuticular wax [PO:0025388] on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340], giving the leaf lamina surface a white-coloured [PATO:0000014] or blue-coloured waxy [PATO:0002384] appearance, termed glaucous.;Variable describing whether mature adult phase leaves have a glaucous appearance, the colour due to being covered with a smooth, usually whitish to bluish epicuticular wax that can easily be rubbed off.
  • type: categorical
  • allowable values:
    • glaucous: Mature (fully expanded) leaves on adult plants have a waxy coating, giving them a glaucous appearance. (Synonyms, waxy bloom, pruinose)
    • not_glaucous: Mature (fully expanded) leaves on adult plants do not have a glaucous appearance.

seed_surface_reflectivity

  • label: Seed surface reflectivity
  • description: A seed morphology trait [TO:0000184] which is the reflectivity [PATO:0001297] of the surface [NCIT:C25245] of a mature [PATO:0001701] seed [PO:0009010].;Surface reflectivity of a seed, indicating whether a seed is shiny or dull.
  • type: categorical
  • allowable values:
    • dull: Seed surface appears dull, indicating it has a low reflectivity.
    • shiny: Seed surface appears shiny or glossy, indicating it has a high reflectivity. (Synonyms, glossy, lustrous, polished)

seed_surface_texture

  • label: Seed surface texture
  • description: A seed morphology trait [TO:0000184] which is the texture [PATO:0000150] of the surface [NCIT:C25245] of a mature [PATO:0001701] seed [PO:0009010].;Surface texture of a mature seed, including descriptions of configuration (overall surface patterns of the epidermal cells) and epidermal excrescence (surface patterns caused by structural outgrowths of the epidermis).
  • type: categorical
  • allowable values:
    • bumpy: Seed surface covered with minute, rounded protuberances. (Synonyms, colliculate, verrucate, papillate, tuberculate, undulate)
    • grooved: Seed surface covered with grooves.
    • netted: Seed surface covered with raised, reticulate, interconnected ridges. (Synonyms, reticulate, honey-combed)
    • papery: Seed surface with a thin and papery texture. (Synonyms, chartaceous)
    • pitted: Seed surface pitted. (Synonyms, foveolate, foveate, dimpled, punctate, lacunose)
    • ribbed: Seed surface with longitudinal raised ribs or ridges. (Synonyms, costate, carinate, lineate, fluted, striate, strigose, lineolate, scalariform, ridged)
    • rough: Seed surface rough with no organisation of markings, resembling sandpaper. (Synonyms, scabrous)
    • scaly: Seed surface rough with loose scales differing in direction or not parallel in direction. (Synonyms, scurfy, squarrose)
    • smooth: Seed surface smooth, lacking texture. (Synonyms, glabrous)
    • spiny: Seed surface with spines. (Synonyms, echinulate)
    • wrinkled: Seed surface with a wrinkled, rugose texture. (Synonyms, rugose, rugulose, bullate)

leaf_arrangement

  • label: Leaf arrangement
  • description: A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the position [TO:0000541] and arrangement [NCIT:C61532] of leaves between stem nodes [PO:0020141], indicating how leaves align and cluster along the vegetative shoot system [PO:0025607].;Leaf arrangement across stem nodes, indicating the 3-dimensional arrangement of leaves along the length of the stem.
  • type: categorical
  • allowable values:
    • clustered: When leaves are in any type of cluster, where the type of cluster is not further specified. The leaves may be clustered toward the base of the plant (a rosette), toward the top of the plant, toward the end of branches, or in distinct tuft (fascicles), but more information is not provided.
    • clustered_basal: When leaves are clustered or crowded to the base of the stem, arranged in a rosette or other basal clustering. (Synonyms, rosette)
    • crowded: When internode length varies considerably along a stem, such that leaves appear crowded or tufted. (Synonyms, congested)
    • decussate: When sequential pairs of opposite leaves attach at right angles to the pair above or below.
    • distichous: When alternate or opposite leaves are arranged in two opposite rows.
    • fasciculate: A tight bundle or cluster of leaves attaches to the stem in a single location, generally due to the shortening of internodes. (Synonyms, tufted)
    • imbricate: When leaves lay tightly against the stem and are overlapping.
    • rosette: When leaves radiate from a central point at or near ground level.
    • scattered: When leaves are attached along the entire length of stem.
    • spiral: When alternate leaves are attached in a progressive spiral pattern along the stem.

leaf_margin_posture

  • label: Leaf margin posture
  • description: A leaf shape trait [TO:0000492] which is the degree of curvature [PATO:0001591] or curling [PATO:0000405] of the leaf margin [PO:0020128] along the side of the leaf, at the leaf apex [PO:0020137], and at the leaf base [PO:0020040].;The posture (curvature or curling) of the leaf margin.
  • type: categorical
  • allowable values:
    • flat: Leaf lamina, including margins, is a 2-dimensional plane. [PATO:0000407]
    • involute: Leaf margins along the side of the leaf are rolled upward, such that the margin of the adaxial (upper) leaf surface is rolled inward. (Synonyms, inrolled, incurved)
    • revolute: Leaf margins along the side of the leaf are rolled under or downward, such that the margin of the abaxial (lower) leaf surface is rolled inward. [PATO:0001963] (Synonyms, recurved)
    • undulate: Leaf margins are wavy in their 2-dimensional plane. (Synonyms, wavy, rippled, sinuate)

leaf_pendulousness

  • label: Leaf pendulousness
  • description: A leaf position trait [TO:0000541] which indicates whether leaves [PO:0025034] are pendulous [APD:glossary/40066], hanging loosely downward and possessing the ability to flutter.;Binary variable describing whether leaves are pendulous, hanging loosely downward and possessing the ability to flutter.
  • type: categorical
  • allowable values:
    • not_pendulous: Leaves are not pendulous.
    • pendulous: Leaves are pendulous.

storage_organ

  • label: Storage organ
  • description: A plant structure morphology trait [TO:0000839] which is the type [NCIT:C25284] of storage organ [EFO:0000997] a plant has, where a storage organ is a plant part (organ [PO:0009008], tissue [PO:0009007], or other plant structure [PO:0009011]) whose function [NCIT:C25518] is carbohydrate [CHEBI:16646] (energy [PATO:0001021]) storage.;Plant structures (tissues, organs, or other structures) where a plant stores excess carbohydrates (energy). The stored energy may be used for future growth, including clonal spread and regrowth following an annual unfavourable season, or for regrowth following a disturbance event (including post-fire regeneration or another disturbance event that causes leaf loss). Trait values include both generic terms and quite specific ones to accommodate the variation in detail provided across datasets.
  • type: categorical
  • allowable values:
    • bulb: Below ground storage organ derived from stem and usually leaf tissue.
    • caudex: Swollen stem that usually functions in water storage, although the term can also apply to below ground structures or to swollen roots.
    • corm: Below ground storage organ that is a short, rounded, vertical, fleshy, underground stem.
    • fleshy_underground_organ: Plants that use fleshy, generally underground organs (swellings) to regenerate, but the type of organ is not specified; examples include corms, tubers, bulbs, and caudex.
    • lignotuber: Swollen woody structures are located at the transition between the stem base and root crown of woody shrubs, mallees and small trees, and are formed from stem tissues. Buds are located over the entire structure. No colonisation ability.
    • no_storage_organ: No storage organ present
    • pseudobulb: An above ground storage organ present in many terrestrial and epiphytic orchids created throough the thickening of the stem between leaf nodes.
    • rhizome: A type of continually growing or creeping underground stem, but it is not specified if the rhizome is woody or non-woody.
    • rhizome_fibrous: A type of continually growing or creeping fibrous, non-woody underground stem.
    • rhizome_fleshy: A type of continually growing or creeping fleshy, non-woody underground stem.
    • rhizome_woody: A type of continually growing or creeping woody underground stem.
    • root_tuber: A swollen root the stores nutrients, carbohydrates and water. (Synonyms, taproot tuber)
    • stem_tuber: A swollen underground stem that stores nutrients, carbohydrates and water.
    • storage_organ: Plant has a storage organ, but the type is unknown.
    • tuber: A swollen underground organ that stores nutrients, carbohydrates and water, where it is not specified if the tuber develops from root or stem tissue.

seed_surface_hairs

  • label: Seed surface hairs
  • description: A seed morphology trait [TO:0000184] which is the presence or absence of trichomes [PO:0006504] on the surface [NCIT:C25245] of a mature [PATO:0001701] seed [PO:0009010].;Surface hairiness of a mature seed, including vestiture (trichome cover) and bristles.
  • type: categorical
  • allowable values:
    • glabrous: Seed surface lacks trichomes or bristles.
    • hairs: Seed surface with trichomes or bristles.
    • hairs_branched: Seed surface with branched trichomes or bristles.(Synonyms, stellate, plumose)
    • hairs_simple: Seed surface with simple trichomes or bristles. (Synonyms, tomentose, velutinous, velvety, sericeous)

petiole_width

  • label: Petiole width
  • description: Width [PATO:0000921] of the petiole [PO:0020038].;Width of the petiole.
  • type: numeric
  • units: cm
  • allowable range: 0 - 200 cm

root_system_type

  • label: Root system type (presence of taproot)
  • description: A root system morphology trait [TO:0000985] which indicates if the root system [PO:0025025] is one where the primary root [PO:0020127] remains the dominant root, developing into a taproot, a long [PATO:0000573], thick [PATO:0000591] root with few branches, or whether the lateral roots [PO:0020121] are of similar length to the primary root.;Variable describing whether a plant has a tap root.
  • type: categorical
  • allowable values:
    • tap_root_absent: Tap root not present.
    • tap_root_present: Tap root present.
    • tap_root_sometimes_present: Tap root sometimes present.

leaf_N_per_dry_mass

  • label: Leaf nitrogen (N) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of nitrogen in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.5 - 100 mg/g

leaf_area

  • label: Leaf area
  • description: A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of the leaf lamina [PO:0020039] and petiole [PO:0020038]. For some species the area of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335].;The 2-dimensional projected (scanned) area of the surface of a leaf or a leaf analogue (leaflet, cladode, phyllode).
  • type: numeric
  • units: mm2
  • allowable range: 0.1 - 1e+07 mm2

leaf_delta13C

  • label: Leaf stable carbon isotope composition (delta13C)
  • description: Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of leaves [PO:0025034].;Carbon stable isotope ratio, d13C, of leaves.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -50 - 0 {delta}[ppth]

leaf_mass_per_area

  • label: Leaf mass per area
  • description: A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to the leaf [PO:0025034] area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038].;The ratio of leaf dry mass to leaf area, measured on an entire leaf including the petiole; LMA. (The inverse of specific leaf area, SLA.)
  • type: numeric
  • units: g/m2
  • allowable range: 1 - 3000 g/m2

leaf_photochemical_reflectance_index

  • label: Photochemical reflectance index (PRI)
  • description: A remote sensing [EnvThes:21762] index [NCIT:C25390] of the efficiency [PATO:0001029] with which a whole plant [PO:0000003] uses photosynthetically active radiation [ENVO:01001864], calculated from the reflectance [EnvThes:21580] of radiation [ENVO:01001023] off the plant canopy [ENVO:01001242] at 531 nm (a xanthophyll [CHEBI:27325] band) and 570 nm (a reference band): PRI = (R531 - R570) / (R531 + R570). It is recorded with a spectroradiometer [MOSAIC:00000974] and a higher value indicates a higher radiation use efficiency.;Photochemical reflectance index measures plant responses to stress and the efficiency with which it uses photosynthetically active radiation, by indicating changes in carotenoid pigments in live foliage. It is calculated from the reflectance of radiation off the plant canopy at 531 nm (a xanthophyll band) and 570 nm (a reference band): PRI = (R531 - R570) / (R531 + R570), with a higher value indicating higher radiation use efficiency.
  • type: numeric
  • units: {dimensionless}
  • allowable range: -1 - 1 {dimensionless}

leaf_water_band_index

  • label: Water band index
  • description: A remote sensing [EnvThes:21762] index [NCIT:C25390] of whole plant [PO:0000003] water status, which is the ratio [PATO:0001470] of the reflectance [EnvThes:21580] of radiation [ENVO:01001023] off the plant canopy [ENVO:01001242] at 970 nm to the reflectance at 900 nm, recorded with a spectroradiometer [MOSAIC:00000974], where a higher value indicates a higher canopy water [CHEBI:15377] content.;Water band index measures whole plant water status. It is the ratio of the reflectance of radiation off the plant canopy at 970 nm to the reflectance at 900 nm; a higher value indicates a higher canopy water content.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 1 - 1.1 {dimensionless}

modified_NDVI

  • label: Modified normalized difference vegetation index (modified NDVI)
  • description: A remote sensing [EnvThes:21762] index [NCIT:C25390] of whole plant [PO:0000003] vegetation status, which is a ratio [PATO:0001470] that considers the relative reflectance [EnvThes:21580] of red light versus short-wave infrared radiation (SWIR) [ENVO:21001214] off the plant canopy [ENVO:01001242], where a higher value indicates greater vegetation health. The formula is: (SWIR - Red) / (SWIR + Red).;A remote sensing index of whole plant vegetation status, which is a ratio that considers the relative reflectance of red light versus short-wave infrared radiation (SWIR) off the plant canopy, where a higher value indicates greater vegetation health. The formula is: (SWIR - Red) / (SWIR + Red).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.1 - 1 {dimensionless}

wood_density

  • label: Wood density
  • description: A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of a unit of wood [NCIT:C93062] from a stem [PO:0009047] assessed after drying to its volume [PATO:0000918] assessed without drying.;The ratio of wood dry mass to wood fresh volume, measured on a stem segment without bark.
  • type: numeric
  • units: mg/mm3
  • allowable range: 0.05 - 1.4 mg/mm3

seed_dry_mass

  • label: Seed dry mass
  • description: A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a mature [PATO:0001701] seed [PO:0009010].;Dry mass of a mature seed, including both oven dried and air-dried samples.
  • type: numeric
  • units: mg
  • allowable range: 1e-05 - 1e+06 mg

seed_germination_treatment

  • label: Seed germination treatment
  • description: The treatment [EFO:0000727] or experimental conditions [XCO:0000000] required for a high [NCIT:C25227] proportion [PATO:0001470] of seeds [PO:0009010] to germinate [GO:0009845] .;Treatment required for seed to display high germination rates. The data included in AusTraits are mostly from seed banks that have recorded the protocols they use to encourage seeds to germinate. The treatments may therefore be only loosely connected to the cues required for a species’ to break dormancy.
  • type: categorical
  • allowable values:
    • after_ripening: Seed germinates more readily after a period of warm, dry storage.
    • chemical_treatment: Seed germinates more readily after a chemical treatment.
    • cold: Seed germinates more readily after cold treatment.
    • dark: Seed germinates more readily after being stored in the dark.
    • heat: Seed germinates more readily after being heated.
    • heat+smoke: Seed germinates more readily after a treatment that merges heat and smoke.
    • heat_lethal: Seed is heat sensitive. (A fire response, more than a treatment to break dormancy.)
    • imbibe_hot: Seed germinates more readily after being soaked in hot water; water temperature usually between 70 deg C to just below boiling.
    • leaching: Seed germinates more readily after leaching out certain chemicals.
    • light: Seed germinates more readily after light exposure.
    • mycorrhizae: Seed germinates more readily if infected by a mycorrhizal symbiont.
    • none: No treatment required for seed to display high germination rates.
    • rain: Seed germinates more readily after rain.
    • scarify: Seed coat is removed, nicked, or otherwise punctured to increase germination.
    • smoke: Seed germinates more readily after being subjected to a smoke treatment; includes smoked water.
    • stratification: Seed germinates more readily after being stored in moist conditions; not specified if the treatment was cold stratification or warm stratification.
    • stratification_cold: Seed germinates more readily after a period of cold, wet storage.
    • stratification_warm: Seed germinates more readily after a period of warm, wet storage.

leaf_dry_matter_content

  • label: Leaf dry matter content (LDMC)
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of leaf dry mass [TO:0001014] to leaf [PO:0025034] fresh mass [APD:glossary/40044].;The ratio of leaf dry mass to leaf fresh mass; LDMC.
  • type: numeric
  • units: g/g
  • allowable range: 0.001 - 2 g/g

dispersal_syndrome

  • label: Diaspore dispersal syndrome
  • description: A classification [NCIT:C25161] based on a taxon’s [NCIT:C40098] seed morphology traits [TO:0000184] or fruit morphology traits [TO:0002629] that indicates the taxa (biotic vectors) or abiotic mechanisms [NCIT:C25343] that most likely disperse [OMIT: 0027297] seeds [PO:0009010] and fruit [PO:0009001] away from the parent plant.;Plant dispersal syndromes are generally assigned to species based on suites of diaspore characteristics associated with specific mechanisms of dispersal. Trait values are mostly given as the formal dispersal syndromes ending in ‘chory’, a suffix meaning ‘a specified method of plant dispersal’.
  • type: categorical
  • allowable values:
    • anemochory: Diaspore is dispersed by wind. (Synonyms, wind)
    • anthrochory: Diaspore is dispersed by humans, either intentionally or unintentionally.
    • atelochory: Diaspore dispersal is prevented. (Synonyms, antitelochory)
    • autochory: Diaspore is dispersed by methods originating from the parent plant or diaspore.
    • ballistic: Seeds are launched away from the plant by explosion as soon as the seed capsule opens.
    • barochory: Diaspores are dispersed without assistance. (Synonyms, gravity, mobile, unassisted, passive)
    • chamaechory: Diaspore is dispersed by being rolled along ground surface by wind.
    • endozoochory: Diaspore is ingested by animals, either intentionally or accidentally, then transported before being deposited. (Synonyms, ingestion)
    • epizoochory: Diaspore is dispersed by accidentally attaching itself to the outside of an animal vector. (Synonyms, ectozoochory, exozoochory)
    • hydrochory: Diaspore is dispersed on the surface of water. (Synonyms, water)
    • myrmecochory: Diaspores have elaiosomes (specialised nutritious appendages) that make them attractive for capture, transport and use by ants or related insects.
    • undefined: Dispersal mechanism unknown. (Synonyms, unknown)
    • zoochory: Diaspore is dispersed by animals, by an undescribed mechanism.

leaf_thickness

  • label: Leaf thickness
  • description: The thickness [PATO:0000915] of the leaf lamina [PO:0020039], measured away from the midrib [PO:0020139].;Thickness of the leaf lamina, generally measured away from the midrib.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 51 mm

sapwood_specific_hydraulic_conductivity_theoretical

  • label: Theoretical sapwood specific hydraulic conductivity (Ks)
  • description: A stem [PO:0009047] water transport [GO:0006833] trait which is a theoretical [MESH:D008962] measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a stem under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), calculated from the Hagen Poiseuille equation [hupson:SCAIVPH_00000131], then normalised to the diameter [PATO:0001334] of water transport tissues [CARO:0000043] and accounting for the stem length [PATO:0000122]. It is the calculated sapwood hydraulic conductance, divided by the sapwood [PO:0004513] cross-sectional [NCIT:C63795] area [PATO:0001323] (m2), multiplied by the path length measured (m2).;Theoretical Ks; A calculation of the theoretical flow rate of water (kg/s) along a stem for a given drop in pressure (MPa), normalised to the microscopically determined cross-sectional area without bark and pith (the diameter of water transport tissues). The calculation is based on Hagen-Poiseuille’s law.
  • type: numeric
  • units: kg/m/MPa/s
  • allowable range: 0.05 - 5000 kg/m/MPa/s

stem_vessel_density

  • label: Stem xylem vessel density
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of xylem vessels [PO:0025417] in a stem [PO:0009047] cross-section [NCIT:C63795] to stem cross-sectional area [PATO:0001323].;The ratio of the count of xylem vessels in a stem cross-section to stem cross-sectional area.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 0.4 - 2000 {count}/mm2

stem_vessel_diameter

  • label: Stem xylem vessel diameter
  • description: A vascular tissue morphology trait [TO:0000470] which is the diameter [PATO:0001334] of xylem vessels [PO:0025417] in a stem [PO:0009047].;Diameter of xylem vessels in a stem.
  • type: numeric
  • units: um
  • allowable range: 1 - 1000 um

stem_vessel_lumen_fraction

  • label: Stem xylem vessel lumen fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of total xylem vessel [PO:0025417] lumen [PO:0025117] cross-sectional [NCIT:C63795] area [PATO:0001323] to stem [PO:0009047] cross-sectional area.;The fraction of a stem cross-section comprised of lumen and calculated as the ratio of the total cross-sectional area comprised of xylem vessels (lumen fraction) to the total stem cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.001 - 1 mm2/mm2

leaf_water_use_efficiency_intrinsic

  • label: Intrinsic water use efficiency (WUEi)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its stomatal conductance [EnvThes:21211], with photosynthetic rate and stomatal conductance determined by gas exchange measurements.;The ratio of photosynthetic rate (A; CO2 assimilation rate) to stomatal conductance (gs; gaseous diffusion), with photosynthetic rate and stomatal conductance determined from gas exchange measurements.
  • type: numeric
  • units: umol{CO2}/mmol{H2O}
  • allowable range: 0.001 - 2 umol{CO2}/mmol{H2O}

stem_vessel_diameter_hydraulic

  • label: Stem xylem vessel hydraulic mean diameter
  • description: A vascular tissue morphology trait [TO:0000470] which is a weighted arithmetic mean [STATO:0000398] diameter [PATO:0001334] of xylem vessels [PO:0025417] in a stem [PO:0009047], reflecting the greater relative contribution of wider vessels to hydraulic conductivity [EnvThes:22133] as described by the Hagen-Poiseuille equation [hupson:SCAIVPH_00000131] .;Hydraulic diameter (or hydraulically weighted diameter) is a weighted arithmetic mean diameter of xylem vessels in a stem, reflecting the greater contribution of wide vessels to actual hydraulic conductivity. It is based on the Hagen-Poiseuille law, a fluid dynamics equation that describes laminar flow through a tube. It shows that a few large conduits transport an equal amount of water as many small ones, as flow rate is proportional to the 4th power of the conduit’s diameter.
  • type: numeric
  • units: um
  • allowable range: 1 - 500 um

stem_vessel_multiple_fraction

  • label: Stem xylem vessel multiple fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of groups of xylem vessels [PO:0025417] to the total count of xylem vessels, with a value approximating 1 indicating a tendency towards solitary vessels.;Vmf; A vessel grouping index, calculated as the number of groups of vessels divided by the total number of vessels, where vessels are considered to be part of the same group if they are in contact with one another. A Vmf value approximating 1 denotes a tendency towards solitary vessels while a smaller value indicates a tendency of vessels to occur in clusters.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.001 - 1 {dimensionless}

stem_xylem_vulnerability_index

  • label: Xylem vulnerability index
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of a stem’ [PO:0009047] average xylem vessel [PO:0025417] diameter [PATO:0001334] to the stem’s xylem vessel count density [ECSO:00001160].;The ratio of a stem’s average xylem vessel diameter to the stem’s xylem vessel density, an index of sap conductance and xylem vessel susceptibility to vessel cavitation.
  • type: numeric
  • units: um.m2/{count}
  • allowable range: 0.05 - 1000 um.m2/{count}

leaf_hairs_adult_leaves

  • label: Mature leaf hairiness
  • description: A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340].;Variable describing whether mature (fully expanded) adult phase leaves on adult plants have hairs (trichomes).
  • type: categorical
  • allowable values:
    • glabrous: Mature (fully expanded) leaves on adult plants do not have hairs (trichomes).
    • hairy: Mature (fully expanded) leaves on adult plants have hairs (trichomes).

clonal_spread_mechanism

  • label: Clonal spread mechanism
  • description: A plant structure morphology trait [TO:0000839] which is the mechanism [NCIT:C25343] plants use for clonal growth [EnvThes:21032] (asexual reproduction [GO:0019954]) or for regeneration [GO:0031099] after a disturbance event [EnvThes:20373];Mechanisms and structures (tissues, organs, or other structures) a plant uses to spread vegetatively, including the production of clones.
  • type: categorical
  • allowable values:
    • aboveground_clonal: Vegetative buds or plant fragments that can disperse and produce new plants (including axillary buds, bulbils and turions).
    • adventitious_roots: Adventitious roots can form at nodes on the main roots or aboveground from nodes on stems.
    • belowground_clonal: Plants use an unspecified underground organ for clonal spread.
    • bulb: Below ground storage organ derived from stem (and sometimes leaf) tissue, that divides into new individuals.
    • corm: Below ground organ derived from stem tissue that can function either for vegetative spread or to regenerate following a disturbance.
    • non_clonal: Plant does not display clonal growth.
    • rhizome: Plant has an underground stem that has stem buds and forms adventitious roots at nodes; this stem may be woody or non-woody. Rhizomes are both a mechanism for vegetative spread and a strategy to survive fire.
    • root_buds: Root buds that produce vertical suckers.
    • root_tuber: A swollen root that functions as a storage organ and has buds that allow clonal growth.
    • stem_tuber: A swollen underground stem that functions as a storage organ and has buds that allow clonal growth.
    • stolon: Plants having horizontal branches (stems) from the base of the plant that produce new plants from buds at its tip or nodes.
    • viviparous: Plant produces clonal offspring that grow while still attached to the parent plant.

plant_climbing_mechanism

  • label: Plant climbing mechanisms
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait that indicates stem [PO:0009047] and leaf [PO:0025034] structures [PATO:0000141] and physical mechanisms [NCIT:C25343] a plant uses to climb [NCIT:C81002] up other plants or substrates.;Mechanisms plants use to climb, including both specialised leaf and stem adaptations and stem growth habits.
  • type: categorical
  • allowable values:
    • adventitious_roots: Roots arising from stem nodes which cling to a host plant.
    • hooks: Curved stem outgrowths which allow climbers to cling to a host plant.
    • scrambling: Stem growth habit, where the plant stems are too weak to support themselves, and instead lean on and are supported by host vegetation. There are no specialised structures employed.
    • spines: Straight stem outgrowths (including true spines as well as thorns and prickles) which allow climbers to cling to a host plant.
    • tendrils: Modified leaves, petioles, or specialised axillary shoots that coil tightly, forming a specialised structure for climbing.
    • twining: Stem growth habit, where the plant stems twist or coil around a host plant to provide support.

resprouting_capacity

  • label: Post-fire resprouting capacity
  • description: A fire response trait which indicates whether mature [PATO:0001701] plants [PO:0000003] from a taxon [NCIT:C40098] or population [PCO:0000001] are killed or have vegetative meristematic tissue [PO:0006079] (buds [PO:0000055]) that resprout [Cerrado:fire#PlantResprout] following a fire [ENVO:01000786] that causes 100% leaf scorch, with fire severity index [EnvThes:22120] an important contextual variable.;Trait indicating whether mature individuals of a taxon are killed, resprout, or have a mixed survival response when exposed to fire that causes 100% leaf scorch. Species where at least 70% of plants resprout are designated resprouters, while those where fewer than 30% of plants resprout are designated as fire killed. Species with an intermediate response have a mixed fire response, and are coded as fire_killed resprouts.
  • type: categorical
  • allowable values:
    • fire_killed: Fewer than 30% of plants resprout following a fire with 100% leaf scorch. (Synonyms, fire-sensitive)
    • partial_resprouting: Between 30-70% of plants resprout following a fire with 100% leaf scorch.
    • resprouts: More than 70% of plants resprout following a fire with 100% leaf scorch.

seed_viability

  • label: Seed viability
  • description: The proportion [PATO:0001470] of mature [PATO:0001701] seeds [PO:0009010] that are capable of germinating [GO:0009845] under suitable [NCIT:C116358] environmental conditions [ENVO:01000203] or experimental conditions [XCO:0000000] .;Proportion of mature seeds in a sample that are capable of germinating under suitable conditions.
  • type: numeric
  • units: {count}/{count}
  • allowable range: 0 - 1 {count}/{count}

seedbank_longevity

  • label: Seedbank longevity
  • description: The half-life of the seedbank [MESH:D000068098], with is the time interval (duration [PATO:0001309]) from when seeds [PO:0009010] in a seedbank mature [PATO:0001701] until more than half the seeds in a seedbank are no longer able to germinate [GO:0009845].;The half-life of the seedbank, which is the length of time (duration) from when seeds in a seedbank mature until more than half the seeds in a seedbank are no longer able to germinate.
  • type: numeric
  • units: a
  • allowable range: 0.01 - 1000 a

bud_bank_location

  • label: Bud bank location
  • description: The position [PATO:0000140] and distribution [PATO:0000060] of vegetative meristematic tissue [PO:0006079] (buds [PO:0000055]) on a plant [PO:0000003]. These may be buds that allow a plant to regrow following an unfavourable season or buds that survive [EnvThes:21064] a disturbance event [EnvThes:20373] such as a fire, cyclone or drought. In particular, they may be meristematic tissue that survives a fire [ENVO:01000786] and allows the plant to resprout [Cerrado:fire#PlantResprout] following the fire.;The location of buds (meristematic tissue) on a plant that allows it to resprout following a fire, annual unfavourable season, or other disturbance event, such as a drought or cyclone. Trait values include both generic terms and quite specific ones to accommodate the variation in detail provided across datasets.
  • type: categorical
  • allowable values:
    • apical_buds: Plants resprout from above-ground apical meristems. Also termed a caudex. Term refers to palm-like plants with aerial buds that resprout, including grass trees, palms, tree ferns, and pandanus. (Synonyms, aerial apical sprouter)
    • basal_buds: Plants resprout from its base, with buds originating from stem tissue (e.g. lignotuber) and root tissue (root crown) not distinguished between. Most buds designated as being basal are likely to be resprouts from a lignotuber, but this trait value is used if the dataset reported basal buds rather than basal stem buds or lignotuber.
    • basal_stem_buds: Plants resprout from the base of the stem. Australian plants with basal stem buds usually have a lignotuber, documented in the trait storage_organ. (Synonyms, rootstock, lignotuberous resprouter)
    • bud-bearing_root: Root buds produce vertical suckers (from root crown) or adventitious roots. (Synonyms, root-suckering resprouter, adventitious buds)
    • epicormic_buds: Plants resprout from the stem or trunk.
    • fleshy_underground_organ: Plants resprouts from buds on a fleshy underground organ that is a taproot tuber, bulb, corm, stem tuber, root tuber, or belowground caudex. (Synonyms, geophyte)
    • none: Plants lack regenerative buds.
    • rhizome: Plants resprout from buds on a rhizome, where it is not specified if the rhizome is woody or non-woody.
    • rhizome_non_woody: Plants resprout from buds on a fleshy rhizome, rhizophore, or fibrous rhizome.
    • rhizome_woody: Plants resprout from buds on a woody rhizome.
    • root_crown: Plants resprout from the root crown, the root tissue at the root-stem transition.
    • stem_buds: Plants resprout from stem buds; this trait value is separate to epicormic buds. (Synonyms, aerial axillary resprouter)
    • stem_tuber: Plants resprout from buds on a stem tuber.
    • stolon: Horizontal branches (stems) resprout from the base of the plant.
    • underground_buds: Plants that resprout from underground organ, but the type of organ isn’t specified.

post_fire_recruitment

  • label: Post-fire recruitment
  • description: A fire response trait which is the magnitude [NCIT:C49158] of seed [PO:0009010] germination [GO:0009845] and seedling establishment [EnvThes:20410] after a fire [ENVO:01000786].;A trait indicating the magnitude of seed germination and seedling establishment following a fire.
  • type: categorical
  • allowable values:
    • post_fire_recruitment: Plants that germinate post-fire.
    • post_fire_recruitment_absent: Plants that do not show increased seeding following fire.

leaf_hydraulic_vulnerability

  • label: Leaf xylem pressure, 50% lost conductance (P50)
  • description: The tension [PATO:0002284] (water pressure) (water potential [EnvThes:22311]) in leaf [PO:0025034] xylem vessels [PO:0025417] at which the hydraulic conductance [EnvThes:22133] has declined by 50% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;A measure of leaf hydraulic vulnerability, which is the leaf xylem water pressure at which leaf hydraulic conductance has declined by 50% from the mean maximum rate of hydraulic conductance.
  • type: numeric
  • units: MPa
  • allowable range: -40 - 0 MPa

leaf_specific_hydraulic_conductance

  • label: Leaf specific hydraulic conductance (kleaf)
  • description: A leaf [PO:0025034] water transport [GO:0006833] trait which is a measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a leaf under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), normalised to the leaf area [PATO:0001323]. It is calculated as the ratio [PATO:0001470] of the water flow rate through the leaf (mmol/s) to the water potential gradient gradient from the petiole [PO:0020038] to mesophyll [PO:0006070] substomatal cavities (MPa), divided by the leaf area (m2).;kleaf; Measure of how efficiently water is transported through a leaf. It is calculated as the ratio of water flow rate through the leaf (mmol/s) to the water potential gradient from the petiole to mesophyll substomatal cavities (MPa), divided by the leaf area (1/m2). As this is a measure of “conductance” not “conductivity” the measurement is not normalised to the path length (the leaf length).
  • type: numeric
  • units: mmol/m2/MPa/s
  • allowable range: 0.1 - 100 mmol/m2/MPa/s

leaf_turgor_loss_point

  • label: Leaf turgor loss point
  • description: A parameter [STATO:0000034] calculated from a pressure-volume curve [APD:glossary/40124], which is the leaf water potential [EnvThes:22311] at which the mean turgor pressure [TO:0000701] of cells [CL:0000000] in the leaf/terminal shoot drops to zero. This is the point at which the plant wilts.;A parameter calculated from a pressure-volume curve which is the water potential at which the mean cell turgor pressure in the leaf/shoot declines to zero.
  • type: numeric
  • units: MPa
  • allowable range: -5 - 0 MPa

leaf_NP_ratio

  • label: Leaf nitrogen to phosphorus ratio (N/P) per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to the mass of phosphorus [CHEBI:28659] in the leaf.;The ratio of the mass of nitrogen in a leaf to the mass of phosphorus in the leaf.
  • type: numeric
  • units: g/g
  • allowable range: 1 - 100 g/g

leaf_N_per_area

  • label: Leaf nitrogen (N) content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of nitrogen in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.15 - 50 g/m2

leaf_P_per_area

  • label: Leaf phosphorus (P) content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of phosphorus in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.005 - 1.2 g/m2

leaf_P_per_dry_mass

  • label: Leaf phosphorus (P) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of phosphorus in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.03 - 12 mg/g

leaf_dark_respiration_per_area

  • label: Leaf respiration rate per unit leaf area, in the dark (Rdark)
  • description: The cellular respiration [GO:0045333] rate [PATO:0000161] of plant leaves [PO:0025034] in the dark (absence of any photosynthetically active radiation [ENVO:01001864]), recorded per unit leaf area [PATO:0001323].;The respiration rate of a leaf in the dark (absence of photosynthetically active radiation), calculated as the production of CO2 per unit leaf area.
  • type: numeric
  • units: nmol{CO2}/m2/s
  • allowable range: 1 - 15000 nmol{CO2}/m2/s

leaf_dark_respiration_per_dry_mass

  • label: Leaf respiration rate per unit leaf dry mass, in the dark (Rdark)
  • description: The cellular respiration [GO:0045333] rate [PATO:0000161] of plant leaves [PO:0025034] in the dark (absence of any photosynthetically active radiation [ENVO:01001864]), recorded per unit leaf dry mass [TO:0001014].;The respiration rate of a leaf in the dark (absence of photosynthetically active radiation), calculated as the production of CO2 per unit leaf dry mass.
  • type: numeric
  • units: nmol{CO2}/g/s
  • allowable range: 0.1 - 1000 nmol{CO2}/g/s

leaf_intercellular_CO2_concentration_at_Amax

  • label: Leaf internal CO2 concentration during Amax measurement (ci)
  • description: The intercellular space [PO:0025379] carbon dioxide [CHEBI:16526] concentration [PATO:0000033] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451] concentrations.;The CO2 concentration in intercellular (interstitial) spaces in a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations; ci.
  • type: numeric
  • units: umol{CO2}/mol
  • allowable range: 10 - 1800 umol{CO2}/mol

leaf_intercellular_CO2_concentration_at_Asat

  • label: Internal CO2 concentration during Asat measurement (ci)
  • description: The intercellular space [PO:0025379] carbon dioxide [CHEBI:16526] concentration [PATO:0000033] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations.;The CO2 concentration in intercellular (interstitial) spaces in a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations; ci.
  • type: numeric
  • units: umol{CO2}/mol
  • allowable range: 10 - 1300 umol{CO2}/mol

leaf_intercellular_CO2_concentration_to_atmospheric_CO2_concentration_ratio

  • label: Ratio of internal to external CO2 concentrations (ci/ca)
  • description: The ratio [PATO:0001470] of leaf [PO:0025034] intercellular space [PO:0025379] carbon dioxide [CHEBI:16526] concentration [PATO:0000033] to atmospheric carbon dioxide [ENVO:01000451] concentration.;The ratio of the CO2 concentration in intercellular (interstitial) spaces in leaves to the atmospheric CO2 concentration, an indication of stomatal conductance; ci/ca.
  • type: numeric
  • units: umol{CO2_internal}/umol{CO2_ambient}
  • allowable range: 0.001 - 1.3 umol{CO2_internal}/umol{CO2_ambient}

leaf_photosynthetic_nitrogen_use_efficiency_saturated

  • label: Leaf photosynthetic nitrogen use efficiency during Asat measurement (PNUE)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its nitrogen [CHEBI:25555] content, with the photosynthesis measurements made on leaves which are saturated [NCIT:C61427] by photosynthetically active radiation [ENVO:01001864] and which are exposed to ambient [NCIT:C62306] atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033].;The ratio of photosynthesis (CO2 assimilation rate) to leaf nitrogen content, where photosynthesis has been measured in a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations; PNUE.
  • type: numeric
  • units: umol{CO2}/g{N}/s
  • allowable range: 0.1 - 1000 umol{CO2}/g{N}/s

leaf_photosynthetic_phosphorus_use_efficiency_saturated

  • label: Leaf photosynthetic phosphorus use efficiency during Asat measurement (PPUE)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its phosphorus [CHEBI:28659] content, with the photosynthesis measurements made on leaves which are saturated [NCIT:C61427] by photosynthetically active radiation [ENVO:01001864] and which are exposed to ambient [NCIT:C62306] atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033].;The ratio of photosynthesis (CO2 assimilation rate) to leaf phosphorus content, where photosynthesis has been measured in a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations; PPUE.
  • type: numeric
  • units: umol{CO2}/g{P}/s
  • allowable range: 1 - 10000 umol{CO2}/g{P}/s

leaf_photosynthetic_rate_per_area_maximum

  • label: Leaf photosynthesis rate per unit leaf area under saturating light and CO2 (Amax)
  • description: The photosynthetic rate [TO:0001015] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451; CHEBI:16526] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323].;The photosynthetic rate of a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations, at optimal leaf temperature and VPD conditions, per unit leaf area; maximum photosynthetic rate.
  • type: numeric
  • units: umol{CO2}/m2/s
  • allowable range: 2 - 65 umol{CO2}/m2/s

leaf_photosynthetic_rate_per_area_saturated

  • label: Leaf photosynthesis rate per unit leaf area under saturating light and ambient CO2 (Asat)
  • description: The photosynthetic rate [TO:0001015] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and ambient [NCIT:C62306] atmospheric carbon dioxide [ENVO:01000451; CHEBI:16526] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323].;The photosynthetic rate of a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, air temperature and vapour pressure deficit, per unit leaf area; light-saturated photosynthetic rate.
  • type: numeric
  • units: umol{CO2}/m2/s
  • allowable range: 0.01 - 50 umol{CO2}/m2/s

leaf_photosynthetic_rate_per_dry_mass_maximum

  • label: Leaf photosynthesis rate per unit leaf dry mass under saturating light and CO2 (Amax)
  • description: The photosynthetic rate [TO:0001015] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451; CHEBI:16526] concentrations [PATO:0000033], recorded per unit leaf dry mass [TO:0001014].;The photosynthetic rate of a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations, at optimal leaf temperature and VPD conditions, per unit leaf dry mass; maximum photosynthetic rate.
  • type: numeric
  • units: umol{CO2}/g/s
  • allowable range: 0.01 - 1 umol{CO2}/g/s

leaf_photosynthetic_rate_per_dry_mass_saturated

  • label: Leaf photosynthesis rate per unit leaf dry mass under saturating light and ambient CO2 (Asat)
  • description: The photosynthetic rate [TO:0001015] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and ambient [NCIT:C62306] atmospheric carbon dioxide [ENVO:01000451; CHEBI:16526] concentrations [PATO:0000033], recorded per unit leaf dry mass [TO:0001014].;The photosynthetic rate of a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, air temperature and vapour pressure deficit, per unit leaf dry mass; saturated photosynthetic rate.
  • type: numeric
  • units: umol{CO2}/g/s
  • allowable range: 0.001 - 0.8 umol{CO2}/g/s

leaf_transpiration_per_area_at_Asat

  • label: Leaf transpiration per unit leaf area during Asat measurement (E)
  • description: The transpiration [ENVO:02500035] rate [PATO:0000161] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Transpiration rate is the rate at which plants lose water [CHEBI:15377] vapor [NCIT:C63390] through their stomata [PO:0008032].;The rate of water loss from a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 50 mmol{H2O}/m2/s

leaf_water_use_efficiency_instantaneous

  • label: Instantaneous water use efficiency (WUE)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its transpiration [ENVO:02500035] rate [PATO:0000161], with photosynthetic rate and transpiration rate determined by gas exchange measurements.;The ratio of photosynthetic rate (A; CO2 assimilation rate) to transpiration rate (E; water loss), with photosynthetic rate and transpiration rate determined from gas exchange measurements, inside a leaf chamber. This is usually called instantaneous water use efficiency, but can also be referred to as photosynthetic water use efficiency (PWUE) or transpiration efficiency.
  • type: numeric
  • units: umol{CO2}/mmol{H2O}
  • allowable range: 0.1 - 200 umol{CO2}/mmol{H2O}

leaf_stomatal_conductance_per_area_at_Amax

  • label: Leaf stomatal conductance to water vapour per unit leaf area during Amax measurement (gsw)
  • description: The stomatal conductance [EnvThes:21211] to water vapour of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Stomatal conductance is the rate [PATO:0000161] of gaseous [MESH:D005740] water [CHEBI:15377] diffusion [NCIT:C82333] through stomata [PO:0008032].;The rate of gaseous water diffusion through stomata in a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 2000 mmol{H2O}/m2/s

leaf_stomatal_conductance_per_area_at_Asat

  • label: Leaf stomatal conductance to water vapour per unit leaf area during Asat measurement (gsw)
  • description: The stomatal conductance [EnvThes:21211] to water vapour of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Stomatal conductance is the rate [PATO:0000161] of gaseous [MESH:D005740] water [CHEBI:15377] diffusion [NCIT:C82333] through stomata [PO:0008032].;The rate of gaseous water diffusion through stomata in a leaf exposed to saturating light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 2000 mmol{H2O}/m2/s

leaf_transpiration_per_area_at_Amax

  • label: Leaf transpiration per unit leaf area during Amax measurement (E)
  • description: The transpiration [ENVO:02500035] rate [PATO:0000161] of plant leaves [PO:0025034] whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Transpiration rate is the rate at which plants lose water [CHEBI:15377] vapor [NCIT:C63390] through their stomata [PO:0008032].;The rate of water loss from a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 50 mmol{H2O}/m2/s

leaf_dark_transpiration_per_area

  • label: Leaf transpiration rate per unit leaf area, in the dark
  • description: The transpiration [ENVO:02500035] rate [PATO:0000161] of plant leaves [PO:0025034] in the dark (absence of any photosynthetically active radiation [ENVO:01001864]) and exposed to ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Transpiration rate is the rate at which plants lose water [CHEBI:15377] vapor [NCIT:C63390] through their stomata [PO:0008032].;The rate of water loss from a leaf in the dark (no photosynthetically active radiation) and ambient carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 1e-04 - 10 mmol{H2O}/m2/s

leaf_dry_mass

  • label: Leaf dry mass
  • description: A leaf mass trait [TO:0000505] which is the dry [PATO:0001824] mass [PATO:0000125] of a leaf [PO:0025034] or leaf analogue (a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335]).;The oven dry mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
  • type: numeric
  • units: mg
  • allowable range: 0.01 - 15000 mg

recruitment_time

  • label: Seedling recruitment time, by month
  • description: The months [UO:0000035] during which seed [PO:0009010] germination [GO:0009845] and seedling establishment [EnvThes:20410] occurs.;The months during which seed germination and seedling establishment occurs.
  • type: categorical
  • allowable values:
    • **:

leaf_lamina_division

  • label: Leaf divisions
  • description: A leaf shape trait [TO:0000492] which indicates types [NCIT:C25284] of lobed leaves [PO:0025517] and compound leaves [PO:0020043], including the arrangement [NCIT:C61532] of the lobes or leaflets [PO:0020049] with respect to the midrib [PO:0020139] or other primary leaf veins [PO:0025413] and the number of orders of axes [NCIT:C25154] across which there are incisions or divisions.;A trait that captures the types of lobed leaves and compound leaves, including the arrangement of the lobes or leaflets relative to the midrib or other primary veins and the number of orders of axes across which there are incisions or divisions.
  • type: categorical
  • allowable values:
    • bipartite: Leaf divided into two lobes.
    • bipinnate: Leaf that is twice pinnate, having pinnate leaflets that are themselves pinnately divided.
    • bipinnatifid: Leaf that is twice pinnatifid, with two series of lobes that extend less than half-way to the midrib.
    • bipinnatipartite: Leaf that is twice pinnatipartite, with two series of lobes that extend more than half-way to the midrib.
    • bipinnatisect: Leaf that is twice pinnatisect, with two series of lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets.
    • dichotomously_lobed: Leaf divided repeatedly into two lobes.
    • palmately_compound: Leaf divided into leaflets that attach to a central point. (Synonyms, digitate)
    • palmately_lobed: Leaf with divisions that radiate from a single point, usually the basal junction of major veins.
    • pinnately_compound: Leaf divided into leaflets that attach along a central axis. [PATO_0000410]
    • pinnately_lobed: Leaf with divisions that are symmetrical across a central axis, usually the midvein.
    • pinnatifid: Leaf with lobes that extend less than half-way to the midrib.
    • pinnatipartite: Leaf with lobes that extend more than half-way to the midrib.
    • pinnatisect: Leaf with lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets.
    • trichotomously_lobed: Leaf divided repeatedly into three lobes.
    • trifoliate: Leaf divided into three leaflets. (Synonyms, ternate, trifoliolate)
    • tripartite: Leaf divided into three lobes.
    • tripinnate: Leaf that is three times pinnate, with pinnate leaflets that are twice more pinnately divided.

leaf_type

  • label: Leaf type
  • description: A leaf shape trait [TO:0000492] which is inclusive [NCIT:C166400] categories describing the shape [PATO:0000052] of a leaf [PO:0025034] as broad, needle-like, or scale-like.;Variable distinguishing between general leaf form (shape) categories.
  • type: categorical
  • allowable values:
    • broadleaf: Flat leaf lamina, regardless of whether the leaves are or are not toothed, lobed, or compound.
    • leafless: An adult stage plant that lacks leaves throughout the year.
    • needle: Needle or awl-shaped leaf that is generally 3-dimensional.
    • scale: Leaf reduced to a small scale.

huber_value

  • label: Huber value
  • description: A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of sapwood [PO:0004513] area [PATO:0001323] to the total distal [PATO:0001234] leaf [PO:0025034] area.;For a single branch, the ratio of sapwood cross-sectional area at the base of the branch to the branch’s total leaf area.
  • type: numeric
  • units: mm2{sapwood}/mm2{leaf}
  • allowable range: 1e-06 - 0.2 mm2{sapwood}/mm2{leaf}

leaf_fluorescence_quantum_yield

  • label: Leaf ambient quantum yield
  • description: Ambient quantum yield [TO:0012004] of photosystem II [GO:0009539], as determined through chlorophyll fluorescence [GO:0090546]. Quantum yield is the ratio [PATO:0001470] of photons [NCIT:C88112] absorbed to photons emitted [SIO:000384] through fluorescence.;A trait that indicates the health of photosystem II in a light adapted leaf. Quantum yield is calculated as the ratio of (maximum fluorescence under ambient conditions)-(fluorescence under ambient conditions)/(maximum fluorescence under ambient conditions).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.01 - 1 {dimensionless}

leaf_specific_hydraulic_conductivity

  • label: Leaf specific hydraulic conductivity (Kl)
  • description: A stem [PO:0009047] water transport [GO:0006833] trait which is a measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a stem under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), normalised to the branches’ leaf [PO:0025034] area [PATO:0001323] and accounting for the path (stem) length [PATO:0000122]. It is calculated as the ratio of the water flow rate (kg/s) to the water potential gradient across the stem segment (MPa), divided by the total leaf area (1/m2), then multiplied by the stem length (m).;Kl; Measure of the efficiency of water flow through a stem segment, normalised to the total leaf area distil to the stem segment. It is calculated as the ratio of the water flow rate (kg/s) to the water potential gradient across the stem segment (MPa), divided by the total leaf area (1/m2), then multiplied by the stem length (m).
  • type: numeric
  • units: 10^4.kg/m/MPa/s
  • allowable range: 0.1 - 500 10^4.kg/m/MPa/s

sapwood_specific_hydraulic_conductivity

  • label: Sapwood specific hydraulic conductivity (Ks)
  • description: A stem [PO:0009047] water transport [GO:0006833] trait which is a measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a stem under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), normalised to the diameter [PATO:0001334] of water transport tissues [CARO:0000043] and accounting for the stem length [PATO:0000122]. It is calculated as the ratio [PATO:0001470] of the water flow [NCIT:C41141] rate through the stem segment (kg/s) to the water potential gradient across the stem segment (MPa), divided by the sapwood [PO:0004513] cross-sectional [NCIT:C63795] area [PATO:0001323] (m2), then multiplied by the path length measured (m).;Ks; Measure of the efficiency of water flow through a stem segment normalised to the sapwood cross-sectional area across which the flow is measured. It is calculated as the ratio of the water flow rate (kg/s) to the water potential gradient across the stem segment (MPa), divided by the sapwood cross sectional area (1/m2), then multiplied by the stem length (m).
  • type: numeric
  • units: kg/m/MPa/s
  • allowable range: 0.01 - 500 kg/m/MPa/s

leaf_CN_ratio

  • label: Leaf carbon to nitrogen ratio (C/N)
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to the mass of nitrogen [CHEBI:25555] in the leaf.;The ratio of the mass of carbon in a leaf to the mass of nitrogen in the leaf.
  • type: numeric
  • units: g/g
  • allowable range: 10 - 1000 g/g

leaf_C_per_dry_mass

  • label: Leaf carbon (C) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of carbon in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 50 - 750 mg/g

leaf_delta15N

  • label: Leaf stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of leaves [PO:0025034].;Nitrogen stable isotope ratio, d15N, of leaves.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

wood_C_per_dry_mass

  • label: Wood carbon (C) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of carbon in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 50 - 750 mg/g

wood_N_per_dry_mass

  • label: Wood nitrogen (N) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of nitrogen in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

wood_delta13C

  • label: Wood stable carbon isotope composition (delta13C)
  • description: Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of wood [NCIT:C93062].;Carbon stable isotope ratio, d13C, of wood.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -50 - 0 {delta}[ppth]

wood_delta15N

  • label: Wood stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of wood [NCIT:C93062].;Nitrogen stable isotope ratio, d15N, of wood.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

leaf_fresh_mass

  • label: Leaf fresh mass
  • description: A leaf mass trait [TO:0000505] which is the mass [PATO:0000125] of a fresh [EnvThes:21976] leaf [PO:0025034], indicating its mass at the time it is picked. For some species the mass of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335].;The fresh mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
  • type: numeric
  • units: mg
  • allowable range: 0.1 - 30000 mg

stem_water_delta18O

  • label: Stem water stable oxygen isotope composition (delta18O)
  • description: Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of water [CHEBI:15377] in stems [PO:0009047].;Oxygen stable isotope ratio, d18O, of water in stems.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -10 - 50 {delta}[ppth]

seedbank_location

  • label: Seed storage location
  • description: The location [NCIT:C25341] of a plant’s [PO:0000003] seedbank [MESH:D000068098], indicating locations where mature [PATO:0001701] seeds [PO:0009010] are stored and remain viable [PATO:0000169] until germination [GO:0009845].;Location of a plant’s seedbank, the location where mature seeds are stored and viable until germinating.
  • type: categorical
  • allowable values:
    • canopy_seedbank: Seeds stored in the canopy, usually still within a woody cone.
    • canopy_seedbank_absent: Seed explicitly not stored in the canopy.
    • none: Seeds are not stored.
    • seedbank: A seedbank exists, but its locations is not specified.
    • soil_seedbank: Seeds stored in the soil.
    • soil_seedbank_absent: Seed explicitly not stored in the soil.

fire_time_from_fire_to_50_percent_flowering

  • label: Time from fire until 50% of individuals are flowering
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until a 50% of the individuals [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] reaches whole plant flowering stage [PO_0007016].;The time interval from a fire until 50% of the individuals in a population produce flowers.
  • type: numeric
  • units: mo
  • allowable range: 1 - 500 mo

reproductive_maturity

  • label: Age of reproductive maturity
  • description: The age [NCIT:C25150] of a plant when it is capable of sexual reproduction [GO:0000003], which is the time interval (duration) [PATO:0001309] between seed germination [GO:0009845] and when a plant is able to produce mature [PATO:0001701] seed [PO:0009010].;The age of a plant when they are able to set seed, indicating it is reproductively mature.
  • type: numeric
  • units: a
  • allowable range: 0.1 - 1000 a

fire_time_from_fire_to_flowering

  • label: Time from fire to first flowering
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until one individual [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] reaches whole plant flowering stage [PO_0007016].;The time interval from a fire until the first individual in a population begins flowering.
  • type: numeric
  • units: mo
  • allowable range: 1 - 1000 mo

fire_time_from_fire_to_flowering_decline

  • label: Time from fire until flowering declines
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until the individuals [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] begin to decrease their level of flowering [PO_0007016].;The time interval from a fire until the degree of flowering by individuals in a population begin to decline.
  • type: numeric
  • units: mo
  • allowable range: 1 - 1000 mo

fire_time_from_fire_to_peak_flowering

  • label: Time from fire to peak flowering
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until the individuals [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] reach maximum flowering [PO_0007016] levels.;The time interval from a fire until the individuals in a population display maximum flowering.
  • type: numeric
  • units: mo
  • allowable range: 1 - 1000 mo

dispersal_unit

  • label: Dispersal unit
  • description: A reproductive shoot system morphology trait [TO:0000786] which is the plant structure [PO:0009011] involved in dispersal [EnvThes:21038], broadly distinguishing between taxa [NCIT:C40098] whose main dispersal units are spores [NCIT:C79672], seeds [PO:0009010] or fruit [PO:0009001].;Broad terms indicating the plant structure involved in dispersal, distinguishing between taxonomic groups whose main dispersal units are spores, seeds, or fruit.
  • type: categorical
  • allowable values:
    • fruit: Dispersal unit is a fruit.
    • seed: Dispersal unit is a seed.
    • seedling: Dispersal unit is a seedling. (Synonyms, viviparous)
    • spore: Dispersal unit is a spore.

leaf_water_content_per_dry_mass

  • label: Leaf water content per unit leaf dry mass
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of water in a fresh leaf (leaf water content) to leaf dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 20 g/g

dispersers

  • label: Diaspore dispersal agents
  • description: The taxa [NCIT:C40098] (biotic vectors) and abiotic mechanisms [NCIT:C25343] that disperse [OMIT: 0027297] seeds [PO:0009010] and fruit [PO:0009001] away from a parent plant.;Dominant diaspore dispersal agents, including both biotic agents (specific taxa) or abiotic mechanisms responsible for dispersing seeds (or other dispersal units) away from the parent plant.
  • type: categorical
  • allowable values:
    • abiotic: Fruits are dispersed by unspecified abiotic mechanisms.
    • animals: Animals are an important fruit dispersal agent.
    • ants: Ants are an important fruit dispersal agent.
    • bats: Bats are an important fruit dispersal agent.
    • birds: Birds are an important fruit dispersal agent.
    • cassowaries: Cassowaries, in contrast to smaller flying birds, are an important fruit dispersal agent.
    • fish: Fish are an important fruit dispersal agent.
    • floods: Floods are an important fruit dispersal agent.
    • flying_foxes: Flying foxes are an important fruit dispersal agent.
    • flying_vertebrates: Flying birds and bats are important fruit dispersal agents.
    • garden_refuse: Fruit are dispersed through garden refuse.
    • indigenous_people: Indigenous people disperse the fruit through traditional practices.
    • insects: Insects are an important fruit dispersal agent.
    • invertebrates: Invertebrates are an important fruit dispersal agent.
    • mammals: Mammals are an important fruit dispersal agent.
    • mammals_domestic: Domesticated ammals are an important fruit dispersal agent.
    • non-flying_vertebrates: Mammals and non-flying birds (i.e. cassowary, emu) are important fruit dispersal agents.
    • passive: There is no special dispersal agent.
    • rain: Rain is an important fruit dispersal agent; the diaspore is propelled by action of rain on plant structure or the wetting of the plant structure by rain or dew.
    • rodents: Rodents are an important fruit dispersal agent.
    • vehicles: Vehicles are an important dispersal agent, such as through mud on tyres.
    • vertebrates: Vertebrates are an important fruit dispersal agent.
    • water: Water is an important fruit dispersal agent.
    • water_currents: Water currents are an important fruit dispersal agent; the diaspore may be either floating or submerged in fresh/saltwater currents.
    • wind: Wind is an important fruit dispersal agent.

stem_specific_hydraulic_conductivity

  • label: Stem specific hydraulic conductivity (Ks)
  • description: A stem [PO:0009047] water transport [GO:0006833] trait which is a measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a stem under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), normalised to the diameter [PATO:0001334] of water transport tissues [CARO:0000043] and accounting for the stem length [PATO:0000122]. It is calculated as the ratio [PATO:0001470] of the water flow [NCIT:C41141] rate through the stem segment (kg/s) to the water potential gradient across the stem segment (MPa), divided by the stem cross-sectional [NCIT:C63795] area [PATO:0001323] (m2), then multiplied by the path length measured (m).;Ks; Measure of the efficiency of water flow through a stem segment normalised to the stem (sapwood + bark) cross-sectional area across which the flow is measured. It is calculated as the ratio of the water flow rate (kg/s) to the water potential gradient across the stem segment (MPa), divided by the stem cross sectional area (1/m2), then multiplied by the stem length (m).
  • type: numeric
  • units: kg/m/MPa/s
  • allowable range: 0.01 - 500 kg/m/MPa/s

water_potential_50percent_lost_conductivity

  • label: Stem xylem pressure, 50% lost conductivity (P50)
  • description: The tension [PATO:0002284] (water pressure) (water potential [EnvThes:22311]) in stem [PO:0009047] xylem vessels [PO:0025417] at which the hydraulic conductivity [EnvThes:22133] has declined by 50% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;A measure of stem hydraulic vulnerability, which is the stem xylem water pressure value at which stem hydraulic conductivity has declined by 50% from the mean maximum rate of hydraulic conductivity.
  • type: numeric
  • units: MPa
  • allowable range: -20 - 0 MPa

osmotic_potential_at_full_turgor

  • label: Osmotic potential at full tugor
  • description: A parameter [STATO:0000034] calculated from a pressure-volume curve [APD:glossary/40124], which is, for a shoot or leaf [PO:0025034], the osmotic potential (solute potential) when they are fully hydrated (relative water content = 100%). Osmotic potential is always negative and is the potential energy [ENVO:2000016] of water [CHEBI:15377] molecules to flow [NCIT:C41141] across a semi-permeable [PATO:0000982] membrane [GO:0016020] due to the differences in solute [SIO:010418] concentration [PATO:0000033] across the membrane.;A parameter calculated from a pressure-volume curve which is the osmotic potential (solute potential) in leaf cells at full turgor (when relative water content = 100%). Osmotic potential is the energy of water molecules to move across a semi-permeable membrane due to differences in solute concentration across a membrane and is least negative (largest) when plants are at full turgor.
  • type: numeric
  • units: MPa
  • allowable range: -10 - -0.1 MPa

water_potential_predawn

  • label: Pre-dawn water potential
  • description: The shoot system [PO:0009006] water potential [EnvThes:22311] just before sunrise, a proxy for soil water potential [EnvThes:22283] and representing the plant’s daily maximum [NCIT:C25564] (least negative) water potential.;The water potential of a plant’s shoot system just before sunrise, a proxy for soil water potential and the plant’s highest (least negative) water potential for the day.
  • type: numeric
  • units: MPa
  • allowable range: -20 - 0 MPa

plant_flood_regime_classification

  • label: Plant flood regime response
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which categorises plants based on the depth [PATO:0001595], duration [PATO:0001309] and frequency [PATO:0000044] of inundation [SWEET:phenSystem/Inundation] a plant [PO:0000003] is able to tolerate. It incorporates their ability to tolerate or respond to fluctuations in flooding [ENVO:01000710] and drying, including growth [GO:0040007] and seed germination [GO:0009845] responses.;Classification scheme of plant flood regime requirements and tolerances, categorising taxa into seven groups based on their germination and growth responses to flood regimes.
  • type: categorical
  • allowable values:
    • amphibious: Species which germinate in damp or flooded conditions and can grow in or our of water.
    • amphibious_fluctuation_responders_floating: Species which germinate in flooded conditions, grow in both flooded and damp conditions, reproduce above the surface of the water, and which have floating leaves when inundated. (Species with floating leaves; ARf)
    • amphibious_fluctuation_responders_plastic: Species which germinate in flooded conditions, grow in both flooded and damp conditions, reproduce above the surface of the water, and which have morphological plasticity (e.g. heterophylly) in response to water-level variation. (Morphologically plastic species; ARp)
    • amphibious_fluctuation_tolerators_emergent: Species which germinate in damp or flooded conditions, which tolerate variation in water level, and which grow with their basal portions under water and reproduce out of water. (Emergent species; ATe)
    • amphibious_fluctuation_tolerators_low: Species which germinate in damp or flooded conditions, which tolerate variation in water level, which are low-growing and tolerate complete submersion when water-levels rise. (Low-growing species; ATl)
    • submerged: Species which germinate, grow and reproduce underwater. (S
    • terrestrial_damp: Species which germinate, grow and reproduce on saturated soil. (Damp species; Tda)
    • terrestrial_dry: Species which germinate, grow and reproduce where there is no surface water and the water table is below the soil surface. (Dry species; Tdr)

vegetative_reproduction_ability

  • label: Vegetative reproduction ability
  • description: A whole plant [PO:0000003] life history [ECOCORE:00000018] trait which is the ability [NCIT:C78209] of individual [EFO:0000542] plants to reproduce [GO:0000003] through asexual reproduction [GO:0019954] (vegetative reproduction).;A binary trait, indicating whether a plant taxon is able to reproduce and spread through the growth and division of vegetative material.
  • type: categorical
  • allowable values:
    • not_vegetative: Plant taxa that regenerate and spread only by seed.
    • vegetative: Plant taxa that can regenerate and spread through the growth and division of vegetative material, not just from seed; examples include runners, rhizomes, and bulbs.
    • vegetative_obligate: Plant taxa that never (or very rarely) regenerate from seed, instead regenerating and spreading entirely through the growth and division of vegetative material.

stem_dark_respiration_per_area

  • label: Stem respiration rate per unit stem area, in the dark
  • description: The cellular respiration [GO:0045333] rate [PATO:0000161] of plant stems [PO:0009047] in the dark (absence of any photosynthetically active radiation [ENVO:01001864]), recorded per unit stem area [PATO:0001323].;The respiration rate of a stem in the dark (absence of photosynthetically active radiation), calculated as the production of CO2 per unit stem area.
  • type: numeric
  • units: umol{CO2}/m2/s
  • allowable range: 0.01 - 10 umol{CO2}/m2/s

bark_thickness

  • label: Bark thickness
  • description: A bark morphology trait which is the thickness [PATO:0000915] of the bark [PO:0004518]. The corresponding stem [PO:0009047] diameter [PATO:0001334] should also be recorded so that a scaled measure, the bark thickness to stem diameter ratio, can also be calculated.;Thickness of the bark on the stem.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 200 mm

leaf_photosynthesis_Jmax_per_area_25C

  • label: Leaf Jmax per unit leaf area at 25 deg C (Jmax25)
  • description: Maximum [NCIT:C25564] electron transport [MESH:D004579] rate [PATO:0000161] through the photosynthetic [GO:0015979] electron transport chain [GO:0022900], recorded per unit leaf [PO:0025034] area [PATO:0001323] and measured at 25 Celsius. The value is a model parameter [STATO:0000034] derived from the portion of the photosynthesis A-ci response curve at high carbon dioxide concentrations where ATP and NADPH production by the light reactions [GO:0019684] limits the rate of carboxylation [MOP:0000591].;The maximum potential photosynthetic electron transport rate, measured at 25 Celsius and recorded per unit leaf area. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at high carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the rate of ATP and NADPH production by the light reactions.
  • type: numeric
  • units: umol/m2/s
  • allowable range: 35 - 800 umol/m2/s

leaf_photosynthesis_Vcmax_per_area

  • label: Leaf Vcmax per unit leaf area (Vcmax)
  • description: Maximum [NCIT:C25564] carboxylation [MOP:0000591] rate [PATO:0000161] by the Rubisco complex [GO:0048492] in plant leaves [PO:0025034], recorded per unit leaf [PO:0025034] area [PATO:0001323]. The value is a model parameter [STATO:0000034] derived from the initial slope of a photosynthesis [GO:0015979] A-ci response curve, the component that is carbon dioxide [CHEBI:16526] limited and hence limited by Rubisco activity.;The apparent maximum rate of Rubisco (ribulose 1,5-bisphosphate carboxylase/oxygenase) activity carboxylase activity (carboxylation capacity), recorded per unit leaf dry mass. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at low carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the Rubisco activity.
  • type: numeric
  • units: umol/m2/s
  • allowable range: 10 - 300 umol/m2/s

lifespan

  • label: Lifespan
  • description: The duration [PATO:0001309] of a plant’s lifespan (longevity [NCIT:C153298]), from seed germination [GO:0009845] to death [GO:0016265].;Plant lifespan, the duration from germination to death. Lifespan refers to the death of both aboveground and belowground biomass, indicating the death of the genetic individual.
  • type: numeric
  • units: a
  • allowable range: 0.1 - 10000 a

resprouting_capacity_juvenile

  • label: Post-fire resprouting capacity of juvenile plants
  • description: A fire response trait which indicates whether juvenile [PATO:0001190] plants [PO:0000003] from a taxon [NCIT:C40098] or population [PCO:0000001] are killed or have vegetative meristematic tissue [PO:0006079] (buds [PO:0000055]) that resprouts [Cerrado:fire#PlantResprout] following a fire [ENVO:01000786] with 100% leaf scorch.;Trait indicating whether juvenile phase individuals of a taxon are killed, resprout, or have a mixed survival response when exposed to a fire with 100% leaf scorch.
  • type: categorical
  • allowable values:
    • juvenile_fire_killed: Few to no juvenile plants resprout following fire with 100% leaf scorch.
    • juvenile_moderate_fire_killed: 25-50% of juvenile plants resprout following fire with 100% leaf scorch.
    • juvenile_resprout: More than 50% of juvenile plants survive and resprout following fire with 100% leaf scorch.

seedbank_longevity_class

  • label: Seedbank longevity class
  • description: A seedbank’s [MESH:D000068098] longevity [EnvThes:22228], expressed as a categorical variable [STATO:0000252], where seedbank longevity is the time interval (duration [PATO:0001309]) a mature [PATO:0001701] seed [PO:0009010] remains in the seedbank, viable [PATO_0000169] and able to germinate [GO:0009845].;Categorical groupings indicating the length of time (duration) a mature seed remains in the seedbank, viable and able to germinate.
  • type: categorical
  • allowable values:
    • long_persistent: Seed that persists in the seedbank for more than 5 years, although the time interval varies by study.
    • persistent: Seed that survives in the seedbank until at least its second germination season.
    • short_persistent: Seed that persists in the seedbank for approximately 1 to 5 years, although the time interval varies by study.
    • transient: Seed that survives in the seedbank until its first annual germination season, but not until its second annual germination season. Some transient seeds will germinate within weeks to months, while others may remain in the seedbank for slightly over a year, depending on how the annual recruitment season aligns with seed maturation. Transient seeds are non-dormant, remaining in the seedbank until environmental conditions are appropriate for germination.
    • widely_dispersed: Seed longevity not important, as seeds are widely dispersed and always available for germination.

seedling_establishment_conditions

  • label: Seedling establishment conditions
  • description: The environmental conditions [ENVO:01000203] under which a taxon [NCIT:C40098] displays seed [PO:0009010] germination [GO:0009845] and seedling establishment [EnvThes:20410], with trait values reflecting successional changes to the vegetation community.;The environmental conditions under which a taxon’s seedlings can establish, with trait values reflecting successional changes to the vegetation community.
  • type: categorical
  • allowable values:
    • establish_anytime: Seedlings can establish immediately following a disturbance and in later years as vegetation ages, right thru to mature and over-mature vegetation
    • establish_intermediate_to_mature_vegetation: Seedlings require some environmental characteristics not found in vegetation straight after fires to establish, for they cannot establish within the first season or two after fire but can establish in older vegetation (including mature to over-mature vegetation).
    • establish_post_disturbance: Seedling establishment occurs following disturbances that increase light levels at the ground surface, but the type of disturbance is not documented.
    • establish_post_fire: Seedling establishment occurs following fire.

life_form

  • label: Life form
  • description: Raunkiaer classification of life form, which is a feature of the whole plant [PO:0000003] defining the position [PATO:0000140] of the perennating meristematic tissue [PO:0006079] (buds [PO:0000055] or shoot apices [PO:0000037]) on the shoot system [PO:0009006], their distance [PATO:0000040] to the ground surface [sweetontology.net/realmSoil/Ground] and the protection afforded to survive the unfavourable season, or the production of seeds [PO:0009010] to avoid the unfavourable season.;Raunkiaer classification of life form, indicating the location of and protection provided to surviving meristematic tissue (buds) during the unfavourable (cold or dry) season.
  • type: categorical
  • allowable values:
    • aerophyte: Plant that obtains moisture and nutrients from the air and rain, resting on other plants or the ground surface.
    • chamaephyte: Plant whose perennating buds are on persistent shoots that do not extend more than 50 cm above the soil surface. These are normally woody perennials.
    • cryptophyte: Plant whose perennating buds are subterranean or under water. This grouping can be further divided into geophyte, helophyte, and hydrophyte.
    • epiphyte: Plant that grows on the surface of another plant.
    • geophyte: Plant that perennates as an organ (rhizome, bulb, corm, or tuber) buried in dry soil, with all leaves dying back each year; a subcategory of cryptophyte.
    • helophyte: Plant that perennates as an organ (rhizome, bulb, corm, or tuber) buried in marshy or wet soil or lying under water, with buds emerging during the growing season and shoots and leaves rising about the water surface; a subcategory of cryptophyte. (Synonyms, emergent_hydrophyte)
    • hemicryptophyte: Plant whose perennating buds are at or near the soil surface.
    • hydrophyte: Plant that perennates as buds or as an organ (rhizome, bulb, corm, or tuber) buried in marshy or wet soil or lying under water, with shoots emerging during the growing season that remain below or at the water surface; a subcategory of cryptophyte.
    • phanerophyte: Plant whose perennating buds are on persistent shoots which rise at least 50 cm above the soil surface. These are normally woody perennials.
    • therophyte: Annual plants whose shoot and root systems die completely at the end of flowering and seed production, such that they survive the unfavourable cold or dry season in the form of seeds.

leaf_photosynthesis_Jmax_per_area

  • label: Leaf Jmax per unit leaf area (Jmax)
  • description: Maximum [NCIT:C25564] electron transport [MESH:D004579] rate [PATO:0000161] through the photosynthetic [GO:0015979] electron transport chain [GO:0022900], recorded per unit leaf [PO:0025034] area [PATO:0001323]. The value is a model parameter [STATO:0000034] derived from the portion of the photosynthesis A-ci response curve at high carbon dioxide concentrations where ATP and NADPH production by the light reactions [GO:0019684] limits the rate of carboxylation [MOP:0000591].;The maximum potential photosynthetic electron transport rate, recorded per unit leaf area. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at high carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the rate of ATP and NADPH production by the light reactions.
  • type: numeric
  • units: umol/m2/s
  • allowable range: 10 - 800 umol/m2/s

diaspore_fleshiness

  • label: Diaspore fleshiness
  • description: A seed morphology trait [TO:0002629] which indicates whether a mature [PATO:0001701] fresh [EnvThes:21976] diaspore is fleshy [PATO:0002351], where a diaspore is a plant’s dispersal [EnvThes:21038] unit and includes both the seed [PO:0009010] and the seed dispersal [EnvThes:21042] structures [PATO:0000141].;Binary variable, dividing diaspores (dispersal units) into ‘dry’ versus ‘fleshy’ based on whether they have fleshy pericarps, fleshy accessory tissues (such as the receptacle in Podocarpus), or fleshy appendages (e.g. aril, thalamus, receptacle, calyx, rachis or bract or succulent pedicel) when mature. Dispersal units where none of these tissues is fleshy are designated as ‘non-fleshy’.
  • type: categorical
  • allowable values:
    • dry: Diaspores where the true fruit, accessory tissues and dispersal appendages are all dry when mature.
    • fleshy: Diaspores where either the true fruit, accessory tissues or dispersal appendages are fleshy when mature.

fruit_fleshiness

  • label: Fruit fleshiness
  • description: A fruit morphology trait [TO:0002629] which indicates whether a mature [PATO:0001701] fresh [EnvThes:21976] fruit [PO:0009001] is fleshy [PATO:0002351].;Binary variable, dividing entire, true fruits into ‘dry’ versus ‘fleshy’ based on whether their botanical fruit type is fleshy when mature and fresh.
  • type: categorical
  • allowable values:
    • dry: Fruits are dry when mature and fresh.
    • fleshy: Fruits are fleshy when mature and fresh.

leaf_photosynthetic_rate_per_area_ambient

  • label: Leaf photosynthesis rate per unit leaf area under ambient light and CO2 (A)
  • description: The photosynthetic rate [TO:0001015] of plant leaves [PO:0025034] exposed to ambient [NCIT:C62306] photosynthetically active radiation [ENVO:01001864] levels and ambient atmospheric carbon dioxide [ENVO:01000451; CHEBI:16526] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323].;The photosynthetic rate of a leaf exposed to ambient light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, air temperature and vapour pressure deficit, per unit leaf area.
  • type: numeric
  • units: umol{CO2}/m2/s
  • allowable range: 0.01 - 40 umol{CO2}/m2/s

leaf_stomatal_conductance_per_area_ambient

  • label: Leaf stomatal conductance to water vapour per unit leaf area under ambient conditions (gsw)
  • description: The stomatal conductance [EnvThes:21211] to water vapour of plant leaves [PO:0025034] exposed to ambient [NCIT:C62306] photosynthetically active radiation [ENVO:01001864] levels and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Stomatal conductance is the rate [PATO:0000161] of gaseous [MESH:D005740] water [CHEBI:15377] diffusion [NCIT:C82333] through stomata [PO:0008032].;The rate of gaseous water diffusion through stomata in a leaf exposed to ambient light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 2000 mmol{H2O}/m2/s

leaf_transpiration_per_area_ambient

  • label: Leaf transpiration per unit leaf area under ambient conditions (E)
  • description: The transpiration [ENVO:02500035] rate [PATO:0000161] of plant leaves [PO:0025034] exposed to ambient [NCIT:C62306] photosynthetically active radiation [ENVO:01001864] levels and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Transpiration rate is the rate at which plants lose water [CHEBI:15377] vapor [NCIT:C63390] through their stomata [PO:0008032].;The rate of water loss from a leaf exposed to ambient light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations, recorded per unit leaf area.
  • type: numeric
  • units: mmol{H2O}/m2/s
  • allowable range: 0.1 - 50 mmol{H2O}/m2/s

water_potential_midday

  • label: Midday water potential
  • description: The average water potential [EnvThes:22311] of a plant’s leaf [PO:0025034] and shoot [PO:0009006] mesophyll [PO:0006070] tissue [CARO:0000043] during maximum midday transpiration [ENVO:02500035] and assumed to be the plant’s daily minimum [NCIT:C25570] water potential.;The water potential of a plant’s leaf or shoot photosynthetic tissue during maximum midday transpiration, the time of day when the plant is most water stressed and has the lowest (most negative) water potential.
  • type: numeric
  • units: MPa
  • allowable range: -20 - 0 MPa

stem_hydraulic_conductivity

  • label: Stem hydraulic conductivity (Kh)
  • description: A stem [PO:0009047] water transport [GO:0006833] trait which is a measure of the rate [PATO:0000161] of water [CHEBI:15377] movement [NCIT:C37994] through a stem under a specific [NCIT:C38024] driving force [PATO:0001035] (water potential [EnvThes:22311] gradient [NCIT:C70693]), accounting for the stem length [PATO:0000122]. It is calculated as the ratio [PATO:0001470] of the water flow [NCIT:C41141] rate through the stem segment (kg/s) to the water potential gradient across the stem segment (MPa), multiplied by the path length measured (m).;Kh; Measure of the efficiency of water flow through a stem segment. It is calculated as the ratio of the water flow rate (kg/s) to the water potential gradient (MPa) across the stem segment, multiplied by the stem length (m).
  • type: numeric
  • units: 10^6.kg.m/MPa/s
  • allowable range: 0.01 - 2000 10^6.kg.m/MPa/s

hydraulic_safety_margin_50

  • label: Hydraulic safety margin, 50%
  • description: The difference between the shoot water potential [EnvThes:22311] (tension [PATO:0002284] (water pressure) in stem [PO:0009047] xylem vessels [PO:0025417]) at which the mean turgor pressure [TO:0000701] of cells [CL:0000000] drops to zero and the water potential at which the hydraulic conductivity [EnvThes:22133] has declined by 50% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;The difference between the stem water potential at turgor loss and the water potential at which 50% of conductivity is lost.
  • type: numeric
  • units: MPa
  • allowable range: -5 - 10 MPa

hydraulic_safety_margin_88

  • label: Hydraulic safety margin, 88%
  • description: The difference between the shoot water potential [EnvThes:22311] (tension [PATO:0002284] (water pressure) in stem [PO:0009047] xylem vessels [PO:0025417]) at which the mean turgor pressure [TO:0000701] of cells [CL:0000000] drops to zero and the water potential at which the hydraulic conductivity [EnvThes:22133] has declined by 88% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;The difference between the stem water potential at turgor loss and the water potential at which 88% of conductivity is lost.
  • type: numeric
  • units: MPa
  • allowable range: -5 - 10 MPa

water_potential_88percent_lost_conductivity

  • label: Stem xylem pressure, 88% lost conductivity (P88)
  • description: The tension [PATO:0002284] (water pressure) (water potential [EnvThes:22311]) in stem [PO:0009047] xylem vessels [PO:0025417] at which the hydraulic conductivity [EnvThes:22133] has declined by 88% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;A measure of stem hydraulic vulnerability, which is the stem xylem water pressure value at which stem hydraulic conductivity has declined by 88% from the mean maximum rate of hydraulic conductivity.
  • type: numeric
  • units: MPa
  • allowable range: -20 - 0 MPa

fire_time_from_fire_to_fruiting

  • label: Time from fire to fruiting
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until one individual [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] produces mature [PATO:0001701] fruit [PO:0009001].;The time interval from a fire until the first individual in a population produces a mature fruit with viable propagules.
  • type: numeric
  • units: mo
  • allowable range: 1 - 500 mo

seed_dormancy_class

  • label: Dormancy type
  • description: Physical [PATO:0002079], physiological [NCIT:C16990], and morphological [PATO:0000051] mechanisms [NCIT:C25343] that inhibit mature [PATO:0001701] seeds [PO:0009010] from germinating [GO:0009845].;Mechanisms that inhibit mature seeds from germinating and the environmental or experimental cues that allow a seed to break dormancy and germinate.
  • type: categorical
  • allowable values:
    • morphological_dormancy: Seeds exhibit morphological dormancy, generally because an underdeveloped embryo needs to grow.
    • morphophysiological_dormancy: Seeds exhibit morphophysiological dormancy, with both morphological and physiological dormancy mechanisms that must be overcome.
    • non_dormant: Seeds are non-dormant.
    • physical_dormancy: Seeds exhibit physical dormancy, due to one or more water-impermeable layers in the seed coat.
    • physiological_dormancy: Seeds exhibit physiological dormancy, whereby chemical changes are required within the seed to break dormancy, including temperature stratification and after-ripening.

fruiting_time

  • label: Fruiting time, by month
  • description: The months [UO:0000035] during which a plant [PO:0000003] produces fruit [PO:0009001] with mature [PATO:0001701] seeds [PO:0009010].;Months during which taxon produces fruit with mature seeds.
  • type: categorical
  • allowable values:
    • **:

plant_diameter_breast_height

  • label: Stem diameter at breast height
  • description: A stem morphology trait [TO:0000361] which is the maximum [NCIT:C25564] diameter at breast height [ECSO:00000549] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341].;Maximum diameter at breast height (DBH) for a taxon in a given location. It is commonly referred to as “DBH”.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 5000 mm

seed_germination

  • label: Seed germination proportion
  • description: The proportion [PATO:0001470] of mature [PATO:0001701] seeds [PO:0009010] in a sample that germinated [GO:0009845] under the specified [NCIT:C38024] environmental conditions [ENVO:01000203] or experimental conditions [XCO:0000000].;Proportion of mature seeds in a sample that germinated under specified environmental or experimental conditions.
  • type: numeric
  • units: {count}/{count}
  • allowable range: 0 - 1 {count}/{count}

serotiny

  • label: Serotiny
  • description: A fire response trait which indicates whether a fruit [PO:0009001] or cone [strobilus; PO:0025083] remains closed in the canopy [ENVO:01001242] after maturation [TO:0002661], with the seed [PO:0009010] only being released [NCIT:C42882] after a specific [NCIT:C38024] environmental conditions [ENVO:01000203], usually an environmental cue associated with a fire [ENVO:01000786], but possibly also drought or extreme heat.;Categorical variable describing whether a fruit or cone remains closed in the plant canopy after maturation until an environmental cue, usually a fire, but possibly also drought or extreme heat, triggers it to release its seeds.
  • type: categorical
  • allowable values:
    • not_serotinous: Plant does not display serotiny.
    • serotinous: Plant serotinous, level not specified.
    • serotiny_high: Plant displays a strong level of serotiny, indicating all seeds are retained as a canopy seedbank until an environmental cue triggers their release.
    • serotiny_low: Plant displays a weak level of serotiny, indicating it releases most of its seeds gradually, with only a small proportion of the seeds retained as a canopy seedbank until an environmental cue triggers the release of the entire seedbank.
    • serotiny_moderate: Plant displays a moderate level of serotiny, indicating it releases some of its seeds gradually, with the remainder retained as a canopy seedbank until an environmental cue triggers the release of the entire seedbank.

leaf_light_respiration_per_area

  • label: Leaf respiration rate per unit leaf area, in the light (Rday)
  • description: The cellular respiration [GO:0045333] rate [PATO:0000161] of plant leaves [PO:0025034] exposed to ambient [NCIT:C62306] photosynthetically active radiation [ENVO:01001864] levels, recorded per unit leaf area [PATO:0001323].;The respiration rate of a leaf in ambient light (ambient photosynthetically active radiation), calculated as the production of CO2 per unit leaf area.
  • type: numeric
  • units: nmol{CO2}/m2/s
  • allowable range: 10 - 10000 nmol{CO2}/m2/s

leaf_senesced_N_per_dry_mass

  • label: Senesced leaf nitrogen (N) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of nitrogen in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 100 mg/g

leaf_senesced_P_per_dry_mass

  • label: Senesced leaf phosphorus (P) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of phosphorus in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.005 - 15 mg/g

wood_P_per_dry_mass

  • label: Wood phosphorus (P) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of phosphorus in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

cell_cross-sectional_area

  • label: Cell cross-sectional area
  • description: A plant cell size [TO:0002684] trait which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a cell [CL:0000000].;Cell cross-sectional area.
  • type: numeric
  • units: um2
  • allowable range: 100 - 10000 um2

leaf_cuticle_thickness_abaxial

  • label: Cuticle thickness on the lower leaf surface
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039].;Thickness of the cuticle on the lower leaf surface.
  • type: numeric
  • units: um
  • allowable range: 0.1 - 100 um

leaf_cuticle_thickness_adaxial

  • label: Cuticle thickness on the upper leaf surface
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039].;Thickness of the cuticle on the upper leaf surface.
  • type: numeric
  • units: um
  • allowable range: 0.1 - 100 um

leaf_epidermis_thickness_abaxial

  • label: Lower leaf side epidermis thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].;Thickness of the epidermis on the lower side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 1 - 100 um

leaf_epidermis_thickness_adaxial

  • label: Upper leaf side epidermis thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].;Thickness of the epidermis on the upper side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 1 - 100 um

leaf_palisade_tissue_thickness_abaxial

  • label: Lower palisade mesophyll thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].;Thickness of palisade mesophyll tissue on the lower side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 10 - 1000 um

leaf_palisade_tissue_thickness_adaxial

  • label: Upper palisade mesophyll thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].;Thickness of palisade mesophyll tissue on the upper side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 10 - 1000 um

leaf_spongy_mesophyll_thickness

  • label: Spongy mesophyll cell thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of spongy mesophyll [PO:0005647] tissue [CARO:0000043] within the leaf lamina [PO:0020039].;Thickness of spongy mesophyll tissue in a leaf.
  • type: numeric
  • units: um
  • allowable range: 10 - 1000 um

nitrogen_fixing

  • label: Plant nitrogen fixation capacity
  • description: A trait which indicates whether a plant has the capacity to engage in symbioses [GO:0044403] with microorganisms [NCIT:C14329] (bacteria [NCIT:C14187], cyanobacteria [NCBITaxon:1117], and fungi [NCBITaxon:4751]) capable of nitrogen fixation [GO:0009399] and usually identified by nodulation [GO:0009877] on the plant roots [PO:0009005].;Binary variable describing whether a plant has the capacity for nitrogen fixatation, indicating it engages in symbioses with microorganisms (most commonly bacteria or cyanobacteria) that are capable of nitrogen fixation. Nitrogen fixation capacity is usually identified by nodules on the plant roots.
  • type: categorical
  • allowable values:
    • nitrogen_fixer: Plant hosts nitrogen-fixing microbes in root nodules.
    • non_nitrogen_fixer: Plant does not host nitrogen-fixing microbes.

photosynthetic_pathway

  • label: Plant photosynthetic pathway
  • description: The photosynthetic pathway, or biochemical reaction [NCIT:C19899], used by a plant to convert carbon dioxide [ENVO:01000451] into carbohydrates [CHEBI:16646], with options being C3 photosynthesis [GO:0019253], C4 photosynthesis [GO:0009760], and CAM photosynthesis [GO:0009761].;Type of photosynthetic pathway displayed by a plant.
  • type: categorical
  • allowable values:
    • c3: Plant using the ‘standard’ photosynthetic pathway, where a 3-carbon compound is produced after the first stage in the photosynthetic pathway.
    • c3-c4: Taxa that shift between C3 and C4 photosynthesis.
    • c3-cam: Taxa that shift between C3 and cam photosynthesis.
    • c4: Plants in which the photosynthetic light-dependent reactions and the Calvin cycle are physically separated to reduce photorespiration.
    • c4-cam: Taxa that shift between C4 and cam photosynthesis.
    • cam: Plants which display crassulacean acid metabolism.
    • facultative_cam: Species that shifts between C3 and CAM photosynthesis depending on water availability.
    • unknown: Photosynthetic pathway is unknown. (This trait value should be used if the photosynthetic pathway is assessed but unknown, not to record a lack of data.)

leaf_hypodermis_thickness_adaxial

  • label: Upper leaf side hypodermis thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].;Thickness of the hypodermis on the upper side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 1 - 100 um

leaf_hypodermis_thickness_abaxial

  • label: Lower leaf side hypodermis thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].;Thickness of the hypodermis on the lower side of the leaf.
  • type: numeric
  • units: um
  • allowable range: 1 - 100 um

leaf_inclination_angle

  • label: Leaf inclination angle
  • description: A leaf position trait [TO:0000541] which is the inclination angle [PATO:0002326] of the adaxial [PATO:0002047] leaf lamina [PO:0020039] surface relative to a horizontal [PATO:0001855] plane, such that an adaxial leaf surface facing the solar zenith [ECSO:00002723] has an angle of 0.;The inclination angle of the upper (adaxial) leaf lamina surface relative to a horizontal plane, such that a leaf surface facing the solar zenith has an angle of 0.
  • type: numeric
  • units: deg
  • allowable range: -90 - 90 deg

leaf_stomatal_resistance_ambient

  • label: Leaf stomatal water vapour resistance under ambient conditions
  • description: The stomatal [PO:0008032] water vapour resistance [SWEET:propConductivity/Resistance] of plant leaves [PO:0025034] exposed to ambient [NCIT:C62306] photosynthetically active radiation [ENVO:01001864] levels and ambient atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033], recorded per unit leaf area [PATO:0001323]. Stomatal resistance is the resistance to the diffusion [NCIT:C82333] of carbon dioxide [CHEBI:16526] through stomata.;Leaf stomatal water vapour resistance under ambient light (photosynthetically active radiation) levels and ambient carbon dioxide concentrations.
  • type: numeric
  • units: s/m
  • allowable range: 10 - 1e+06 s/m

woodiness

  • label: Woodiness
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait which indicates whether the plant stem [PO:0009047] (the vegetative shoot system [PO:0025607]) is woody, with “woody” defined to include both plant stems with secondary xylem (true wood) [PO:0005848] and those with wood-like lignification [CHEBI:6457] (woodiness in monocots and ferns).;A binary trait, indicating whether a plant’s stems are woody, with woody broadly defined to include both true wood (secondary xylem) and taxa that do not produce secondary xylem (i.e. monocots and ferns) but have thick, stiff, robust lignified stems.
  • type: categorical
  • allowable values:
    • herbaceous: Plant with non-lignified stems.
    • woody: Plant whose stems are stiff and lignified, including both stems that produce secondary xylem and those that produce only lignified primary tissues but are notably stiff and robust.

leaf_reflectance

  • label: Leaf visible light reflection
  • description: A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is reflected [SWEET:procWave/Reflection] by a leaf.;Proportion of incoming visible light (between 400-700 nm) that is reflected by a leaf.
  • type: numeric
  • units: umol/umol
  • allowable range: 0.01 - 1 umol/umol

leaf_reflectance_near_infrared

  • label: Leaf infra-red light reflection
  • description: A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming infrared radiation [ENVO:21001214] that is reflected [SWEET:procWave/Reflection] by a leaf.;Proportion of incoming near infra-red light (between 750-10500 nm) that is reflected by a leaf.
  • type: numeric
  • units: umol/umol
  • allowable range: 0.1 - 1 umol/umol

leaf_water_content_per_fresh_mass

  • label: Leaf water content per unit leaf fresh mass
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf fresh mass [APD:glossary/40044].;The ratio of the mass of water in a fresh leaf (leaf water content) to leaf fresh mass
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 10 g/g

leaf_P_resorption

  • label: Leaf phosphorus resorption efficiency
  • description: A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf phosphorus [CHEBI:28659] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf phosphorus content in a live [PATO:0001421] leaf, indicating what proportion of phosphorus is resorbed [NCIT:C93204] from the senescent leaf before it is shed.;A leaf nutrient recycling trait which is one minus the ratio of leaf phosphorus content in a senescent leaf (leaf litter) to leaf phosphorus content in a live leaf, indicating what proportion of phosphorus is resorbed from the senescent leaf before it is shed.
  • type: numeric
  • units: mg/mg
  • allowable range: -1.1 - 1 mg/mg

leaf_lifespan

  • label: Leaf lifespan
  • description: The duration [PATO:0001309] that an individual leaf [PO:0025034] is attached to the stem [PO:0009047] and is physiologically functional [PATO:0001510].;Duration of time that an individual leaf is attached to the stem and physiologically functional; leaf longevity.
  • type: numeric
  • units: mo
  • allowable range: 1 - 1000 mo

seed_P_per_seed_dry_mass

  • label: Seed phosphorus (P) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of phosphorus in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 100 mg/g

leaf_B_per_dry_mass

  • label: Leaf boron (B) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of boron in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.01 - 1000 mg/kg

leaf_Ca_per_dry_mass

  • label: Leaf calcium (Ca) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of calcium in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 1000 mg/g

leaf_Cu_per_dry_mass

  • label: Leaf copper (Cu) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of copper in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.01 - 150 mg/kg

leaf_Fe_per_dry_mass

  • label: Leaf iron (Fe) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of iron in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 1 - 10000 mg/kg

leaf_K_per_dry_mass

  • label: Leaf potassium (K) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of potassium in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

leaf_Mg_per_dry_mass

  • label: Leaf magnesium (Mg) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of magnesium in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10000 mg/g

leaf_Mn_per_dry_mass

  • label: Leaf manganese (Mn) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of manganese in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 1 - 20000 mg/kg

leaf_Ni_per_dry_mass

  • label: Leaf nickel (Ni) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nickel [CHEBI:28112] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of nickel in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.1 - 100 mg/kg

leaf_S_per_dry_mass

  • label: Leaf sulphur (S) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of sulphur in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 50 - 50000 mg/kg

leaf_Zn_per_dry_mass

  • label: Leaf zinc (Zn) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of zinc in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.1 - 1000 mg/kg

leaf_Mo_per_dry_mass

  • label: Leaf molybdenum (Mo) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of molybdenum in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0 - 100 mg/kg

leaf_Si_per_dry_mass

  • label: Leaf silicon (Si) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of silicon [CHEBI:27573] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of silicon in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 200 mg/g

leaf_ash_per_dry_mass

  • label: Leaf ash content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of leaf [PO:0025034] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the leaf dry mass before the combustion process.;The ratio of the mass of leaf ash remaining after a combustion process to the leaf dry mass before the combustion process.
  • type: numeric
  • units: g/g
  • allowable range: 1e-04 - 1 g/g

leaf_phenol_per_dry_mass

  • label: Total leaf phenolic content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phenol [CHEBI:33853] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where phenol content is measured as gallic acid [CHEBI:30778] equivalents.;The ratio of the mass of phenol in a leaf to leaf dry mass, where phenol content is measured as gallic acid equivalents.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 1000 mg/g

pollination_syndrome

  • label: Pollination syndrome
  • description: A classification [NCIT:C25161] based on a taxon’s [NCIT:C40098] flower morphology traits [TO:0000499] that indicates the taxa (biotic vectors) or abiotic mechanisms [NCIT:C25343] that most likely pollinate [GO:0009856] a flower [PO:0009046].;For a given taxon, the biotic vectors (including explicit taxa) or abiotic mechanisms that are most likely to pollinate its flowers, based on the taxon’s flower morphology.
  • type: categorical
  • allowable values:
    • abiotic: Floral syndrome indicates pollination occurs through wind, water, or collision of free-floating particles.
    • animal: Floral syndrome suggests flowers are animal-pollinated.
    • ballistic: Floral syndrome suggests pollen is launched away from the plant by explosion.
    • bat: Floral syndrome suggests flowers are bat-pollinated.
    • bee: Floral syndrome suggests flowers are bee-pollinated.
    • beetle: Floral syndrome suggests flowers are beetle-pollinated.
    • biotic: Floral syndrome suggests flowers are pollinated by a biotic agent.
    • bird: Floral syndrome suggests flowers are bird-pollinated.
    • butterfly: Floral syndrome suggests flowers are butterfly-pollinated.
    • fly: Floral syndrome suggests flowers are fly-pollinated.
    • insect: Floral syndrome suggests flowers are insect-pollinated, type of insect not specified.
    • mammal: Floral syndrome suggests flowers are mammal-pollinated.
    • moth: Floral syndrome suggests flowers are moth-pollinated.
    • pollination_rare: Plants mostly reproduce through vegetative spread and flowering is rare.
    • self: Floral syndrome suggests flowers are primarily self-pollinating.
    • thrip: Floral syndrome suggests flowers are thrip-pollinated.
    • vertebrate: Floral syndrome suggests flowers are vertebrate-pollinated (bats, birds, mammals or lizards).
    • wasp: Floral syndrome suggests flowers are wasp-pollinated.
    • water: Floral syndrome suggests flowers are water-pollinated.
    • wind: Floral syndrome suggests flowers are wind-pollinated.

leaf_area_ratio

  • label: Leaf area ratio (LAR)
  • description: A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of a plant’s [PO:0000003] total [NCIT:C25304] leaf [PO:0025034] area [PATO:0001323] to total plant dry mass [TO_0000352].;The ratio of total plant leaf area to total plant dry mass; LAR.
  • type: numeric
  • units: mm2/mg
  • allowable range: 0.05 - 100 mm2/mg

integrated_plant_transpiration

  • label: Integrated plant transpiration
  • description: Rate [PATO:0000161] of water [CHEBI:15377] loss integrated across an entire plant [PO:0000003], recorded per unit leaf [PO:0025034] area [PATO:0001323] and measured across a duration [PATO:0001309] of at least one day.;The rate of water loss integrated across an entire plant, as calculated across longer periods of time under experimental conditions, recorded per unit leaf area.
  • type: numeric
  • units: g/cm2{leaf}/d
  • allowable range: 0.01 - 1 g/cm2{leaf}/d

bulk_modulus_of_elasticity

  • label: Bulk modulus of elasticity (e)
  • description: A parameter [STATO:0000034] calculated from a pressure-volume curve [APD:glossary/40124], which is, for a shoot or leaf [PO:0025034], the ratio [PATO:0001470] of the change in cell [CL:0000000] turgor potential (negative turgor pressure [TO:0000701]) over the relative change in cell volume (change in volume/initial volume). It is an estimation of cell wall [GO:0009505] rigidity.;A parameter calculated from a pressure-volume curve which indicates the elasticity or rigidity of cells in leaves. It is the ratio of the change in cell turgor potential to the change in relative cell volume (change in volume/initial volume).
  • type: numeric
  • units: MPa
  • allowable range: 0.1 - 100 MPa

leaf_relative_water_content_at_turgor_loss_point

  • label: Leaf relative water content at turgor loss point
  • description: A parameter [STATO:0000034] calculated from a pressure-volume curve [APD:glossary/40124], which is, for a leaf [PO:0025034], the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in the leaf at the turgor [PATO:0001620] loss point to the mass of water in the leaf at full hydration (saturated [NCIT:C61427]; relative water content = 100%).;Ratio of the mass of water in leaf at the point the leaf cells lose turgor to the mass of water in a fully saturated leaf. A fully saturated leaf is one which has been rehydrated to hold the maximum amount of water and is considered to have a relative water content of 100%. The plant’s turgor loss point is calculated from a pressure-volume curve.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 1 g/g

root_shoot_ratio

  • label: Root to shoot ratio
  • description: A whole plant morphology trait [TO:0000398] which is the ratio [PATO:0001470] of a plant’s root dry mass [TO:0000078] to its shoot dry mass [TO:0000552].;The ratio of total root system dry mass to total shoot system dry mass.
  • type: numeric
  • units: mg{root}/mg{shoot}
  • allowable range: 0.01 - 15 mg{root}/mg{shoot}

leaf_soluble_starch_per_mass

  • label: Leaf soluble starch content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of water soluble starch in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 200 mg/g

leaf_soluble_sugars_per_mass

  • label: Leaf soluble sugar content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].;The ratio of the mass of sugar in a leaf to leaf dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 200 mg/g

root_soluble_starch_per_mass

  • label: Root soluble starch content per unit root dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in roots [PO:0009005] to root dry mass.;The ratio of the mass of water soluble starch in roots to root dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

root_soluble_sugars_per_mass

  • label: Root soluble sugar content per unit root dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in roots [PO:0009005] to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].;The ratio of the mass of sugar in roots to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 200 mg/g

stem_soluble_starch_per_mass

  • label: Stem soluble starch content per unit stem dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a stem [PO:0009047] to stem dry mass.;The ratio of the mass of water soluble starch in a stem to stem dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

stem_soluble_sugars_per_mass

  • label: Stem soluble sugar content per unit stem dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a stem [PO:0009047] to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].;The ratio of the mass of sugar in a stem to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 200 mg/g

leaf_stomatal_distribution

  • label: Stomatal distribution
  • description: A leaf stomatal complex frequency trait [TO:0000566] which indicates whether stomata [PO:0008032] occur on the abaxial [PATO:0002046] and/or adaxial [PATO:0002047] leaf lamina [PO:0020039].;Distribution of stomatal across the two leaf surfaces.
  • type: categorical
  • allowable values:
    • amphistomatic: Leaves with stomata on both leaf surfaces.
    • amphistomatic_uneven: Leaves with stomata on both leaf surfaces but more stomata on the lower leaf surface.
    • epistomatic: Leaves that only have stomata on the upper (adaxial) leaf surface.
    • hypostomatic: Leaves that only have stomata on the lower (abaxial) leaf surface.

leaf_work_to_punch

  • label: Leaf work to punch
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] compressive strength [SWEET:propPressure/CompressiveStrength; MESH:D019245]. It is the force [PATO:0001035] (work) required to punch a hole in a leaf lamina (fracture [SWEET:phenSolid/Fracture]) with a punch and die.;The force (work) required to punch a hole through a leaf lamina, generally with a punch and die. This is a measure of leaf compressive strength or toughness.
  • type: numeric
  • units: N/m
  • allowable range: 0.5 - 10000 N/m

leaf_work_to_punch_adjusted

  • label: Leaf specific work to punch
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] compressive strength [SWEET:propPressure/CompressiveStrength; MESH:D019245]. It is the force [PATO:0001035] (work) required to punch a hole in a leaf lamina (fracture [SWEET:phenSolid/Fracture]) with a punch and die, scaled by leaf thickness [PATO:0000915].;The force (work) required to punch a hole through a leaf lamina, generally with a punch and die, adjusted for leaf thickness. This is a measure of leaf compressive strength or toughness.
  • type: numeric
  • units: N/m2
  • allowable range: 1000 - 3e+07 N/m2

leaf_work_to_shear

  • label: Leaf work to shear
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] compressive strength [SWEET:propPressure/CompressiveStrength; MESH:D019245]. It is the force [PATO:0001035] required to shear [SWEET:phenSolid/Shear] a leaf lamina with a pair of scissors.;The force (work) required to shear a leaf. This is a measure of leaf compressive strength or toughness that is equivalent to cutting a leaf with scissors.
  • type: numeric
  • units: N
  • allowable range: 0.01 - 10 N

leaf_work_to_shear_adjusted

  • label: Leaf specific work to shear (fracture toughness)
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] compressive strength [SWEET:propPressure/CompressiveStrength; MESH:D019245]. It is the force [PATO:0001035] required to shear [SWEET:phenSolid/Shear] a leaf lamina with a pair of scissors, scaled by leaf thickness [PATO:0000915].;The force (work) required to shear a leaf, adjusted to leaf thickness; also referred to as ‘fracture toughness’. This is a measure of leaf compressive strength or toughness that is equivalent to cutting a leaf with scissors.
  • type: numeric
  • units: N/m
  • allowable range: 100 - 10000 N/m

leaf_work_to_tear_adjusted

  • label: Leaf specific work to tear
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] tensile strength [SWEET:propPressure/TensileStrength; MESH:D013718]. It is the force [PATO:0001035] required to tear or rip [SWEET:phenSolid/Rip] a leaf lamina, scaled by leaf thickness [PATO:0000915].;The force (work) required to tear/rip a leaf, adjusted to leaf thickness. This is a measure of leaf tensile strength.
  • type: numeric
  • units: N/m
  • allowable range: 10 - 15000 N/m

flower_pollen_aperture_shape

  • label: Pollen grain aperture shape
  • description: A floral organ morphology trait [TO:0000862] which is the shape [PATO:0000052], structure [PATO:0000141], and position [PATO:0000140] of the pollen grain [PO:0025281] apertures [GO:0062074].;Pollen grains are often described according to the shape, structure, and position of their apertures. This trait describes the main shape of the many described aperture types.
  • type: categorical
  • allowable values:
    • absent: Pollen that lacks an aperture.
    • colporus: Pollen aperture shape that combines the groove of colpate and the pore of porate apertures.
    • colpus: Pollen aperture shape is elongate and grooved and may be positioned globally or equatorially.
    • pore: Pollen aperture shape is round and pore-like and may be positioned globally or equatorially.
    • ring_like: Aperture that completely encircles the pollen.
    • spiraperturate: Pollen grain with one or more spiral apertures.
    • sulcus: Pollen aperture shape is elongate and grooved and positioned at the pole.
    • syncolpus: Pollen grain with two or more simple (or compound) colpi, the ends of which join at the pole.
    • ulcus: Pollen aperture shape is round and pore-like and positioned at the pole.

flower_pollen_apertures_count

  • label: Number of pollen grain apertures
  • description: A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of pollen apertures [GO:0062074] per pollen [PO:0025281] grain.;The number of apertures per pollen grain as a continuous character.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 100 {count}

flower_ovary_position

  • label: Ovary position
  • description: A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the plant ovary [PO:0009072] relative to the other flower [PO:0009046] organs [PO:0009008], in particular whether the ovary is attached above or below the perianth [PO:0009058] organs (the petals [PO:0009032] and sepals [PO:0009031]).;The position of the flower ovary relative to the insertion level of the remaining floral organs.
  • type: categorical
  • allowable values:
    • half_inferior: Flower where the receptacle is surrounded by the ovary to its mid-level.
    • inferior: Flower with inferior ovary, where the ovary is embedded in the receptacle and therefore located below the insertion level of the remaining floral organs.
    • one_quarter_inferior: Flower where the receptacle surrounds up to one quarter of the ovary.
    • superior: Flower with superior ovary, where the ovary is positioned above the insertion level of the remaining floral organs.
    • three_quarters_inferior: Flower where the receptacle surrounds more than three quarters of the ovary.

flower_perianth_merism

  • label: Number of perianth parts in each whorl (Perianth merism)
  • description: A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) in each floral whorl [PO:0025023].;The number of perianth parts in each whorl, recorded as a continuous trait (with integer values of 1 and above).
  • type: numeric
  • units: {count}
  • allowable range: 0 - 50 {count}

flower_perianth_parts_count

  • label: Number of perianth parts
  • description: A perianth morphology trait [TO:0000275] which is the total count [NCIT:C25463] of perianth [PO:0009058] parts, including sepals [PO:0009031], petals [PO:0009032], and any form of tepal [PO:0009033].;The total number of perianth parts, including sepals, petals, or any form of tepal.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 1000 {count}

flower_anther_orientation

  • label: Anther orientation
  • description: An androecium morphology trait [TO:0000994] which is the position [PATO:0000140] and orientation [PATO:0000133] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released.;Orientation of anther at anthesis.
  • type: categorical
  • allowable values:
    • apical: When thecae are positioned in a transverse position at the tip of the connective and thus dehisce upward in the flower (e.g., Sinofranchetia, Endress and Hufford, 1989).
    • extrorse: When the stomia of the thecae face the floral periphery.
    • introrse: When the stomia of the thecae face the floral centre.
    • latrorse: When pollen is released toward the side (i.e., toward neighbouring anthers).

flower_filament

  • label: Filament presence and shape
  • description: A floral organ morphology trait [TO:0000862] which is whether a filament [PO:0009067] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the relative length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the filament.;Trait that considers the absence or presence of the filament, and in the latter case, the shape of the filament. Shape is considered in terms of length and width and is defined in relation to anther length/width.
  • type: categorical
  • allowable values:
    • absent: No filaments present
    • fused_into_fascicle_stalks: Special case where the stamens are fused into the stalk of a fascicle of flowers.
    • fused_into_synandrium: Filament structure formed when stamens fused together.
    • long_and_narrow: Filaments have a long and narrow shape. (Synonyms, filiform, slender)
    • long_and_narrow_divided_in_two: Filaments have a long and narrow shape that is divided in two; e.g. Adoxa.
    • long_and_very_wide: Filaments have a long and very wide shape; for instance, the petal-like (outer) stamens of Nymphaea.
    • long_and_wide: Filaments have a long and wide laminar shape.
    • petaloid: Filaments are long and wide and colorful, e.g. Canna.
    • present_general: Filaments present, with no specific defining characteristics.
    • short_and_narrow: Filaments have a short and narrow shape; subsessile anthers.
    • short_and_wide: Filaments have a short and wide laminar shape.
    • short_general: Filament is short relative to the anther.

flower_filament_fusion

  • label: Fusion of filaments
  • description: A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that is fused [NCIT:C124537] together at anthesis [PO:0009068].;Fusion of stamen (and staminode) filaments among each other at anthesis (congenitally or postgenitally) is recorded on a continuous scale, from 0 (free filaments) to 1 (filaments fused along their entire length).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 1 {dimensionless}

flower_perianth_symmetry

  • label: Symmetry of perianth
  • description: A perianth morphology trait [TO:0000275] which is the symmetry [PATO:0000965] of the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), whereby a perianth is only considered actinomorphic [PATO:0001328] if all perianth whorls [PO:0025023] are actinomorphic.;Trait capturing the symmetry of the perianth.
  • type: categorical
  • allowable values:
    • actinomorphic_general: Perianth displaying radial symmetry.
    • actinomorphic_rotational: Perianth that can be equally divided into three or more identical sections that, when rotated around the center of the flower by some number of degrees, exactly match each other in orientation and shape.
    • actinomorphic_spiral: Perianth lacking a distinct plane of symmetry because their perianth parts are spirally arranged.
    • actinomorphic_strictly_polysymmetric: Polysymmetry, with three or more planes of bilateral symmetry.
    • asymmetric: Perianth lacking any planes of symmetry.
    • disymmetric: Perianth with two orthogonal planes of bilateral symmetry.
    • zygomorphic: Monosymmetric, with a single plane of bilateral symmetry.

flower_style_differentiation

  • label: Style differentiation
  • description: A floral organ morphology trait [TO:0000862] which is whether a style [PO:0006469] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the style relative to the dimensions of the ovary [PO:0009072].;Presence or absence of a style, and, when present, the shape of the style in terms of length and width in relation to ovary length and width.
  • type: categorical
  • allowable values:
    • absent: Stigma sessile.
    • continuous: When the style is an apical extension of carpel with decurrent stigma.
    • present_length_and_shape_unknown: Style present but shape unknown.
    • present_long_and_narrow: Style present, and long and narrow, including long filiform styles.
    • present_long_and_wide: Style present, and long and wide.
    • present_petaloid: Style present and petaloid.
    • present_short_and_narrow: Style present, and short and narrow, including short filiform styles.
    • present_short_and_thick: Style present, and short and thick.

flower_filament_fusion_to_inner_perianth

  • label: Fusion of filaments to inner perianth series
  • description: A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that are fused [NCIT:C124537] to the innermost perianth [PO:0009058] organ [PO:0009008] at anthesis [PO:0009068].;The fusion of filaments with the innermost perianth organs at anthesis on a continuous scale, from 0 (filaments completely free from perianth) to 1 (filaments fused along their entire length with the perianth; the ‘entire length’ of the filament is defined as the distance between the floral base and the joint between filament and anther).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 1 {dimensionless}

flower_structural_sex_type

  • label: Floral structural sex
  • description: A floral organ morphology trait [TO:0000862] which is the phenotypic sex [PATO:0001894] of a flower [PO:0009046], indicating whether individual flowers have both male [PATO:0000384] and female [PATO:0000383] reproductive structures [UBERON:0005156] (bisexual) or only one type of reproductive organ (unisexual).;Trait that captures floral sex, distinguishing among three states, bisexual, incompletely unisexual, and unisexual.
  • type: categorical
  • allowable values:
    • bisexual: Hermaphrodite; flowers with both male and female reproductive organs.
    • incompletely_unisexual: Male flowers with pistillodes and/or female flowers with staminodes.
    • unisexual: Flowers with either male or female reproductive organs.

flower_androecium_structural_merism

  • label: Number of androecium parts in each whorl (Androecium structural merism)
  • description: An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of stamens [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023].;The number of stamens or stamen bundles (fascicles) in one whorl, considering both fertile stamens and staminodes.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 100 {count}

flower_gynoecium_placentation

  • label: Placentation
  • description: A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the ovules [PO:0020003] within the ovary [PO:0009072], known as placentation [APD:glossary/40074].;The different types of placentation in apocarpous/unicarpellate and in syncarpous gynoecia.
  • type: categorical
  • allowable values:
    • apical: Ovules attach at the top of the ovary; pendulous; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
    • axile: Ovules are placed in the angle between carpel flanks in the center of the ovary; in syncarpous gynoecia.
    • basal: Ovules attach at the base of the ovary; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
    • free_central: Ovules are attached to a central column that emerges from the base of the ovary and protrudes into the non-septate ovary; in syncarpous gynoecia.
    • laminar: When there is more than one series of ovules at the flanks of a carpel; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
    • marginal: Ovules attach along the ventral slit; in apocarpous and unicarpellate gynoecia.
    • parietal: Ovules attach to the ovary wall where two carpels meet; in syncarpous gynoecia.

flower_ovules_per_functional_carpel_count

  • label: Number of ovules per functional carpel
  • description: A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of ovules [PO:0020003] per fertile [PATO:0000955] carpel [PO:0009030].;Number of ovules per carpel recorded as a continuous character (with integer values of 1 and above).
  • type: numeric
  • units: {count}
  • allowable range: 0 - 1e+05 {count}

flower_structural_carpels_count

  • label: Number of structural carpels
  • description: A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of both fertile [PATO:0000955] and female sterile [PATO:0000892] carpels [PO:0009030] in a hermaphrodite [PATO:0001340] flower [PO:0009046] or female [PATO:0000383] flower.;Number of fertile or sterile carpels in bisexual or female flowers, recorded as a continuous character (with integer values of 1 and above).
  • type: numeric
  • units: {count}
  • allowable range: 0 - 10000 {count}

flower_perianth_fusion

  • label: Fusion of perianth
  • description: A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) that are fused [NCIT:C124537] together at anthesis [PO:0009068].;Fusion of perianth organs (congenital or postgenital) at anthesis, recorded on a continuous scale, from 0 (free parts) to 1 (parts fused along their entire length).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 1 {dimensionless}

flower_anther_attachment

  • label: Anther attachment
  • description: An androecium morphology trait [TO:0000994] which indicates how the filament [PO:0009067] is attached to [PATO:0001667] the tissue [CARO:0000043] connecting the two thecae [PO:0009069] (pollen [PO:0025281] sacs) of the anthers [PO:0009066], affecting the spatial pattern [PATO:0000060] of the stamens [PO:0009029].;Anther attachment refers to the area of insertion of the filament on the anther connective (i.e. the tissue connecting the two thecae of an anther).
  • type: categorical
  • allowable values:
    • basifixed: Filament attached to the base of the connective. Laminar stamens are classified as basifixed.
    • dorsifixed: Filament attached to the dorsal side of the anther.
    • dorsifixed_at_base: Filament attached to the dorsal side of the anther, at the base.
    • ventrifixed: Filament attached to the ventral side of the anther.

flower_fertile_stamens_count

  • label: Number of fertile stamens
  • description: A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of fertile [PATO:0000955] stamens [PO:0009029] on a hermaphrodite [PATO:0001340] flower [PO:0009046] or male [PATO:0000384] flower.;Count of fertile stamens in bisexual or male flowers; does not score female flowers.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 10000 {count}

flower_anther_dehiscence

  • label: Anther dehiscence
  • description: An androecium morphology trait [TO:0000994] which is the length [PATO:0000122] and shape [PATO:0000052] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the usually slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released.;Anther dehiscence refers to the type of opening of the anther when releasing its pollen through the stomia.
  • type: categorical
  • allowable values:
    • circular: Anther with a transverse slit that goes entirely around the anther.
    • common_stomium_confluent_thecae: The two thecae of a single anther are fused at one end with a single pore through which pollen exits.
    • flap_valvate: Anther with one or more flap-like valves on horizontal hinges, opening vertically.
    • h_valvate: Anther with two valves on longitudinal hinges, opening horizontally as salon doors; when the stomium bifurcates at its distal and/or proximal end and thus a valve is formed.
    • longitudinal_slit: Anther with longitudinal slits that extend along the entire length of each theca.
    • pores_many: Anther with pollen exiting from many pores along its length.
    • poricidal: Anther where dehiscence of longitudinal slits is incomplete, with pollen exiting via a small pore at the tip.
    • short_apical_slits: Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the apical end of the theca.
    • short_basal_slits: Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the base of the theca.
    • short_central_slits: Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the center of the theca.
    • t_valvate: Anther with T-shaped stomium; reduced transverse slit at base.
    • transverse_slit: Anther with horizontal slit.

flower_androecium_structural_phyllotaxis

  • label: Androecium structural phyllotaxis
  • description: An androecium morphology trait [TO:0000994] which is the spatial pattern or distribution [PATO:0000060] (phyllotaxy [TO:0006014]) of stamen [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023].;Structural phyllotaxy of the androecium, considering both fertile stamens and staminodes. In cases of stamen fascicles, it is the phyllotaxis of fascicles, not individual stamens that are recorded.
  • type: categorical
  • allowable values:
    • irregular: Stamens and staminodes have an irregular arrangement.
    • spiral: Stamens and staminodes have a spiral arrangement.
    • whorled: Stamens and staminodes arranged in whorls.

flower_androecium_structural_whorls_count

  • label: Number of androecium structural whorls
  • description: An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of whorls [PO:0025023] comprising the androecium [PO:0009061], considering both stamen [PO:0009029] and staminodes [PO:0009077]. If stamens are grouped into fascicles [NCIT:C79692], the trait is the count of whorls of fascicles.;The number of whorls, considering both fertile stamens and staminodes. In cases of stamen fascicles, the count of whorls of fascicles is recorded.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 100 {count}

flower_gynoecium_phyllotaxis

  • label: Gynoecium phyllotaxis
  • description: A gynoecium [PO:0009062] morphology trait which is either the count [NCIT:C25463] of whorls [PO:0025023] of carpels [PO:0009030] comprising the gynoecium or other descriptors of carpel phyllotaxy [TO:0006014].;Structural phyllotaxy of the gynoecium.
  • type: categorical
  • allowable values:
    • one_whorl: Flowers with a single whorl of carpels.
    • spiral: Flowers with carpels in a spiral arrangement.
    • two_or_more_whorls: Flowers with two or more whorls of carpels.

flower_ovary_fusion

  • label: Fusion of ovaries
  • description: A floral organ morphology trait [TO:0000862] which, for flowers with multiple carpels, is the proportion [PATO:0001470] of the length [PATO:0000122] of the ovaries [PO:0009072] that are fused [NCIT:C124537] together.;Degree of ovary fusion expressed as a fraction of the total length of the ovary (from the floral base to the apex of the ovary).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 1 {dimensionless}

flower_perianth_differentiation

  • label: Perianth differentiation
  • description: A floral organ morphology trait [TO:0000862] which indicates whether petals [PO:0009032] and sepals [PO:0009031] are differentiated or identical and the various ways they can be identical or different.;The ways in which perianth organs may look different from each other in a given flower.
  • type: categorical
  • allowable values:
    • continuous_differentiation_among_petaloid_tepals: A flower with series of tepals that all resemble petals, which display continuous, gradual differentiation between whorls.
    • continuous_differentiation_among_sepaloid_tepals: A flower with series of tepals that all resemble sepals, which display continuous, gradual differentiation between whorls.
    • continuous_differentiation_outer_sepaloid_inner_petaloid: A flower with multiple whorls, where there is continuous differentiation between the outer sepaloid parts and the inner petaloid parts.
    • marked_differentiation: A flower with marked differentiation between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
    • marked_differentiation_among_petaloid_tepals: A flower with tepals that all resemble petals, but with marked differentiation between whorls of tepals.
    • marked_differentiation_among_sepaloid_tepals: A flower with tepals that all resemble sepals, but with marked differentiation between whorls of tepals.
    • marked_differentiation_outer_sepaloid_inner_petaloid: A typical flower with a distinct calyx and corolla.
    • undifferentiated: Flower were all tepals are alike, including flowers with a single, undifferentiated whorl.
    • very_weak_differentiation: A flower with very weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
    • weak_differentiation: A flower with weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.

flower_perianth_phyllotaxis

  • label: Perianth phyllotaxis
  • description: A perianth morphology trait [TO:0000275] which is the phyllotaxy [TO:0006014] of the perianth [PO:0009058] parts (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), at anthesis [PO:0009068];Perianth phyllotaxis at anthesis.
  • type: categorical
  • allowable values:
    • irregular: Perianth parts lacking any symmetrical organisation.
    • spiral: Perianth parts organised along a continuous spiral, usually with wide divergence angles more or less equal to 137.5 degrees, the Fibonacci ‘golden angle’.
    • whorled: Perianth parts organised in one or more whorls.

flower_perianth_whorls_count

  • label: Number of perianth whorls
  • description: A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of floral whorls [PO:0025023] that comprise the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]).;The number of perianth whorls as a continuous trait (with integer values of 1 and above).
  • type: numeric
  • units: {count}
  • allowable range: 0 - 15 {count}

flower_style_fusion

  • label: Fusion of styles
  • description: A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of styles [PO:0006469] that are fused [NCIT:C124537] together.;The degree of fusion of styles at anthesis, recorded on a continuous scale, from 0 (free styles) to 1 (styles fused along their entire length, but excluding the stigmatic region).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 1 {dimensionless}

flower_anther_connective_extension

  • label: Connective extension (apical)
  • description: An androecium morphology trait [TO:0000994] which is whether anthers [PO:0009066] have protrusions [NCIT:C50710] that extend above the two theca [PO:0009069] (pollen [PO:0025281] sacs), and, if present, the shape [PATO:0000052] and the relative length [PATO:0000122] of the protrusion relative to the length of the entire anther.;Apical (distal) connective extensions (also called ‘distal connective protrusions’) are sterile anther structures that distally extend beyond the level of the thecae (i.e., the two lateral pairs of pollen sacs of a tetrasporangiate anther).
  • type: categorical
  • allowable values:
    • absent: Apical connective extensions are lacking.
    • long_extension: Connective extension more than a third but less than length of anther; e.g. Idiospermum australiense.
    • present_general: Apical connective extensions are present but no information on their length relative to the anthers.
    • short_extension: Connective extension less than a third of the length of anther.
    • very_long_extension: Connective extension more than length of anther; e.g. Galbulimima belgraveana.

flower_colour

  • label: Flower colour
  • description: A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the flower [PO:0009046. The petals [PO:0009032] (corolla [PO:0009059]), sepals [PO:0009031] (calyx [PO:0009060]), or the entire perianth [PO:0009058] may be responsible for the main colour observed.;The dominant flower colour, where the entire flower, the petals, or the sepals may be responsible for the observed colour.
  • type: categorical
  • allowable values:
    • black: Flower colour is black.
    • blue_purple: Flower colour is blue or purple.
    • green: Flower colour is green.
    • grey: Flower colour is grey.
    • pink: Flower colour is pink.
    • red_brown: Flower colour is red or brown.
    • white_cream: Flower colour is white or cream.
    • yellow_orange: Flower colour is yellow or orange.

perianth_colour

  • label: Perianth colour
  • description: A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]).;The dominant colour of the perianth parts (sepals, petals and tepals).
  • type: categorical
  • allowable values:
    • black: Perianth colour is black.
    • blue: Perianth colour is blue.
    • brown: Perianth colour is brown.
    • green: Perianth colour is green.
    • grey: Perianth colour is grey.
    • orange: Perianth colour is orange.
    • pink: Perianth colour is pink.
    • purple: Perianth colour is purple.
    • red: Perianth colour is red.
    • white: Perianth colour is white.
    • yellow: Perianth colour is yellow.

flower_diameter

  • label: Flower diameter
  • description: A floral organ morphology trait [TO:0000862] which is the maximum diameter [PATO:0001334] of the flower [PO:0009046] at anthesis [PO:0009068], considering all floral organs, but most commonly the diameter from petal [PO:0009032] tip to petal tip.;The maximum diameter of the flower at anthesis, considering all floral organs, but most commonly the diameter from petal tip to petal tip.
  • type: numeric
  • units: cm
  • allowable range: 0.01 - 200 cm

flower_pollen_length

  • label: Pollen grain length
  • description: A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of a pollen [PO:0025281] grain.;The largest dimension of the pollen grain.
  • type: numeric
  • units: um
  • allowable range: 0 - 1000 um

flower_length

  • label: Flower length
  • description: A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the flower [PO:0009046] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the flower, measured at anthesis [PO:0009068]. It excludes the length of the pedicel [PO:0030112].;The length of the flower.
  • type: numeric
  • units: cm
  • allowable range: 0.01 - 200 cm

flower_orientation

  • label: Floral orientation
  • description: A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] and orientation [PATO:0000133] of the flower [PO:0009046].;The position and orientation of the flower.
  • type: categorical
  • allowable values:
    • down: Flower orientation is pendant, with the floral entrance pointing down.
    • lateral: Flower orientation is horizontal, with the floral entrance perpendicular to main vertical axis.
    • mixed: Flower orientation is mixed, with different flowers exhibiting different orientations.
    • up: Flower orientation is vertical, with the floral entrance pointing up.

atmospheric_CO2_concentration

  • label: Ambient CO2 concentration (ca)
  • description: The atmospheric carbon dioxide [ENVO:01000451] concentration [PATO:0000033].;Atmospheric CO2 concentration (external CO2 concentration); ca.
  • type: numeric
  • units: umol{CO2}/mol
  • allowable range: 50 - 2000 umol{CO2}/mol

leaf_chloroplast_CO2_concentration

  • label: CO2 concentration inside chloroplasts (cc)
  • description: The concentration [PATO:0000033] of carbon dioxide [ENVO:01000451] inside chloroplasts [GO:0009507] in leaves [PO:0025034].;CO2 concentration inside chloroplasts in leaves; cc.
  • type: numeric
  • units: umol{CO2}/umol
  • allowable range: 10 - 1000 umol{CO2}/umol

leaf_photosynthesis_Jmax_per_mass

  • label: Leaf Jmax per unit leaf mass (Jmax)
  • description: Maximum [NCIT:C25564] electron transport [MESH:D004579] rate [PATO:0000161] through the photosynthetic [GO:0015979] electron transport chain [GO:0022900], recorded per unit leaf [PO:0025034] mass [PATO:0000125]. The value is a model parameter [STATO:0000034] derived from the portion of the photosynthesis A-ci response curve at high carbon dioxide concentrations where ATP and NADPH production by the light reactions [GO:0019684] limits the rate of carboxylation [MOP:0000591].;The maximum potential photosynthetic electron transport rate, recorded per unit leaf mass. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at high carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the rate of ATP and NADPH production by the light reactions.
  • type: numeric
  • units: umol/g/s
  • allowable range: 0 - 3 umol/g/s

leaf_photosynthesis_Vcmax_per_mass

  • label: Leaf Vcmax per unit leaf mass (Vcmax)
  • description: Maximum [NCIT:C25564] carboxylation [MOP:0000591] rate [PATO:0000161] by the Rubisco complex [GO:0048492] in plant leaves [PO:0025034], recorded per unit leaf [PO:0025034] mass [PATO:0000125]. The value is a model parameter [STATO:0000034] derived from the initial slope of a photosynthesis [GO:0015979] A-ci response curve, the component that is carbon dioxide [CHEBI:16526] limited and hence limited by Rubisco activity.;The apparent maximum rate of Rubisco (ribulose 1,5-bisphosphate carboxylase/oxygenase) activity carboxylase activity (carboxylation capacity), recorded per unit leaf mass. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at low carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the Rubisco activity.
  • type: numeric
  • units: umol/g/s
  • allowable range: 0 - 2 umol/g/s

leaf_fluorescence_fv_over_fm

  • label: Leaf maximum quantum yield (Fv/Fm)
  • description: Maximum [NCIT:C25564] quantum yield [TO:0012004] of photosystem II [GO:0009539], as determined through chlorophyll fluorescence [GO:0090546]. Quantum yield is the ratio [PATO:0001470] of photons [NCIT:C88112] absorbed to photons emitted [SIO:000384] through fluorescence.;A trait that indicates the health of photosystem II in a dark adapted leaf. Quantum yield is calculated as the ratio of variable fluorescence (Fv) to maximum fluorescence (Fm), with a higher value indicating a healthier leaf. Variable fluorescence (Fv) is calculated as the difference in fluorescence between a leaf where all photsystems have been saturated by light (Fm) and a leaf in a dark-adapted state (minimum fluorescence; Fo).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.2 - 1 {dimensionless}

osmotic_potential

  • label: Osmotic potential
  • description: In a plant tissue [CARO:0000043], the potential energy [ENVO:2000016] of water [CHEBI:15377] molecules to flow [NCIT:C41141] across a semi-permeable [PATO:0000982] membrane [GO:0016020] due to the differences in solute [SIO:010418] concentration [PATO:0000033] across the membrane, such that the osmotic potential becomes more negative as the solute concentration increases or the cell water content decreases. Osmotic potential is equal to negative osmotic pressure [EnvThes:21558] and is also referred to as solute potential.;The potential energy of water molecules to move across a semi-permeable membrane due to differences in solute concentration across the membrane; solute potential
  • type: numeric
  • units: MPa
  • allowable range: -10 - 0 MPa

root_mass_fraction

  • label: Root mass fraction
  • description: A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of the total root dry mass [TO:0000078] to whole plant [PO:0000003] dry mass [APD:glossary/40080].;The ratio of total root system dry mass to whole plant dry mass.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.01 - 1 mg/mg

stem_mass_fraction

  • label: Stem mass fraction
  • description: A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of whole plant [PO:0000003] stem [PO:0009047] dry [PATO:0001824] mass [PATO:0000125] to whole plant dry mass [APD:glossary/40080].;The ratio of stem dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of stems and branches.
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 1 g/g

leaflet_dry_mass

  • label: Leaflet dry mass
  • description: A leaf dry mass trait [TO:0001014] which is the dry mass of a single leaflet [PO:0020049]. This measurement will not include the petiole [PO:0020038].;The oven dry mass of a single leaflet.
  • type: numeric
  • units: mg
  • allowable range: 0.005 - 15000 mg

leaflet_area

  • label: Leaflet area
  • description: A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of a single leaflet [PO:0020049] lamina [PO:0025060]. This measurement will not include the petiole [PO:0020038].;The 2-dimensional projected (scanned) area of the surface of the lamina of a single leaflet.
  • type: numeric
  • units: mm2
  • allowable range: 0.1 - 1e+06 mm2

branch_mass_fraction

  • label: Side branch dry mass to whole plant dry mass ratio
  • description: A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of side branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to whole plant [PO:0000003] dry mass.;The ratio of side branch dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of all branches off the main shoot axis.
  • type: numeric
  • units: mg/mg
  • allowable range: 0 - 1 mg/mg

leaf_water_content_per_area

  • label: Leaf water content per unit leaf area (leaf succulence)
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of water in a fresh leaf (leaf water content) to leaf surface area; leaf succulence.
  • type: numeric
  • units: g/m2
  • allowable range: 10 - 5000 g/m2

plant_succulence

  • label: Plant succulence
  • description: A whole plant [PO:0000003] phenotype [NCIT:C16977] trait that indicates whether any plant tissues [CARO:0000043] are succulent [FLOPO:0900041], indicating they are thick [PATO:0000591] and fleshy [PATO:0002351], generally storing water [CHEBI:15377] as a drought tolerance [TO:0000276] adaptation or filling with water as a salt tolerance [TO:0006001] adaptation.;Trait indicating whether the leaves or stems of a plant are thick and fleshy because they store water, generally an adaptation to arid or salty environments.
  • type: categorical
  • allowable values:
    • not_succulent: Neither stems nor leaves are thick and fleshy.
    • semi-succulent: A plant has tissues that are somewhat thick and fleshy, including many chenopods.
    • succulent: A plant is designated as succulent but it is not specified which tissues are thick and fleshy.
    • succulent_leaves: Leaves are thick and fleshy.
    • succulent_stems: Stems are thick and fleshy.

leaf_mass_fraction

  • label: Leaf mass fraction
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of a plant’s total leaf dry mass [APD:glossary/40080] to total plant dry mass [TO:0000352].;The ratio of leaf dry mass to total plant dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 1 g/g

leaf_relative_water_content_predawn

  • label: Leaf relative water content predawn
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] harvested pre-dawn [SWEET:reprTimeDay/Dawn] to the mass of water in a leaf at full hydration (saturated [NCIT:C61427]; relative water content = 100%).;Ratio of the mass of water in a fresh leaf collected pre-dawn to the mass of water in a fully saturated leaf. A fully saturated leaf is one which has been rehydrated to hold the maximum amount of water and is considered to have a relative water content of 100%.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 1.4 g/g

leaf_water_use_efficiency_integrated

  • label: Integrated water use efficiency
  • description: The ratio [PATO:0001470] of biomass [ENVO:01000155] production [EnvThes:21071] (carbon dioxide [ENVO:01000451] fixation) to transpiration [ENVO:02500035] (water [CHEBI:15377] loss) across a given time interval [TimeInterval].;The rate of carbon dioxide fixation relative to water loss, calculated as biomass production / transpiration. This measures how much biomass is produced relative to transpiration across a given time interval, and is therefore an integrated measure of water use efficiency; WUE, transpiration ratio
  • type: numeric
  • units: g{biomass}/g{H2O}
  • allowable range: 0.1 - 50 g{biomass}/g{H2O}

leaf_stomatal_density_abaxial

  • label: Stomatal density on the lower leaf surface
  • description: A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323].;Density of stomata on the lower leaf surface.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 1 - 1000 {count}/mm2

leaf_stomatal_density_adaxial

  • label: Stomatal density on the upper leaf surface
  • description: A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323].;Density of stomata on the upper leaf surface.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 0 - 1000 {count}/mm2

leaf_stomatal_density_average

  • label: Stomatal density averaged across both leaf surfaces
  • description: A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] to leaf lamina [PO:0020039] area [PATO:0001323], averaged the across the abaxial [PATO:0002046] and adaxial [PATO:0002047] sides of the leaf lamina.;Density of stomata averaged across both leaf surfaces.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 0 - 1000 {count}/mm2

resprouting_capacity_proportion_individuals

  • label: Post-fire proportion resprouting individuals
  • description: A fire response trait which is the proportion [PATO:0001470] of mature [PATO:0001701] individuals [EFO:0000542] in a population [PCO:0000001] that have vegetative meristematic tissue [PO:0006079] (buds [PO:0000055]) that resprouts [Cerrado:fire#PlantResprout] following a fire [ENVO:01000786].;Proportion of mature individuals in a population that survive a fire and resprout following a fire.
  • type: numeric
  • units: {count}/{count}
  • allowable range: 0 - 1 {count}/{count}

root_structure

  • label: Plant root structures
  • description: A trait which indicates types [NCIT:C25284] of root [PO:0009005] structures [PATO:0000141], including structures associated with mutualisms [GO:0085030] with mycorrhizae [EnvThes:20656].;Specific specialized types of root structures and root symbioses, especially symbioses with mycorrhizal fungi.
  • type: categorical
  • allowable values:
    • arbuscular_mycorrhizal: Symbiosis in which the mycorrhizal fungi does penetrate the cortex cells of the plant roots and its hyphae form arbuscules and vesicles inside the plant root; endomycorrhizal, AM, VAM.
    • carnivorous: Plant has non-mycorrhizal carnivorous root.
    • cluster_roots: Non-proteoid taxa with cluster roots.
    • coiling_vam: Associations that spread predominantly by intracellular hyphal coils within roots.
    • dauciform_root: A type of cluster root found in the family Cyperaceae.
    • ectomycorrhizal: Symbiosis in which the mycorrhizal fungi does not penetrate the cortex cells of the plant roots.
    • ericoid_mycorrhizal: A form of arbuscular mycorrhizal relationship only found on plants in the order Ericales. The fungi’s hyphae penetrate the plant root but do not form arbuscules.
    • fine_roots: Particularly fine roots for nutrient uptake.
    • haustorium: A highly modified stem or root of a parasitic plant, such as mistletoe , or a specialized branch or tube originating from a hairlike filament (hypha) of a fungus. The haustorium penetrates the tissues of a host and absorbs nutrients and water.
    • hemiparasitic_root: Plant with a partially parasitic root.
    • long_root_hairs: Plant with specialised long root hairs.
    • mycorrhizal: Plant mycorrhizal but type not specified.
    • non_mycorrhizal: Plants lacking a mycorrhizal symbiont.
    • orchid_mycorrhizal: A mycorrhizal relationship specific to orchid taxa, and for most orchids, essential for seedlings to establish.
    • parasitic_root: Plant with a parasitic root.
    • proteoid_root: Also known as cluster roots, are plant roots that form clusters of closely spaced short lateral rootlets and aid in nutrient upake in nutrient-poor soils; common in members of the family Proteaceae but present in other families as well.
    • root_hairs: Plant with specialised root hairs.
    • sand-binding: Persistent sheaths of sand that form around the roots of taxa in the families Restionaceae, Cyperaceae, Haemodoraceae, and Lomandraceae in western Australia. Assumed to function in nutrient uptake.
    • saprophyte: Plant that acquires nutrients from decaying biomass.
    • subepidermal_mycorrhizal: Mycorrhizae where hyphae grow in a cavity under epidermal cells. Only in the genus Thysanotus (family Laxmaniaceae).
    • unknown: Root structures unknown.

root_system_classification

  • label: Root system morphology
  • description: A root system morphology trait [TO:0000985] which are categorical [STATO:0000252] trait values that capture the branchiness [PATO:0002009] and texture [PATO:0000150] of the root system [PO:0025025] and the root system depth [TO:0001058].;Categorical root system descriptions that indicate the branchiness and texture of the root system and the depth of the root system.
  • type: categorical
  • allowable values:
    • type_i: The primary root is fibrous or fleshy and is relatively long; mesophytic.
    • type_ii: The primary root is relatively short and is fibrous. Roots are well branched in a dry habitat but branch little in one that is wet.
    • type_iii: The primary root is usually long and fibrous. The first order lateral roots are well branched and are usually short; they are rather evenly distributed along the primary root; xerophytic.
    • type_iv: The primary root is usually long and slender like that of Type II. The first order lateral roots are well branched. Those closest to the surface of the ground, or not far below, are relatively long; xerophytic and mesophytic.
    • type_ix: Adventitious roots that arise on the aerial stem and branches of some herbaceous taxa, shrubs, and trees; on creepers, vines, and runners; and on subterranean rhizomes and stolons.
    • type_v: The primary root is divided-forked or branched-and fibrous, and with few first order lateral roots, or, in age, apparently with none; xerophytic.
    • type_vi: The primary root is usually as in Type V. The first order lateral roots are mostly on the upper portion of the primary root and are thus relatively superficial; xerophytic or in well aerated sandy soil where the rainfall is not light.
    • type_vii: Adventitious roots that are in one group and arise on a short horizontal, or vertical, axis of the shoot. They are fibrous or fleshly; mesophytic and hydrophytic.
    • type_viii: Adventitious roots that are in one group and are fibrous and fleshy, or fibrous and thick. The fibrous adventitious roots are absorbing-anchoring roots and are usually with laterals to the second order; meophytic.

root_specific_root_length

  • label: Specific root length (SRL)
  • description: A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] length [PATO:0000122] to root dry mass [TO:0000078].;The ratio of root length to root dry mass; SRL.
  • type: numeric
  • units: m/g
  • allowable range: 0.5 - 5000 m/g

genome_size

  • label: Plant genome size
  • description: The total mass [PATO:0000125] of chromosomes [NCIT:C13202] that comprise a plant’s genome [OMIT:0027643] in a somatic cell [CL:0002371].;Mass of all chromosomes, the genome, in a somatic cell.
  • type: numeric
  • units: pg
  • allowable range: 0.5 - 100 pg

root_wood_density

  • label: Root wood density
  • description: A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of the dry mass [PATO:0000125] of a unit of wood [PO_0005848] from a root [PO:0009005]to its fresh fresh [EnvThes:21976] volume [PATO:0000918].;The ratio of woody root dry mass to woody root fresh volume.
  • type: numeric
  • units: mg/mm3
  • allowable range: 0.05 - 10 mg/mm3

leaf_water_content_per_saturated_mass

  • label: Leaf water content per unit saturated leaf mass
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a saturated leaf [PO:0025034] to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].;The ratio of the mass of water in a leaf (leaf water content) to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximal water holding capacity and is at full turgor.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 10 g/g

leaf_chlorophyll_A_B_ratio

  • label: Ratio of leaf chlorophyll A content to leaf chlorophyll B content
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to the mass of chlorophyll b [CHEBI:27888] in the leaf.;The ratio of the mass of chlorophyll a in a leaf to the mass of chlorophyll b in the leaf.
  • type: numeric
  • units: umol/umol
  • allowable range: 0.1 - 6 umol/umol

leaf_chlorophyll_per_area

  • label: Leaf total chlorophyll content (chlorophyll A + B) per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf area [PATO:0001323], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content.;The ratio of the mass of chlorophyll in a leaf to leaf area, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
  • type: numeric
  • units: umol/m2
  • allowable range: 10 - 2000 umol/m2

leaf_soluble_protein_per_area

  • label: Leaf soluble protein content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of water soluble protein in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 20 g/m2

leaf_soluble_starch_per_area

  • label: Leaf soluble starch content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of water soluble starch in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 100 g/m2

leaf_soluble_sugars_per_area

  • label: Leaf soluble sugar content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf area [PATO:0001323], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].;The ratio of the mass of sugar in a leaf to leaf area, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 100 g/m2

fruit_P_per_dry_mass

  • label: Fruit phosphorus (P) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of phosphorus in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

fruit_dry_mass

  • label: Fruit dry mass
  • description: A fruit morphology trait [TO:0002629] which is the dry [PATO:0001824] mass [PATO:0000125] of a fruit [PO:0009001], including seeds [PO:0009010].;Fruit dry mass, including seeds. Fruits will generally be oven-dried.
  • type: numeric
  • units: mg
  • allowable range: 1e-04 - 1e+06 mg

competitive_stratum

  • label: Competitive stratum
  • description: A classification [NCIT:C25161] that indicates the vegetation layer [ENVO:01000355] a taxon [NCIT:C40098] reaches within its community [ECSO:00000310], used to assess competition [EnvThes:21097] within the community.;Categorical descriptions of a taxon’s relative stature in its community, used to assess competitive heirarchies within a community.
  • type: categorical
  • allowable values:
    • ground: Graminoids, herbs and ferns reaching maximum heights of c. 0.2-1 m.
    • mid: Shrubs reaching maximum heights of c. 1-2 m.
    • post_fire_ephemeral: Taxa that avoid competition with other strata by rapidly completing their life cycle after fire; a sub-category of ‘ground’.
    • upper: Tall shrubs reaching maximum heights of c. 3-5 m.

fire_time_from_fire_to_50_percent_fruiting

  • label: Time from fire until 50% of individuals are fruiting
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] after a fire [ENVO:01000786] until a 50% of the individuals [EFO:0000542] of a taxon [NCIT:C40098] in a population [PCO:0000001] produce mature [PATO:0001701] fruit [PO:0009001].;The time interval from a fire until 50% of the individuals in a population produce mature fruit.
  • type: numeric
  • units: mo
  • allowable range: 1 - 500 mo

leaf_Na_per_dry_mass

  • label: Leaf sodium (Na) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of sodium in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 1 - 20000 mg/kg

fire_flame_duration

  • label: Leaf flame duration
  • description: A fire response trait which is the duration [PATO:0001309] of the portion of fire [ENVO:01000786]-induced combustion [NCIT:C157201] process from the start of combustion until a flame [ENVO:01000789] is no longer visible.;Flame duration for a single leaf, which is the time from the first visible flame until the last visible flame.
  • type: numeric
  • units: s
  • allowable range: 0.1 - 200 s

fire_smoulder_duration

  • label: Leaf smoulder duration
  • description: A fire response trait which is the duration [PATO:0001309] of the portion of fire [ENVO:01000786]-induced combustion [NCIT:C157201] process from the time when a flame [ENVO:01000789] is no longer visible until smoulder combustion ends.;Smoulder duration for a single leaf, which is the time from the end of the last visible flame until the glowing phase dies out.
  • type: numeric
  • units: s
  • allowable range: 0.1 - 1000 s

fire_time_to_ignition

  • label: Fire time to ignition
  • description: A fire response trait which is the time interval (duration) [PATO:0001309] from a leaf’s [PO:0025034] exposure [NCIT:C17941] to a flame [ENVO:01000789] until the leaf ignites [APD:glossary/40040].;For a single leaf, the time from an ignition source contacting the leaf until ignition occurs.
  • type: numeric
  • units: s
  • allowable range: 0.1 - 100 s

fire_total_burn_duration

  • label: Leaf flame and smoulder duration
  • description: A fire response trait which is the duration [PATO:0001309] of the entire fire [ENVO:01000786]-induced combustion [NCIT:C157201] process from the time of the start of combustion until smoulder combustion [APD:glossary/40094] ends.;For a single leaf, the sum of flame duration and smoulder duration, which is the time from the first visible flame until the glowing phase dies out.
  • type: numeric
  • units: s
  • allowable range: 0.1 - 1500 s

leaf_density

  • label: Leaf tissue density
  • description: The ratio [PATO:0001470] of leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to leaf fresh [EnvThes:21976] volume [PATO:0000918].;The ratio of leaf dry mass to leaf fresh volume.
  • type: numeric
  • units: mg/mm3
  • allowable range: 0.05 - 1.1 mg/mm3

leaf_lignin_per_dry_mass

  • label: Leaf lignin content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of lignin in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 10 - 1000 mg/g

leaf_tannin_per_dry_mass

  • label: Leaf tannin content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (a class of polyphenols [CHEBI:26195]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of tannins (a class of polyphenols) in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 1000 mg/g

bark_Al_per_dry_mass

  • label: Bark aluminium (Al) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of aluminium in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

bark_B_per_dry_mass

  • label: Bark boron (B) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of boron in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 1 mg/g

bark_C_per_dry_mass

  • label: Bark carbon (C) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of carbon in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 50 - 750 mg/g

bark_Ca_per_dry_mass

  • label: Bark calcium (Ca) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of calcium in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

bark_Cu_per_dry_mass

  • label: Bark copper (Cu) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of copper in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1e-04 - 1 mg/g

bark_Fe_per_dry_mass

  • label: Bark iron (Fe) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of iron in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

bark_K_per_dry_mass

  • label: Bark potassium (K) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of potassium in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 100 mg/g

bark_Mg_per_dry_mass

  • label: Bark magnesium (Mg) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of magnesium in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

bark_Mn_per_dry_mass

  • label: Bark manganese (Mn) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of manganese in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 1 mg/g

bark_N_per_dry_mass

  • label: Bark nitrogen (N) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of nitrogen in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

bark_Na_per_dry_mass

  • label: Bark sodium (Na) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of sodium in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 100 mg/g

bark_P_per_dry_mass

  • label: Bark phosphorus (P) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of phosphorus in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

bark_S_per_dry_mass

  • label: Bark sulphur (S) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of sulphur in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

bark_Zn_per_dry_mass

  • label: Bark zinc (Zn) content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of zinc in bark to bark dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

bark_ash_per_dry_mass

  • label: Bark ash content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of bark [PO:0004518] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the bark dry mass before the combustion process.;The ratio of the mass of bark ash remaining after a combustion process to the bark dry mass before the combustion process.
  • type: numeric
  • units: g/g
  • allowable range: 1e-04 - 1 g/g

bark_cellulose_per_dry_mass

  • label: Bark cellulose content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of cellulose in bark to bark dry mass.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.1 - 1 mg/mg

bark_density

  • label: Bark density
  • description: The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding fresh [EnvThes:21976] bark volume [PATO:0000918].;The ratio of bark dry mass to bark fresh volume.
  • type: numeric
  • units: mg/mm3
  • allowable range: 0.1 - 1.1 mg/mm3

bark_lignin_per_dry_mass

  • label: Bark lignin content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in bark [PO:0004518] to bark dry mass.;The ratio of the mass of lignin in bark to bark dry mass.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.01 - 1 mg/mg

bark_tannin_per_dry_mass

  • label: Bark tannin content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (polyphenols [CHEBI:26195]) in bark [PO:0004518] to bark dry mass.;The ratio of the mass of tannins (polyphenols) in bark to bark dry mass.
  • type: numeric
  • units: mg/mg
  • allowable range: 1e-05 - 1 mg/mg

bark_water_content_per_dry_mass

  • label: Bark water content per unit bark dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] (bark water content [APD:glossary/40014]) to bark dry [PATO:0001824] mass.;The ratio of the mass of water in bark (bark water content) to bark dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 1 g/g

leaf_cellulose_per_dry_mass

  • label: Leaf cellulose content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of cellulose in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.01 - 1 mg/mg

leaf_senesced_Al_per_dry_mass

  • label: Senesced leaf aluminium (Al) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of aluminium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

leaf_senesced_B_per_dry_mass

  • label: Senesced leaf boron (B) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of boron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

leaf_senesced_C_per_dry_mass

  • label: Senesced leaf carbon (C) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of carbon in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 1 g/g

leaf_senesced_Ca_per_dry_mass

  • label: Senesced leaf calcium (Ca) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of calcium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 1000 mg/g

leaf_senesced_Cu_per_dry_mass

  • label: Senesced leaf copper (Cu) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of copper in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.01 - 100 mg/kg

leaf_senesced_Fe_per_dry_mass

  • label: Senesced leaf iron (Fe) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of iron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 1 - 10000 mg/kg

leaf_senesced_K_per_dry_mass

  • label: Senesced leaf potassium (K) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of potassium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

leaf_senesced_Mg_per_dry_mass

  • label: Senesced leaf magnesium (Mg) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of magnesium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 100 mg/g

leaf_senesced_Mn_per_dry_mass

  • label: Senesced leaf manganese (Mn) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of manganese in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 1 - 10000 mg/kg

leaf_senesced_Na_per_dry_mass

  • label: Senesced leaf sodium (Na) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of sodium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 10 - 20000 mg/kg

leaf_senesced_S_per_dry_mass

  • label: Senesced leaf sulphur (S) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of sulphur in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 100 - 50000 mg/kg

leaf_senesced_Zn_per_dry_mass

  • label: Senesced leaf zinc (Zn) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of zinc in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0 - 100 mg/kg

leaf_posture_numeric

  • label: Leaf lamina posture (leaf 3-dimensionality)
  • description: A leaf shape trait [TO:0000492] which is the maximum [NCIT:C25564] perpendicular [PATO:0002434] height [PATO:0000119] of a leaf [PO:0025034] placed on a flat surface, indicating the 3-D extent [PATO:0001710] of the leaf. This generally corresponds with whether the leaf is curled [FLOPO:0010468].;The maximum perpendicular height of a leaf when positioned on a flat surface.
  • type: numeric
  • units: mm
  • allowable range: 0 - 100 mm

fire_fuel_bed_bulk_density

  • label: Fuel bed bulk density
  • description: Fuel bed bulk density (volume density [PATO:0001353]), which is the ratio [PATO:0001470] of the mass [PATO:0000125] of the fuel bed divided by the fuel bed volume [PATO:0000918]; the term fuel bed refers to the accumulated plant litter [ENVO:01000628] on the ground [SWEET:realmSoil/Ground], as this is the fuel [CHEBI:33292] that is burnt in a ground fire [ENVO:01000786].;The density of the fuel bed, the plant litter accumulated on the ground. It is calculated as the ratio of the mass of the fuel bed to the fuel bed volume. This trait should be measured on plant litter from a single taxon.
  • type: numeric
  • units: g/cm3
  • allowable range: 0.001 - 1 g/cm3

fire_fuel_consumption

  • label: Fuel consumption by fire
  • description: For a given taxon [NCIT:C40098], the proportion [PATO:0001470] of plant biomass [ENVO:01000155] (fuel [CHEBI:33292]) that is consumed by a fire [ENVO:01000786].;Proportion of available plant biomass (live and dead) that is consumed by fire. This trait should be measured on biomass from a single taxon.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 100 g/g

fire_rate_of_spread

  • label: Fire rate of spread
  • description: For a given taxon [NCIT:C40098], the horizontal [PATO:0001855] speed [PATO:0000008] of fire [ENVO:01000786] movement [NCIT:C37994].;The horizontal rate of fire spread.
  • type: numeric
  • units: cm/s
  • allowable range: 0.01 - 1 cm/s

fruit_Ca_per_dry_mass

  • label: Fruit calcium (Ca) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of calcium in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.5 - 100 mg/g

fruit_K_per_dry_mass

  • label: Fruit potassium (K) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of potassium in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.5 - 100 mg/g

fruit_Mg_per_dry_mass

  • label: Fruit magnesium (Mg) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of magnesium in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.05 - 10 mg/g

fruit_N_per_dry_mass

  • label: Fruit nitrogen (N) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of nitrogen in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.5 - 100 mg/g

fruit_S_per_dry_mass

  • label: Fruit sulphur (S) content per unit fruit dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in fruit [PO:0009001] to fruit dry mass.;The ratio of the mass of sulphur in fruit to fruit dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.05 - 10 mg/g

seed_Ca_per_seed_dry_mass

  • label: Seed calcium (Ca) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of calcium in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 100 mg/g

seed_K_per_seed_dry_mass

  • label: Seed potassium (K) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of potassium in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 1 - 100 mg/g

seed_Mg_per_seed_dry_mass

  • label: Seed magnesium (Mg) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of magnesium in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

seed_N_per_seed_dry_mass

  • label: Seed nitrogen (N) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of nitrogen in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 10 - 1000 mg/g

seed_S_per_seed_dry_mass

  • label: Seed sulphur (S) content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in seeds [PO:0009010] to seed dry mass.;The ratio of the mass of sulphur in seeds to seed dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

stem_N_per_dry_mass

  • label: Stem nitrogen (N) content per unit stem dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a stem [PO:0009047] to stem dry mass.;The ratio of the mass of nitrogen in a stem to stem dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 1000 mg/g

wood_Ca_per_dry_mass

  • label: Wood calcium (Ca) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of calcium in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

wood_K_per_dry_mass

  • label: Wood potassium (K) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of potassium in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 100 mg/g

wood_Mg_per_dry_mass

  • label: Wood magnesium (Mg) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of magnesium in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

wood_Na_per_dry_mass

  • label: Wood sodium (Na) content per unit wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in wood [PO:0005848] to wood dry mass.;The ratio of the mass of sodium in wood to wood dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

wood_dead_Ca_per_dry_mass

  • label: Dead wood calcium (Ca) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of calcium in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

wood_dead_K_per_dry_mass

  • label: Dead wood potassium (K) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of potassium in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

wood_dead_Mg_per_dry_mass

  • label: Dead wood magnesium (Mg) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of magnesium in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 10 mg/g

wood_dead_N_per_dry_mass

  • label: Dead wood nitrogen (N) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of nitrogen in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

wood_dead_Na_per_dry_mass

  • label: Dead wood sodium (Na) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of sodium in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 1 mg/g

wood_dead_P_per_dry_mass

  • label: Dead wood phosphorus (P) content per unit dead wood dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.;The ratio of the mass of phosphorus in senescent wood (litter) to the dry mass of the senesced wood.
  • type: numeric
  • units: mg/g
  • allowable range: 0.001 - 10 mg/g

flower_N_per_dry_mass

  • label: Flower nitrogen (N) content per unit flower dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in flowers [PO:0009046] to flower dry mass.;The ratio of the mass of nitrogen in flowers to flower dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.5 - 100 mg/g

leaf_Al_per_dry_mass

  • label: Leaf aluminium (Al) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of aluminium in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0.01 - 12000 mg/kg

leaf_fresh_mass_per_area

  • label: Leaf fresh mass per leaf area
  • description: A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of the leaf [PO:0025034] fresh mass [APD:glossary/40044]. to the leaf area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038].;The ratio of leaf fresh mass to leaf area, measured on an entire leaf including the petiole.
  • type: numeric
  • units: g/m2
  • allowable range: 200 - 2000 g/m2

leaf_photosynthetic_nitrogen_use_efficiency_maximum

  • label: Leaf photosynthetic nitrogen use efficiency during Amax measurement (PNUE)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its nitrogen [CHEBI:25555] content, where photosynthesis is measured in plant leaves whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033].;The ratio of photosynthesis (CO2 assimilation rate) to leaf nitrogen content, where photosynthesis has been measured in a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations; PNUE.
  • type: numeric
  • units: umol{CO2}/g{N}/s
  • allowable range: 0.1 - 1000 umol{CO2}/g{N}/s

leaf_photosynthetic_phosphorus_use_efficiency_maximum

  • label: Leaf photosynthetic phosphorus use efficiency during Amax measurement PPUE)
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] photosynthetic rate [TO:0001015] to its phosphorus [CHEBI:28659] content, where photosynthesis is measured in plant leaves whose photosystem [GO:0009521] I and II reaction centers [GO:0009538; GO:0009539] are exposed to saturating [NCIT:C61427] photosynthetically active radiation [ENVO:01001864] and saturating atmospheric carbon dioxide [ENVO:01000451] concentrations [PATO:0000033].;The ratio of photosynthesis (CO2 assimilation rate) to leaf phosphorus content, where photosynthesis has been measured in a leaf exposed to saturating light (photosynthetically active radiation) levels and saturating carbon dioxide concentrations; PPUE.
  • type: numeric
  • units: umol{CO2}/g{P}/s
  • allowable range: 1 - 10000 umol{CO2}/g{P}/s

leaf_N_resorption

  • label: Leaf nitrogen resorption efficiency
  • description: A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf nitrogen [CHEBI:25555] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf nitrogen content in a live [PATO:0001421] leaf, indicating what proportion of nitrogen is resorbed [NCIT:C93204] from the senescent leaf before it is shed.;A leaf nutrient recycling trait which is one minus the ratio of leaf nitrogen content in a senescent leaf (leaf litter) to leaf nitrogen content in a live leaf, indicating what proportion of nitrogen is resorbed from the senescent leaf before it is shed.
  • type: numeric
  • units: mg/mg
  • allowable range: -1.1 - 1 mg/mg

leaf_cell_wall_N_per_cell_wall_dry_mass

  • label: Leaf cell wall nitrogen (N) per unit cell wall dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the cell wall dry mass in the leaf.;The ratio of the mass of nitrogen in the plant cell wall in a leaf to the cell wall dry mass in the leaf.
  • type: numeric
  • units: mmol{N}/g
  • allowable range: 0.1 - 10 mmol{N}/g

leaf_cell_wall_N_per_leaf_N

  • label: Leaf cell wall nitrogen (N) per unit leaf N content
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the mass of nitrogen in the leaf.;The ratio of the mass of nitrogen in the plant cell wall in a leaf to the mass of nitrogen in the leaf.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.01 - 1 mg/mg

leaf_cell_wall_fraction

  • label: Leaf cell wall fraction
  • description: A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of cell wall [GO:0009505] dry mass to leaf dry mass [TO:0001014].;The ratio of leaf cell wall dry mass to leaf dry mass. This is the fraction of cell wall material that can be recovered from total leaf biomass.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.1 - 1 mg/mg

leaf_guard_cell_length

  • label: Guard cell length
  • description: A leaf stomatal complex frequency trait [TO:0000566] which indicates the length [PATO:0000122] of a guard cell [PO:0000293].;Length of guard cells.
  • type: numeric
  • units: um
  • allowable range: 10 - 200 um

leaf_senesced_Mo_per_dry_mass

  • label: Senesced leaf molybdenum (Mo) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].;The ratio of the mass of molybdenum in a senescent leaf (leaf litter) to the senescent leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 0 - 30 mg/kg

leaf_epidermis_Ca_per_fresh_mass

  • label: Leaf epidermis calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in epidermal cells to epidermal cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 500 umol/g

leaf_epidermis_P_per_fresh_mass

  • label: Leaf epidermis phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in epidermal cells to epidermal cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 10 umol/g

leaf_hypodermis_Ca_per_fresh_mass

  • label: Leaf hypodermis calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in the hypodermis to hypodermis fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 500 umol/g

leaf_hypodermis_P_per_fresh_mass

  • label: Leaf hypodermis phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in the hypodermis to hypodermis fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 10 umol/g

leaf_palisade_mesophyll_Ca_per_fresh_mass

  • label: Leaf palisade mesophyll cell calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in palisade mesophyll cells to palisade mesophyll cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 100 umol/g

leaf_palisade_mesophyll_P_per_fresh_mass

  • label: Leaf palisade mesophyll cell phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in palisade mesophyll cells to palisade mesophyll cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 30 umol/g

leaf_sclerenchyma_Ca_per_fresh_mass

  • label: Leaf sclerenchyma cell calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in sclerenchyma cells to sclerenchyma cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 100 umol/g

leaf_sclerenchyma_P_per_fresh_mass

  • label: Leaf sclerenchyma cell phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in sclerenchyma cells to sclerenchyma cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 10 umol/g

leaf_spongy_mesophyll_Ca_per_fresh_mass

  • label: Leaf spongy mesophyll cell calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in spongy mesophyll cells to spongy mesophyll cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 1000 umol/g

leaf_spongy_mesophyll_P_per_fresh_mass

  • label: Leaf spongy mesophyll cell phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in spongy mesophyll cells to spongy mesophyll cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 30 umol/g

leaf_internal_parenchyma_Ca_per_fresh_mass

  • label: Leaf internal parenchyma cell calcium (Ca) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044].;The ratio of the mass of calcium in parenchyma cells to parenchyma cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 1 - 500 umol/g

leaf_internal_parenchyma_P_per_fresh_mass

  • label: Leaf internal parenchyma cell phosphorus (P) content per unit leaf fresh mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044].;The ratio of the mass of phosphorus in parenchyma cells to parenchyma cell fresh mass.
  • type: numeric
  • units: umol/g
  • allowable range: 0 - 10 umol/g

accessory_cost_fraction

  • label: Seed accessory cost fraction
  • description: A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is one minus the ratio [PATO:0001470] of total plant seed [PO:0009010] dry [PATO:0001824] mass [PATO:0000125] to total plant reproductive shoot system tissue dry mass where reproductive tissues include all flower buds [PO:0000056], flowers [PO:0009046], fruits [PO:0009001], dispersal [EnvThes:21038] tissues, and seeds produced during the developmental process involved in reproduction [GO:0003006].;The fraction of total reproductive investment required to mature a seed that is invested in non-seed tissues. It is calculated as one minus, the ratio of total biomass investment in seeds to total biomass investment in all reproductive tissues, including flower buds, flowers, fruits, dispersal tissues, aborted seeds, and successfully matured seeds.
  • type: numeric
  • units: mg/mg
  • allowable range: 0.01 - 1 mg/mg

accessory_cost_mass

  • label: Seed accessory cost mass
  • description: A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is the sum of the dry [PATO:0001824] mass [PATO:0000125] of all flower buds [PO:0000055], flowers [PO:0009046], fruits [PO:0009001], aborted seeds, and dispersal [EnvThes:21038] tissues produced during the developmental process involved in reproduction [GO:0003006], but excludes mature [PATO:0001701] seed [PO:0009010] dry mass.;The mass of seed accessory costs, which is the total biomass investment in all reproductive tissues, excluding biomass invested in successfully matured seeds; it includes all biomass invested in flower buds, flowers, fruits, dispersal tissues and aborted seeds.
  • type: numeric
  • units: mg
  • allowable range: 0.01 - 10000 mg

leaf_Cl_per_dry_mass

  • label: Leaf chlorine (Cl) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorine [CHEBI:23116] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of chlorine in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/kg
  • allowable range: 10 - 10000 mg/kg

leaf_photosynthesis_Jmax_over_Vcmax_25C

  • label: Leaf Jmax to leaf Vcmax ratio at 25 deg C
  • description: The ratio [PATO:0001470] of a leaf’s [PO:0025034] maximum [NCIT:C25564] electron transport [MESH:D004579] rate [PATO:0000161] to its maximum carboxylation [MOP:0000591] rate, reflecting the trade-off between photosynthetic gain and costs of excess energy dissipation, with the ratio affected by temperature, photosynthetically active radiation [ENVO:01001864] (light), and carbon dioxide [CHEBI:16526] concentration. These are both model parameters [STATO:0000034] derived from a photosynthesis [GO:0015979] A-ci response curve.;Ratio of photosynthetic electron transport capacity (Jmax) to maximum Rubisco activity (carboxylation capacity) (Vcmax). These are model-derived parameters, calculated from an A-ci response curve, reflecting, respectively the component of the curve where photosynthetic rates are limited by the light reactions versus low carbon dioxide concentrations.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 1 - 5 {dimensionless}

leaf_photosynthesis_Vcmax_per_area_25C

  • label: Leaf Vcmax per unit leaf area at 25 deg C (Vcmax25)
  • description: Maximum [NCIT:C25564] carboxylation [MOP:0000591] rate [PATO:0000161] by the Rubisco complex [GO:0048492] in plant leaves [PO:0025034], recorded per unit leaf [PO:0025034] area [PATO:0001323] and measured at 25 Celsius. The value is a model parameter [STATO:0000034] derived from the initial slope of a photosynthesis [GO:0015979] A-ci response curve, the component that is carbon dioxide [CHEBI:16526] limited and hence limited by Rubisco activity.;The apparent maximum rate of Rubisco (ribulose 1,5-bisphosphate carboxylase/oxygenase) activity carboxylase activity (carboxylation capacity), measured at 25 Celsius and recorded per unit leaf area. This is a model-derived value, calculated from the portion of an A-ci response curve (curve showing the photosynthetic response to increasing carbon dioxide concentrations) at low carbon dioxide concentrations. This is the portion of the curve where the photosynthetic rate is limited by the Rubisco activity.
  • type: numeric
  • units: umol/m2/s
  • allowable range: 10 - 300 umol/m2/s

ploidy

  • label: Chromosome ploidy
  • description: The count [NCIT:C25463] of copies of each chromosome [NCIT:C13202] present in a somatic cell [CL:0002371].;Chromosome ploidy, the number of copies of each chromosome present in a somatic cell.
  • type: numeric
  • units: {count}
  • allowable range: 1 - 4 {count}

leaf_epidermis_thickness

  • label: Leaf epidermis thickness
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] , in samples where the leaf lamina [PO:0020039] side (abaxial [PATO:0002046] versus adaxial [PATO:0002047]) is not specified.;Thickness of the epidermis, where the leaf lamina side is not specified.
  • type: numeric
  • units: um
  • allowable range: 1 - 100 um

leaf_palisade_cell_width

  • label: Palisade cell width
  • description: A plant cell morphology trait [TO:0002683] which is the width [PATO:0000921] of individual palisade mesophyll cells [PO:0006206].;Width of individual leaf palisade cells.
  • type: numeric
  • units: um
  • allowable range: 1 - 50 um

leaf_stomatal_hairs

  • label: Stomatal hairiness
  • description: A leaf stomatal complex frequency trait [TO:0000566] which is the pilosity [PATO:0000066] (hairiness) of the stomatal pores [PO:0008032].;Variable indicating if stomata are covered by hairs and, if hairy, the density of hairs present.
  • type: categorical
  • allowable values:
    • stomata_covered_with_dense_hairs: Stomata densely covered by hairs.
    • stomata_not_covered_by_hairs: Stomata not hair-covered.

leaf_total_vein_density

  • label: Length of all minor and major leaf lamina veins per unit area
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of the total length [PATO:0000122] of all leaf lamina veins [PO:0020138] to leaf lamina [PO_0020039] area [PATO:0001323].;The length of all minor and major leaf lamina veins per unit leaf area.
  • type: numeric
  • units: mm/mm2
  • allowable range: 0.01 - 20 mm/mm2

leaf_Cr_per_dry_mass

  • label: Leaf chromium (Cr) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chromium [CHEBI:28073] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of chromium in a leaf to leaf dry mass.
  • type: numeric
  • units: ppm
  • allowable range: 0.01 - 150 ppm

leaf_Co_per_dry_mass

  • label: Leaf cobalt (Co) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of cobalt [CHEBI:27638] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of cobalt in a leaf to leaf dry mass.
  • type: numeric
  • units: ppm
  • allowable range: 0.01 - 150 ppm

leaf_Se_per_dry_mass

  • label: Leaf selenium (Se) content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of selenium [CHEBI:27568] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of selenium in a leaf to leaf dry mass.
  • type: numeric
  • units: ppm
  • allowable range: 0.01 - 100 ppm

seed_protein_per_seed_dry_mass

  • label: Seed protein content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of protein [CHEBI:36080] in a seed [PO:0009010] to seed dry mass [TO:0000181].;The ratio of the mass of protein in a seed to seed dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 1e-05 - 1 g/g

plant_type_by_resource_use

  • label: Plant resource requirements and tolerance
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which categorises plants based on the environmental conditions in which they survive and thrive, including their tolerance of, sensitivity [NCIT:C25201] to, and requirement [NCIT:C25652] for certain resource availabilities [EnvThes:21497], including water [CHEBI:15377] availability and soil [ENVO:00001998] nutrient [CHEBI:33284] content and salinity [PATO:0085001].;Classification scheme of plant resource requirements and tolerances, categorising taxa based on their water, soil nutrient, and soil salinity requirements and tolerances.
  • type: categorical
  • allowable values:
    • hydrohalophyte: Tidal swamp or ‘mangrove’ and coastal lagoon elements and temperate-zone salt-marsh taxa are classed together.
    • phreatophyte: Deep-rooted plants—usually trees—obtaining water from a deep underground source that may or may not be saline.
    • psammophile: Sand-loving plants commonly found in littoral strand or inland sandfields.
    • weedy: This inexact term is meant to indicate a high degree of adaptability to sites, and proven tendency toward aggressive colonization of either dry or wet sites, or rarely, both. In general, weeds invade and colonize highly disturbed sites or areas.
    • xerohalophyte: Plants adapted to inland salt desert and saline habitats.
    • xerophyte: Drought-tolerant and drought-adapted plants.

plant_tolerance_salt

  • label: Plant salt tolerance strategy
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which categorises plants based on their salinity [PATO:0085001] tolerance [TO:0006001], reflecting the duration [PATO:0001309] and concentration [PATO:0000033] of salt [CHEBI:24866] exposure that a plant is able to tolerate.;Classification scheme of plant salt-tolerance categories, categorising taxa based on their tolerance of or sensitivity to salty soils, salt spray, or salt water.
  • type: categorical
  • allowable values:
    • glycophyte: Plants that cannot survive even one tenth the salt concentration found in seawater.
    • halophyte: Plants that complete their life cycle in a salty environment; many survive in seawater or even higher concentrations of salt.
    • halophyte_moderate: Plant that is moderately tolerant of soil salinity/sodicity.
    • hydrohalophyte: Tidal swamp or ‘mangrove’ and coastal lagoon elements and temperate-zone salt-marsh taxa are classed together.
    • salinity_tolerance_undefined: Plant’s ability to tolerate salt is unknown.
    • salt_spray_tolerant: Plant that is tolerant of salt spray (typically inhabiting near-coastal environments).
    • xerohalophyte: Plants adapted to inland salt desert and saline habitats.

plant_tolerance_soil_salinity

  • label: Plant soil salinity tolerance
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which is the maximum [NCIT:C25564] soil [ENVO:00001998] salinity [PATO:0085001], reported as soil conductivity [PATO:0001585], that a plant is able to tolerate.;Maximum salinity tolerated by a taxon, reported as the conductivity of the soil.
  • type: numeric
  • units: dS/m
  • allowable range: 0.1 - 100 dS/m

seed_oil_per_seed_dry_mass

  • label: Seed oil content per unit seed dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of lipids [CHEBI:18059] in a seed [PO:0009010] to seed dry mass [TO:0000181].;The ratio of the mass of lipids in a seed to seed dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 1e-04 - 1 g/g

bark_dry_mass_per_surface_area

  • label: Bark dry mass per unit bark surface area
  • description: The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding bark surface area [EnvThes:21564].;The ratio of bark dry mass to bark surface area.
  • type: numeric
  • units: g/cm2
  • allowable range: 0.01 - 1 g/cm2

wood_axial_parenchyma_fraction

  • label: Wood axial parenchyma fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the fraction of a cross-section [NCIT:C63795] of sapwood [PO:0004513] comprised of axial secondary xylem parenchyma [PO:0004533], calculated as ratio [PATO:0001470] of axial parenchyma cross-sectional area [PATO:0001323] to sapwood cross-sectional area.;The fraction of a cross-section of sapwood comprised of axial parenchyma. It is calculated as the ratio of total axial parenchyma cross-sectional area to sapwood cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.001 - 1 mm2/mm2

wood_conduit_fraction

  • label: Wood conduit fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the fraction of a cross-section [NCIT:C63795] of sapwood [PO:0004513] comprised of all water [CHEBI:15377] transport conduits (both tracheids [PO:0000301] and xylem vessels [PO:0025417]), calculated as the ratio [PATO:0001470] of total cross-sectional area [PATO:0001323] of conduits to sapwood cross-sectional area.;The fraction of a cross-section of sapwood comprised of all water transport conduits, including both xylem vessels and tracheids. It is calculated as the ratio of total conduit cross-sectional area to sapwood cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.01 - 1 mm2/mm2

wood_fibre_fraction

  • label: Wood fibre fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the fraction of a cross-section [NCIT:C63795] of sapwood [PO:0004513] comprised of plant fiber cells [PO:0025407], calculated as the ratio [PATO:0001470] of total fiber cell cross-sectional area [PATO:0001323] to sapwood cross-sectional area.;The fraction of a cross-section of sapwood comprised of fibres. It is calculated as the ratio of total fibre cross-sectional area to sapwood cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.01 - 1 mm2/mm2

wood_ray_parenchyma_fraction

  • label: Wood ray parenchyma fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the fraction of a cross-section [NCIT:C63795] of sapwood [PO:0004513] comprised of ray secondary xylem parenchyma cells [PO:0004534], calculated as the ratio [PATO:0001470] of total ray parenchyma cross-sectional area [PATO:0001323] to sapwood cross-sectional area.;The fraction of a cross-section of sapwood comprised of rays. It is calculated as the ratio of total ray cross-sectional area to sapwood cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.01 - 1 mm2/mm2

seed_germination_time

  • label: Seed germination time
  • description: The time interval (duration) [PATO:0001309] from when a seed [PO:0009010] is exposed to suitable [NCIT:C116358] environmental conditions [ENVO:01000203] or experimental conditions [XCO:0000000] for germinating [GO:0009845] until it germinates.;The time interval from when a seed is exposed to suitable conditions for germinating until it germinates.
  • type: numeric
  • units: d
  • allowable range: 0 - 1000 d

bark_thickness_index

  • label: Scaled bark thickness
  • description: A bark morphology trait which is the ratio [PATO:0001470] of two times bark [PO:0004518] thickness [PATO:0000915] to the stem [PO:0009047] diameter [PATO:0001334].;The ratio of two times a single bark thickness measure to stem diameter, to adjust for increasing bark thickness with increasing stem diameter.
  • type: numeric
  • units: mm/mm
  • allowable range: 0.001 - 1 mm/mm

stem_dry_matter_content

  • label: Stem dry matter content (SDMC)
  • description: A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of stem [PO:0009047] (or branch [PO:0025073]) dry [PATO:0001824] mass [PATO:0000125] to stem (or branch) fresh mass [APD:glossary/40044].;The ratio of stem (or branch) dry mass to stem (or branch) fresh mass; SDMC.
  • type: numeric
  • units: mg/g
  • allowable range: 100 - 1000 mg/g

stem_C_per_dry_mass

  • label: Stem carbon (C) content per unit stem dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a stem [PO:0009047] to stem dry mass.;The ratio of the mass of carbon in a stem to stem dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 100 - 1000 mg/g

leaf_mass_to_stem_mass_ratio

  • label: Leaf dry mass to stem dry mass ratio
  • description: A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of a plant’s total leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to its total stem [PO:0009047] and branch [PO:0025073] dry mass.;The ratio of whole plant leaf dry mass to whole plant stem dry mass.
  • type: numeric
  • units: g/g
  • allowable range: 0.01 - 10 g/g

diaspore_dry_mass

  • label: Diaspore dry mass
  • description: A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a seed [PO:0009010] and the seed dispersal [EnvThes:21042] structures [PATO:0000141].;Dry mass of a plant’s dispersal unit, including the seed and any dispersal appendages; synonymous with the term dispersule and often synonymous with the term disseminule.
  • type: numeric
  • units: mg
  • allowable range: 1e-05 - 1e+06 mg

root_C_per_dry_mass

  • label: Root carbon (C) content per unit root dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in roots [PO:0009005] to root dry mass.;The ratio of the mass of carbon in roots to root dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 50 - 750 mg/g

root_N_per_dry_mass

  • label: Root nitrogen (N) content per unit root dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in roots [PO:0009005] to root dry mass.;The ratio of the mass of nitrogen in roots to root dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.1 - 100 mg/g

root_delta13C

  • label: Root stable carbon isotope composition (delta13C)
  • description: Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of roots [PO:0009005].;Carbon stable isotope ratio, d13C, of roots.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -50 - 0 {delta}[ppth]

root_delta15N

  • label: Root stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of roots [PO:0009005].;Nitrogen stable isotope ratio, d15N, of roots.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

leaf_epidermal_cell_density_abaxial

  • label: Lower leaf side epidermal cell density
  • description: A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323].;Epidermal cell density on the lower side of the leaf.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 100 - 10000 {count}/mm2

leaf_epidermal_cell_density_adaxial

  • label: Upper leaf side epidermal cell density
  • description: A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323].;Epidermal cell density on the upper side of the leaf.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 100 - 10000 {count}/mm2

stem_vessel_non_lumen_fraction

  • label: Stem non-lumen fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the proportion of a cross-section of stem that is not comprised of lumen (empty space) and is calculated at 1 minus the ratio [PATO:0001470] of total xylem vessel [PO:0025417] and tracheid [PO:0000301] lumen [PO:0025117] cross-sectional [NCIT:C63795] area [PATO:0001323] to stem [PO:0009047] cross-sectional area.;The fraction of a stem cross-section not comprised of lumen (the empty space in xylem vessels and tracheids) and calculated as one minus the ratio of the total cross-sectional area comprised of xylem vessels and tracheids (lumen fraction) to the total stem cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.001 - 1 mm2/mm2

leaf_lamina_mass_per_area

  • label: Leaf lamina mass per area
  • description: A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to leaf [PO:0025034] area [PATO:0001323], measured on a piece of the leaf lamina [PO:0020039] located away from major leaf lamina veins [PO:0020138].;The ratio of leaf dry mass to leaf area, measured on a piece of the leaf lamina that does not include the petiole or major veins.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 500 g/m2

plant_physical_defence_structures

  • label: Physical defence structures
  • description: A plant structure morphology trait [TO:0000839] which indicates the plant structures [PO:0009011] used for defence [NCIT:C61556], including spine leaves [PO:0025173], spine stipules [PO:0025174], spine petiole [PO:0025613], thorns [PO:0025172], and prickles [PO:0025169].;The presence (or explicit absence) of specific physical defence structures on a plant, including spines, thorns, prickles, and sharp-tipped leaves.
  • type: categorical
  • allowable values:
    • absent: Plant that lacks a physical defence structure.
    • prickle: A sharp-tipped outgrowth from the epidermis, bark or other plant organ that is explicitly not a spine (modified leaf) or thorn (modified stem).
    • pungent_leaf_apex: A sharp-tipped leaf.
    • sharp_pointed_defence: Plant that has either spines, thorns, or prickles but the type of physical defence structure is not specified.
    • spine: A sharp-pointed structure, which is a modified leaf or leaf part, including petioles, midribs, secondary veins, leaflets, or stipules.
    • stinging_or_irritant_hairs: Trichomes that sting or irritate, by injecting a toxin.
    • thorn: A sharp-pointed, stiff, woody modified stem.

root_diameter

  • label: Root diameter
  • description: A root morphology trait [TO:0000043] which is the maximum [NCIT:C25564] root [PO:0009005] diameter [PATO:0001334] of a plant.;Maximum root diameter.
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1000 mm

root_distribution_coefficient

  • label: Root biomass depth distribution coefficient
  • description: A root system morphology trait [TO:0000985] which is a model parameter [STATO:0000034] indicating the distribution [PATO:0000060] of a plant’s roots [PO:0009005] with depth [PATO:0001595]. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter.;Root biomass depth distribution coefficient (‘B’ from Gale & Grigal 1987), where high values indicate that root biomass is allocated deeper in the soil. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.5 - 1 {dimensionless}

root_dry_matter_content

  • label: Root dry matter content (RDMC)
  • description: A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of root dry mass [TO:0000078] to root [PO:0009005] fresh mass [APD:glossary/40044].;The ratio of root dry mass to root fresh mass; root dry matter content, RDMC.
  • type: numeric
  • units: mg/g
  • allowable range: 10 - 1000 mg/g

root_specific_root_area

  • label: Root surface area per unit root dry mass (specific root area)
  • description: A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] surface area [EnvThes:21564] to root dry mass [TO:0000078].;The ratio of root surface area to root dry mass; specific root area.
  • type: numeric
  • units: mm2/mg
  • allowable range: 0.001 - 1 mm2/mg

root_fine_root_coarse_root_ratio

  • label: Fine root volume to coarse root volume ratio
  • description: A root system morphology trait [TO:0000985] which is the ratio [PATO:0001470] of the volume [PATO:0000918] of fine roots [EnvThes:20581] (those <0.5mm in diameter) to the volume of coarse roots [APD:glossary/40026] (those >0.5 mm in diameter).;The ratio of the volume of fine roots (<0.5mm diameter) to the volume of coarse roots (>0.5mm diameter).
  • type: numeric
  • units: mm3/mm3
  • allowable range: 0.05 - 50 mm3/mm3

leaf_hairs_immature_leaves

  • label: Immature leaf hairiness
  • description: A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of an adult phase leaf [PO:0006340] still in the vascular leaf expansion stage [PO:0001052].;Variable describing whether immature (still expanding) adult phase leaves on adult plants have hairs (trichomes).
  • type: categorical
  • allowable values:
    • glabrous: Immature (still expanding) leaves do not have hairs (trichomes).
    • hairy: Immature (still expanding) leaves have hairs (trichomes).

resprouting_capacity_stem_ratio

  • label: Post-fire to pre-fire stem ratio
  • description: A fire response trait which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stems [PO:0009047] per individual [EFO:0000542] following a fire [ENVO:01000786] to the count of stems per individual before the fire, recorded for taxa that have multiple stems emerging from the ground [SWEET:realmSoil/Ground].;Ratio of stem count post-fire to basal stem count (e.g. grass tillers) pre-fire at an individual or population level. This trait is appropriate to score for plants that have many stems from the base (shrubs, herbs, graminoids) where the number of stems is censused both before and after the fire. It is a continuous measure of resprouting strength conditioned on initial size.
  • type: numeric
  • units: {count}/{count}
  • allowable range: 0 - 10 {count}/{count}

flower_count_maximum

  • label: Maximum flower number
  • description: A floral organ morphology trait [TO:0000862] which is the maximum [NCIT:C25564] count [NCIT:C25463] of flowers [PO:0009046] on an individual [EFO:0000542].;Maximum flower number produced.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 1e+06 {count}

leaf_mesophyll_conductance_per_area

  • label: Leaf mesophyll conductance to carbon dioxide per unit leaf area (gm)
  • description: The diffusion [NCIT:C82333] rate [PATO:0000161] of carbon dioxide [CHEBI:16526] in plant leaves [PO:0025034] from intercellular spaces [PO:0025379] to chloroplasts [GO:0009507], recorded per unit leaf area [PATO:0001323].;The rate of carbon dioxide movement between sub-stomatal cavities (intracellular space) and chloroplasts, per unit leaf area
  • type: numeric
  • units: mol{CO2}/m2/s
  • allowable range: 0.01 - 2000 mol{CO2}/m2/s

leaf_mesophyll_conductance_per_mass

  • label: Leaf mesophyll conductance to carbon dioxide per unit leaf mass (gm)
  • description: The diffusion [NCIT:C82333] rate [PATO:0000161] of carbon dioxide [CHEBI:16526] in plant leaves [PO:0025034] from intercellular spaces [PO:0025379] to chloroplasts [GO:0009507], recorded per unit leaf dry mass [TO:0001014].;The rate of carbon dioxide movement between sub-stomatal cavities (intracellular space) and chloroplasts, per unit leaf mass
  • type: numeric
  • units: mmol{CO2}/g/s
  • allowable range: 0.01 - 10 mmol{CO2}/g/s

bark_modulus_of_elasticity

  • label: Bark modulus of elasticity
  • description: A bark [PO:0004518] physical quality [PATO:0002079] trait which is the bark’s modulus of elasticity [SWEET:propPressure/ModulusOfElasticity], its resistance to being deformed [SWEET:phenSolid/Deform] elastically [PATO:0001031] (non-permanently) when a force [PATO:0001035] is applied. It is calculated as the ratio of the force applied to bark per unit area (stress) to the bark’s deformation (bending).;A measure of stem stiffness and elasticity, which is how much the bark is deformed elastically (bends) under stress. It is calculated as the ratio of the force applied to bark per unit area (stress) to the bark’s deformation (bending). The trait is also called Young’s modulus for bark.
  • type: numeric
  • units: MPa
  • allowable range: 10 - 1e+05 MPa

modulus_of_rupture

  • label: Modulus of rupture
  • description: A secondary xylem [PO:0005848] physical quality [PATO:0002079] trait which is the xylem [PO:0005352] cell wall’s [GO:0009505] mechanical [MESH:D055595] bending strength [PATO:0001230], reported as the force [PATO:0001035] applied to a stem [PO:0009047] that causes it to stop bending and break (fail).;A measure of wood bending strength, which is the force per unit surface area applied to a length of bent stem (surface stress) that causes it to break. This is also the force required to rupture xylem vessels.
  • type: numeric
  • units: MPa
  • allowable range: 1 - 10000 MPa

stem_modulus_of_elasticity

  • label: Stem modulus of elasticity
  • description: A stem [PO:0009047] physical quality [PATO:0002079] trait which is the stem’s modulus of elasticity [SWEET:propPressure/ModulusOfElasticity], its resistance to being deformed [SWEET:phenSolid/Deform] elastically [PATO:0001031] (non-permanently) when a force [PATO:0001035] is applied. It is calculated as the ratio of the force applied to wood per unit area (stress) to the stem’s deformation (bending).;A measure of stem stiffness and elasticity, which is how much the stem is deformed elastically (bends) under stress. It is calculated as the ratio of the force applied to stem per unit area (stress) to the stem’s deformation (bending). This is the modulus of a compound tissue made up of bark and wood (or xylem) and potentially pith and could also be called structural modulus of elasticity. Trait is also called Young’s modulus for a stem.
  • type: numeric
  • units: MPa
  • allowable range: 10 - 1e+05 MPa

xylem_modulus_of_elasticity

  • label: Xylem modulus of elasticity
  • description: A secondary xylem [PO:0005848] physical quality [PATO:0002079] trait which is the xylem’s modulus of elasticity [SWEET:propPressure/ModulusOfElasticity], its resistance to being deformed [SWEET:phenSolid/Deform] elastically [PATO:0001031] (non-permanently) when a force [PATO:0001035] is applied.;A measure of wood stiffness and elasticity, which is how much wood (secondary xylem) is deformed elastically (bends) under stress. It is calculated as the ratio of the force applied to wood per unit area (stress) to the stem’s deformation (bending). This is the modulus of elasticity explicitly for xylem (wood). Trait is also called Young’s modulus for xylem.
  • type: numeric
  • units: MPa
  • allowable range: 10 - 1e+05 MPa

stem_vessel_wall_fraction

  • label: xylem vessel wall fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of xylem vessel [PO:0025417] cell wall [GO:0009505] cross-sectional [NCIT:C63795] area [PATO:0001323] to total xylem vessel cross-sectional area.;Fraction of xylem vessels comprised of cell wall, calculated as the ratio of xylem vessel cell wall cross-sectional area to total xylem vessel cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0.001 - 1 mm2/mm2

fruit_wall_thickness

  • label: Fruit wall thickness
  • description: A fruit morphology trait [TO:0002629] which is the thickness [PATO:0000915] of the entire pericarp [PO:0009084] (the exocarp [PO:0009085], mesocarp [PO:0009087], and endocarp [PO:0009086] combined), although it often refers just to the fleshy [PATO:0002351] components of the pericarp (most frequently the exocarp and mesocarp).;Fruit wall thickness of a mature fresh fruit. This term sometimes applies explicitly to the entire pericarp (exocarp + mesocarp + endocarp), but more frequently will have a functional meaning, referring to the outer fleshy layer which might be just some components of the peripcarp (most frequently the exocarp and mesocarp).
  • type: numeric
  • units: mm
  • allowable range: 0.01 - 1000 mm

leaf_chlorophyll_per_dry_mass

  • label: Leaf total chlorophyll content (chlorophyll A + B) per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content.;The ratio of the mass of chlorophyll in a leaf to leaf dry mass, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
  • type: numeric
  • units: mmol/kg
  • allowable range: 0.1 - 15 mmol/kg

leaf_work_to_tear

  • label: Leaf work to tear
  • description: A leaf lamina [PO:0025060] strength [PATO:0001230] trait which is a measure of a leaf’s [PO:0025034] tensile strength [SWEET:propPressure/TensileStrength; MESH:D013718]. It is the force [PATO:0001035] required to tear or rip [SWEET:phenSolid/Rip] a leaf lamina.;The force (work) required to tear/rip a leaf. This is a measure of leaf tensile strength.
  • type: numeric
  • units: N
  • allowable range: 0.01 - 50 N

stem_water_content_per_saturated_mass

  • label: Stem water content per unit saturated stem mass
  • description: A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a stem [PO:0009047] to stem fresh mass [APD:glossary/40044] in a stem that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].;The ratio of the mass of water in a stem (stem water content) to total stem mass, measured in a saturated stem. A saturated stem is one that has been hydrated to its maximal water holding capacity and is at full turgor.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 10 g/g

leaf_chlorophyll_index_modified_ND705

  • label: Modified chlorophyll absorption ratio index 705
  • description: A remote sensing [EnvThes:21762] index [NCIT:C25390] of chlorophyll [CHEBI:28966] absorption [SWEET:procWave/Absorption], which is a ratio [PATO:0001470] that considers the relative reflectance [EnvThes:21580] of different wavelengths off the plant canopy [ENVO:01001242], where a higher value indicates greater chlorophyll absorption. The formula is: (R750-R705)/(R750+R705+2(R445)).;A remote sensing index of chlorophyll absorption, which is a ratio that considers the relative reflectance of different wavelengths off the plant canopy, where a higher value indicates greater chlorophyll absorption. The formula is: (R750-R705)/(R750+R705+2(R445)).
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0.1 - 1 {dimensionless}

bark_photosynthetic_status

  • label: Bark photosynthesis
  • description: A photosynthetic trait [TO:0001015] which indicates whether a plant’s bark [PO:0004518] does or does not have a photosynthetic rate greater than zero.;Binary variable indicating whether bark is photosynthetic.
  • type: categorical
  • allowable values:
    • non_photosynthetic_bark: Plant’s bark is not photosynthetic.
    • photosynthetic_bark: Plant’s bark is photosynthetic.

stem_delta13C

  • label: Stem stable carbon isotope composition (delta13C)
  • description: Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of stems [PO:0009047].;Carbon stable isotope ratio, d13C, of stems.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -50 - 0 {delta}[ppth]

stem_delta15N

  • label: Stem stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of stems [PO:0009047].;Nitrogen stable isotope ratio, d15N, of stems.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

stem_mass_to_shoot_mass_ratio

  • label: Stem dry mass to vegetative shoot dry mass ratio (support fraction)
  • description: A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of stem [PO:0009047] and branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to vegetative shoot system [PO:0025607] dry mass.;The ratio of whole plant stem dry mass to whole plant vegetative shoot dry mass. That is, the proportion of aboveground vegetative biomass that is comprised of stems and branches, rather than leaves; support fraction.
  • type: numeric
  • units: g/g
  • allowable range: 0 - 1 g/g

leaf_chlorophyll_A_per_dry_mass

  • label: Leaf chlorophyll A content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of chlorophyll a in a leaf to leaf dry mass.
  • type: numeric
  • units: mmol/kg
  • allowable range: 0.1 - 10 mmol/kg

leaf_chlorophyll_B_per_dry_mass

  • label: Leaf chlorophyll B content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of chlorophyll b in a leaf to leaf dry mass.
  • type: numeric
  • units: mmol/kg
  • allowable range: 0.01 - 10 mmol/kg

leaf_epidermal_cell_density_both_sides

  • label: Average leaf epidermal cell density
  • description: A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] to leaf lamina [PO:0020039] area [PATO_0001323], averaged across the abaxial [PATO:0002046] (lower) and adaxial [PATO:0002047] (upper) sides of the leaf lamina.;Epidermal cell density averaged across the upper and lower sides of the leaf.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 100 - 10000 {count}/mm2

leaf_insoluble_protein_per_area

  • label: Leaf insoluble protein content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] insoluble [PATO:0001538] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of water insoluble protein in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 20 g/m2

leaf_palisade_cell_length

  • label: Palisade cell length
  • description: A plant cell morphology trait [TO:0002683] which is the length [PATO:0000122] of individual palisade mesophyll cells [PO:0006206].;Length of individual leaf palisade cells.
  • type: numeric
  • units: um
  • allowable range: 10 - 1000 um

leaf_palisade_layer_number

  • label: Number of layers of palisade cells
  • description: A portion of plant tissue morphology trait [TO:0000843] which is the count [NCIT:C25463] of cell layers [UBERON:0000119] of palisade mesophyll cells [PO:0006206].;The number of layers of palisade cells in a leaf.
  • type: numeric
  • units: {count}
  • allowable range: 1 - 10 {count}

leaf_starch_per_area

  • label: Leaf starch content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of total starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of total starch in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 50 g/m2

water_potential_12percent_lost_conductivity

  • label: Stem xylem pressure, 12% lost conductivity (P12)
  • description: The tension [PATO:0002284] (water pressure) (water potential [EnvThes:22311]) in stem [PO:0009047] xylem vessels [PO:0025417] at which the hydraulic conductivity [EnvThes:22133] has declined by 12% from the mean maximum [NCIT:C25564] rate [PATO:0000161].;A measure of stem hydraulic vulnerability, which is the stem xylem water pressure value at which stem hydraulic conductivity has declined by 12% from the mean maximum rate of hydraulic conductivity.
  • type: numeric
  • units: MPa
  • allowable range: -20 - 0 MPa

leaf_xylem_delta15N

  • label: Leaf xylem stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from leaf [PO:0025034] xylem vessels [PO:0025417].;Nitrogen stable isotope ratio, d15N, of xylem sap from leaf xylem vessels.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

root_xylem_delta15N

  • label: Root xylem stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from root [PO:0009005] xylem vessels [PO:0025417].;Nitrogen stable isotope ratio, d15N, of xylem sap from root xylem vessels.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

pollination_system

  • label: Pollination system
  • description: A classification [NCIT:C25161] based on explicit field studies of pollination [GO:0009856] in the taxon’s [NCIT:C40098] native range (preferred) or else a combination of a taxon’s flower morphology traits [TO:0000499] (that suggest specific [NCIT:C38024] taxa (biotic vectors) or abiotic mechanisms [NCIT:C25343] that most likely pollinate [GO:0009856] a flower [PO:0009046]) and the pollinators actually observed visiting the taxon’s flower.;For a given taxon, the biotic vectors (including explicit taxa) or abiotic mechanisms that are most likely to pollinate its flowers, based on explicit field studies of pollination in the taxon’s native range (preferred) or else a combination of possible pollinators observed visiting a taxon’s flowers and the taxon’s flower morphology.
  • type: categorical
  • allowable values:
    • abiotic: Observations indicate pollination occurs through wind, water, or collision of free-floating particles.
    • animal: Flowers are observed to be animal-pollinated.
    • ballistic: Pollen is observed to be launched away from the plant by explosion.
    • bat: Flowers are observed to be bat-pollinated.
    • bee: Flowers are observed to be bee-pollinated.
    • beetle: Flowers are observed to be beetle-pollinated.
    • biotic_specialised: Taxa depend upon a specialist pollinator for pollen transfer.
    • biotic_unspecialised: Taxa are pollinated by a biotic agent, but do not have a specialist pollinator.
    • bird: Flowers are observed to be bird-pollinated.
    • butterfly: Flowers are observed to be butterfly-pollinated.
    • fly: Flowers are observed to be fly-pollinated.
    • insect: Flowers are observed to be insect-pollinated, type of insect not specified.
    • mammal: Flowers are observed to be mammal-pollinated.
    • moth: Flowers are observed to be moth-pollinated.
    • self: Flowers are self-pollinated.
    • thrip: Flowers are observed to be thrip-pollinated.
    • vertebrate: Flowers are observed to be vertebrate-pollinated (bats, birds, mammals or lizards).
    • wasp: Flowers are observed to be wasp-pollinated.
    • water: Flowers are observed to be water-pollinated.
    • wind: Flowers are observed to be wind-pollinated.

bud_length

  • label: Bud length
  • description: A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud.;Linear dimension from the base to the apex of a bud, even if this is not the longest dimension.
  • type: numeric
  • units: cm
  • allowable range: 0.01 - 200 cm

bud_width

  • label: Bud width
  • description: A floral organ morphology trait [TO:0000862] which is the width [PATO:0000921] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud. Width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the bud length [PATO:0000122];Greatest linear width dimension of a bud, sometimes referred to as ‘diameter’ in buds where the width and height are identical; orthogonal to the bud length, the base-to-apex dimension.
  • type: numeric
  • units: cm
  • allowable range: 0.01 - 200 cm

buds_per_inflorescence

  • label: Buds per inflorescence
  • description: A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of buds [PO:0000055] in an inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system [PO:0025082].;The count of buds in an inflorescence, where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system.
  • type: numeric
  • units: {count}/{count}
  • allowable range: 1 - 1000 {count}/{count}

leaf_K_per_area

  • label: Leaf potassium (K) content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of potassium in a leaf to leaf area.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 10 g/m2

root_specific_taproot_length

  • label: Specific tap root length (STRL)
  • description: A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of primary root [PO:0020127] length [PATO:0000122] to primary root dry mass [TO:0000078];The ratio of taproot length (primary root length) to taproot dry mass; STRL.
  • type: numeric
  • units: m/g
  • allowable range: 0.5 - 5000 m/g

plant_alternative_energy_and_nutrient_acquisition_strategy

  • label: Plant alternative energy and nutrient acquisition strategies
  • description: A whole plant [PO:0000003] life history [ECOCORE:00000018] trait which are alternative mechanisms [NCIT:C25343] plants use to obtain [NCIT:C52349] or produce [SIO:001106] energy [PATO:0001021] (carbohydrates [CHEBI:16646]) or nutrients [CHEBI:33284].;Trait indicating the strategies used by a plant to obtain or produce energy (carbohydrates) or nutrients, including photosynthesis and various saprophytic and carnivorous energy and nutrient acquisition mechanisms.
  • type: categorical
  • allowable values:
    • carnivorous: Plant that acquires some or most of its nutrients from animals or protozoa.
    • nutrient_mining: Plant that uses a specialised root structure, such as cluster roots, to mine nutrients inaccessible to other plant taxa.
    • saprophyte: Plant that acquires nutrients from decaying biomass.

stem_count

  • label: Stem count
  • description: A shoot system morphology trait [TO:0000077] which is the count [NCIT:C25463] of stems [PO:0009047] on multi-stemmed plants [PO:0000003].;Count of stems on multi-stemmed plants.
  • type: numeric
  • units: {count}
  • allowable range: 0 - 5 {count}

post_fire_flowering

  • label: Post-fire flowering
  • description: A fire response trait which is a taxon’s [NCIT:C40098] flowering [PO:0007016] response [NCIT:C25755] to fire [ENVO:01000786], indicating if the magnitude [NCIT_C49158] of flowers [PO:0009046] increases, decreases or remains unchanged following a fire.;Indication of whether the magnitude of flowers produced by a taxon increases, decreases or remains unchanged following a fire.
  • type: categorical
  • allowable values:
    • fire_dependent_flowering: Plants predominantly flower after a fire, usually within the first 1-4 growing seasons following the fire.
    • fire_enhanced_flowering: Flowering is enhanced after a fire, with plants flowering more prolifically for the first 1-4 growing seasons following the fire.
    • fire_independent_flowering: Flowering is independent of fire, with the magnitude of flowers produced by plants unchanged following a fire.
    • fire_suppressed_flowering: Flowering is suppressed or absent after a fire, with plants producing fewer flowers for at least the first 1-4 growing seasons following the fire.

sprout_depth

  • label: Sprout depth
  • description: The depth [PATO:0001595] below the ground [SWEET:realmSoil/Ground] surface or height [PATO:0000119] above the ground surface of vegetative meristematic tissue [PO:0006079] (buds [PO:0000055]) that resprouts [Cerrado:fire#PlantResprout] following a fire [ENVO:01000786].;Depth below the soil surface (positive number) or height above ground (negative number) from which buds (meristematic tissue) resprout following a disturbance, such as a fire.
  • type: numeric
  • units: mm
  • allowable range: -100 - 100 mm

leaf_secondary_vein_angle

  • label: Leaf secondary vein angle
  • description: A vascular tissue morphology trait [TO:0000470] which is the angle [PATO:0002326] between the primary leaf vein [PO:0025413] and the secondary leaf vein [PO:0020140] in a leaf lamina [PO:0020039].;Angle between the primary vein (midrib) and a secondary vein in a leaf lamina.
  • type: numeric
  • units: deg
  • allowable range: 0 - 90 deg

leaf_vein_frequency

  • label: Leaf vein frequency
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of all leaf lamina veins [PO:0020138] intersecting the perimeter [PATO:0001711] of a square of the leaf lamina divided by the length of the perimeter of the square.;A measure of vein density, which is the number of veins intersecting the perimeter of a square of the leaf lamina divided by the length of the perimeter of the square.
  • type: numeric
  • units: {count}/mm
  • allowable range: 0.01 - 20 {count}/mm

leaf_delta18O

  • label: Leaf stable oxygen isotope composition (delta18O)
  • description: Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of leaves [PO:0025034].;Oxygen stable isotope ratio, d18O, of leaves.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -10 - 50 {delta}[ppth]

leaf_carotenoid_per_dry_mass

  • label: Leaf carotenoid content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of carotenoids in a leaf to leaf dry mass.
  • type: numeric
  • units: mmol/kg
  • allowable range: 0.1 - 10 mmol/kg

leaf_rubisco_N_per_total_leaf_N

  • label: Leaf rubisco nitrogen (N) content per unit leaf N content
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in rubisco [MESH:D012273] in a leaf [PO:0025034] to the mass of nitrogen in a leaf.;The ratio of the mass of nitrogen in rubisco in a leaf to the mass of nitrogen in a leaf.
  • type: numeric
  • units: g/g
  • allowable range: 0.05 - 0.5 g/g

leaf_rubisco_per_leaf_dry_mass

  • label: Leaf rubisco content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of rubisco [MESH:D012273] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].;The ratio of the mass of rubisco in a leaf to leaf dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 5 - 50 mg/g

leaf_thylakoid_N_per_total_leaf_N

  • label: Leaf thylakoid protein nitrogen (N) content per unit leaf N content
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in thylakoid [GO:0009579] proteins [CHEBI:36080] in a leaf [PO:0025034] to the mass of nitrogen in a leaf.;The ratio of the mass of nitrogen in thylakoid proteins in a leaf to the mass of nitrogen in a leaf.
  • type: numeric
  • units: g/g
  • allowable range: 0.05 - 0.3 g/g

leaf_total_non-structural_carbohydrates_per_area

  • label: Leaf total non-structural carbohydrate content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf area [PATO:0001323], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505].;The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf area, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
  • type: numeric
  • units: g/m2
  • allowable range: 0.1 - 50 g/m2

leaf_total_non-structural_carbohydrates_per_mass

  • label: Leaf total non-structural carbohydrate content per unit leaf dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505].;The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf dry mass, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
  • type: numeric
  • units: mg/g
  • allowable range: 10 - 200 mg/g

reproductive_light_environment_index

  • label: Canopy light environment required for reproduction
  • description: A plant phenological [TO:0000933] life history [ECOCORE:00000018] trait which is an index [NCIT:C25390] that captures the level of photosynthetically active radiation [ENVO:01001864] required by plants for reproduction [GO:0000003].;The minimum light environment required by plants for reproduction, scaled from 0 to 8.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 8 {dimensionless}

establishment_light_environment_index

  • label: Canopy light environment required for seedling establishment
  • description: A plant phenological [TO:0000933] life history [ECOCORE:00000018] trait which is an index [NCIT:C25390] that captures the level of photosynthetically active radiation [ENVO:01001864] required by plants for seed germination [GO:0009845] and seedling establishment [EnvThes:20410].;The minimum light environment required by seedlings for establishment, scaled from 0 to 8.
  • type: numeric
  • units: {dimensionless}
  • allowable range: 0 - 8 {dimensionless}

cotyledon_function

  • label: Cotyledon function
  • description: A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] have a photosynthesis [GO:0015979] or energy [PATO:0001021] storage [OBI:0302893] function [NCIT:C25518].;Binary variable distinguishing between cotyledons that function as leaves and can photosynthesise versus ones that function as energy reserve organs.
  • type: categorical
  • allowable values:
    • foliaceous: Cotyledons are photosynthetic and function as leaves.
    • reserve: Cotyledons function as energy reserve organs.

cotyledon_position

  • label: Cotyledon position at germination
  • description: A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] emerge from the seed coat [PO:0009088] at germination [GO:0009845].;Binary variable distinguishing between seedlings where the cotyledon remains within the seed coat versus emerges from the seed coat at germination.
  • type: categorical
  • allowable values:
    • cryptocotylar: A type of seed germination in which the cotyledons remain within the seed coat at germination.
    • phanerocotylar: A type of seed germination in which the cotyledons emerge from the seed coat.

seedling_germination_location

  • label: Seedling germination location
  • description: A plant embryo cotyledon morphology trait [TO:0000775] which is the position [PATO:0000140] of seed germination [GO:0009845] relative to the soil surface.;Binary variable distinguishing between seedlings that germinate above versus below ground.
  • type: categorical
  • allowable values:
    • epigeal: Germinant with one or more cotyledons emerging aboveground.
    • hypogeal: Germinant with all cotyledons remaining belowground.

flowering_cues

  • label: Environmental flowering cues
  • description: Unpredictable, irregular environmental conditions [ENVO_01000203] that are linked to a plant [PO:0000003] producing flowers [PO:0009046].;A plants’ flowering response to unpredictable, irregular environmental cues.
  • type: categorical
  • allowable values:
    • fire: Plants increase flowering in response to fires.
    • floods: Plants increase flowering in response to floods.
    • rain: Plants increase flowering in response to rain, but the time of year is not specified.
    • rain_all_year: Plants increase flowering in response to rain at any time of the year.
    • rain_autumn: Plants increase flowering in response to autumn rains.
    • rain_obligate: Plants only flower following rain events.
    • rain_spring: Plants increase flowering in response to spring rains.
    • rain_summer: Plants increase flowering in response to summer rains.
    • rain_winter: Plants increase flowering in response to winter rains.

root_P_per_dry_mass

  • label: Root phosphorus (P) content per unit root dry mass
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in roots [PO:0009005] to root dry mass.;The ratio of the mass of phosphorus in roots to root dry mass.
  • type: numeric
  • units: mg/g
  • allowable range: 0.01 - 100 mg/g

branch_terminal_twig_cross_sectional_area

  • label: Terminal twig cross-sectional area
  • description: A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of the most distal [PATO:0001234] branch [PO:0025073].;Cross-sectional area of the terminal twig, measured just beyond the final branching node.
  • type: numeric
  • units: mm2
  • allowable range: 0.05 - 50 mm2

seed_dry_mass_reserve

  • label: Seed embryo and endosperm dry mass
  • description: A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of the components of a seed [PO:0009010] that provide energy [PATO:0001021] to a seedling [EnvThes:20698], generally the embryo [PO:0009009] and, if present, the endosperm [PO:0009089].;Dry mass of a seed’s embryo and endosperm (if present), the components of a seed that provide energy to a seedling. These tissues are distinct from a seed’s dispersal structures or protective tissues.
  • type: numeric
  • units: mg
  • allowable range: 1e-05 - 1e+05 mg

branch_terminal_twig_length

  • label: Terminal twig length
  • description: A stem morphology trait [TO:0000361] which is the length [PATO:0000122] of the most distal [PATO:0001234] branch [PO:0025073].;Length of the terminal twig, the length of stem beyond the final branching node.
  • type: numeric
  • units: mm
  • allowable range: 1 - 1000 mm

leaf_vessel_density

  • label: Leaf xylem vessel density
  • description: A vascular tissue morphology trait [TO:0000470] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of xylem vessels [PO:0025417] in a leaf [PO:0025034] cross-section [NCIT:C63795] to leaf cross-sectional area [PATO:0001323].;The ratio of the count of xylem vessels in a leaf cross-section to leaf cross-sectional area.
  • type: numeric
  • units: {count}/mm2
  • allowable range: 1 - 50000 {count}/mm2

leaf_vessel_diameter

  • label: Leaf xylem vessel diameter
  • description: A vascular tissue morphology trait [TO:0000470] which is the diameter [PATO:0001334] of xylem vessels [PO:0025417] in a leaf [PO:0025034].;Diameter of xylem vessels in a leaf.
  • type: numeric
  • units: um
  • allowable range: 1 - 250 um

plant_tolerance_snow

  • label: Plant snow tolerance
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which is the duration [PATO:0001309] that a plant [PO:0000003] is able to tolerate being covered by snow [ENVO:01000406].;Categorical groupings, indicating the length of time a taxon is able to tolerate being under snow cover.
  • type: categorical
  • allowable values:
    • days: Taxa can tolerate being covered by snow for days (typically less than 1 week).
    • intolerant: Taxa is frost tender and known to be killed by any snow.
    • months: Taxa can tolerate being covered by snow for months.
    • not_applicable: Taxa is virtually never confronted with snow in nature and the response is unknown.
    • weeks: Taxa can tolerate being covered by snow for weeks (typically less than 1 month).

plant_tolerance_water_logged_soils

  • label: Plant water-logging tolerance
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which is the duration [PATO:0001309] that a plant [PO:0000003] is able to tolerate water-logged soils, an indication of a plant’s sensitivity [NCIT:C25201] to flood [ENVO:01000710] conditions. Soil [ENVO:00001998] is considered water-logged if it is saturated [NCIT:C61427] with water [CHEBI:15377], limiting the plant root’s [PO:0009005] ability to respire [GO:0045333] due to a lack of soil oxygen [CHEBI:15379].;Categorical groupings, indicating the length of time a taxon is able to tolerate water-logged soils.
  • type: categorical
  • allowable values:
    • 1-6_months: Taxa able to tolerate 1-6 months of waterlogging.
    • aquatic: Taxa truly aquatic and able to be permanently inundated.
    • greater_than_6_months: Taxa able to tolerate more than 6 months of waterlogging.
    • less_than_1_month: Taxa able to tolerate less than 1 month of waterlogging.
    • not_applicable: Taxa is not typically confronted with waterlogging and its response is unknown.

resprouting_capacity_non_fire_disturbance

  • label: Plant vegetative response to disturbances other than fire
  • description: A whole plant [PO:0000003] life history [ECOCORE:00000018] trait which is the ability [NCIT:C78209] of individual [EFO:0000542] plants to regenerate [GO:0031099] vegetatively (resprout [Cerrado:fire#PlantResprout]) from meristematic tissue [PO:0006079], buds [PO:0000055]) after a disturbance event [EnvThes:20373] other than a fire [ENVO:01000786] that causes 100% defoliation.;A plant’s ability to regenerate vegetatively (resprout) from a disturbance other than fire that causes 100% defoliation.
  • type: categorical
  • allowable values:
    • resprouts_after_cyclone: Plants are able to resprout following 100% leaf loss in a cyclone.
    • resprouts_after_drought: Plants are able to resprout following 100% leaf loss during a drought.
    • resprouts_non_fire_disturbance: Plants that are known to be fire-killed, but have the ability to resprout following a disturbance other than fire that causes 100% leaf loss.

plant_tolerance_inundation

  • label: Plant inundation tolerance
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which is the duration [PATO:0001309] that a plant [PO:0000003] is able to tolerate inundation [SWEET:phenSystem/Inundation], an indication of a plant’s sensitivity [NCIT:C25201] to flood [ENVO:01000710] conditions.;Categorical groupings, indicating the length of time a taxon is able to tolerate being under water.
  • type: categorical
  • allowable values:
    • 1-6_months: Taxa able to tolerate being inundated for 1-6 months.
    • aquatic: Taxa truly aquatic and able to be permanently inundated.
    • greater_than_6_months: Taxa able to tolerate being inundated for more than 6 months.
    • less_than_1_month: Taxa able to tolerate being inundated for less than 1 month.
    • not_applicable: Taxa is not typically confronted with being inundated and its response is unknown.

plant_tolerance_calcicole

  • label: Plant calcium sensitivity
  • description: An environmental [ENVO:01000203] tolerance [NCIT:C43414] trait which is a plant’s [PO:0000003] calcium sensitivity [TO:0000006], corresponding to its sensitivity [NCIT:C25201] to high soil [ENVO:00001998] pH [NCIT:C45997].;Classification scheme of plant soil calcium tolerance, corresponding to its sensitivity to high soil pH. Plants are categorised as either calcifuge (intolerant of basic soils) or calcicole (tolerant of basic soils, such as calcareous sands and limestone derived soils).
  • type: categorical
  • allowable values:
    • calcicole: Plants are tolerant of basic soils, such as calcareous sands and limestone derived soils.
    • calcifuge: Plants are intolerant of basic soils. (Synonyms, acidophilous)

fire_exposure_level

  • label: Fire exposure level
  • description: A fire response trait indicating small-scale variation in plant [PO:0000003] habitat [ENVO:01000739] or growing season that protects the plant’s shoot systems [PO:0009006] or root system [PO:0025025] from exposure [NCIT:C17941] to fire [ENVO:01000786] or being damaged (scorched) by fire, when a fire occurs nearby.;Trait indicating habitat (or microhabitat) variation that protects the shoot or root system of some taxa from being exposed to fire and scorched by the fire, even when a fire burns nearby.
  • type: categorical
  • allowable values:
    • aquatic_taxon: Plants rarely or never experience fires because they are aquatic.
    • fire_avoidance_among_rocks: Plants rarely experience fires because their habitat is restricted to rocky outcrops or rock pavement.
    • fire_avoidance_in_swamp: Plants rarely experience fires because their habitat is usually saturated, but populations can be catastrophically impacted during severe droughts when the swamps dry out.
    • fire_avoidance_near_water: Plants rarely or never experience fires because their habitat is close to a body of water.
    • seasonal_fire_avoidance: Plants avoid fire by only have above-ground parts during seasons when fire is rare or absent, but populations can be catastrophically impacted if fire occurs during an atypical season.

bark_morphology_eucalyptus

  • label: Bark morphology, Eucalyptus
  • description: A bark morphology trait which is the texture [PATO:0000150] of the bark [PO:0004518] of Eucalyptus taxa.;Broad categories that describe the texture of Eucalyptus bark.
  • type: categorical
  • allowable values:
    • eucalypt_box: Eucalyptus, Corymbia or Angophora taxa with “Box” type bark.
    • eucalypt_gum: Eucalyptus, Corymbia or Angophora taxa with “Gum” type bark.
    • eucalypt_ironbark: Eucalyptus, Corymbia or Angophora taxa with “Ironbark” type bark.
    • eucalypt_peppermint: Eucalyptus, Corymbia or Angophora taxa with “Peppermint” type bark.
    • eucalypt_ribbonbark: Eucalyptus, Corymbia or Angophora taxa with “Gum” type bark that sheds in ribbons.
    • eucalypt_stocking: Eucalyptus, Corymbia or Angophora taxa with “Gum” type bark above a rough barked stocking.
    • eucalypt_stringybark: Eucalyptus, Corymbia or Angophora taxa with “Stringy” type bark.

plant_spinescence

  • label: Plant spinescence
  • description: A plant structure morphology trait [TO:0000839] which summarises the extent of physical structures a plant uses for defence [NCIT:C61556], including the size [PATO:0000117], strength [PATO:0001230], and density [PATO:0001019] of spine leaves [PO:0025173], spine stipules [PO:0025174], thorns [PO:0025172], and/or prickles [PO:0025169] and the pain [HP:0012531] inflicted by these structures.;Degree to which a plant is defended by spines, thorns and/or prickles, as defined by Pérez-Harguindeguy et al. 2013.
  • type: categorical
  • allowable values:
    • high_density_hard_spines_to_100mm: Intermediate or high density of hard, sharp spines >100mm long; plant is dangerous to careless large mammals, including humans. (level 5)
    • high_density_hard_spines_to_20mm: Intermediate or high density of hard, sharp spines >20mm long; plant may cause significant wounds when hit carelessly. (level 4)
    • high_density_hard_spines_to_5mm: Intermediate or high density of hard, sharp spines >5mm long; plant causes strong pain when hit carelessly. (level 3)
    • high_density_soft_spines: High density of soft spines, intermediate density of spines of intermediate hardness, or low density of hard, sharp spines >5mm long; plant causes actual pain when hit carelessly. (level 2)
    • low_density_soft_spines: Low or very local density of soft spines <5mm long; plant may sting or prickle when hit carelessly but not impart strong pain. (level 1)
    • no_spines: No spines. (level 0)

seed_volume

  • label: Seed volume
  • description: A seed morphology trait [TO:0000184] which is the volume [PATO:0000918] of a seed [PO:0009010].;Volume of a seed.
  • type: numeric
  • units: mm3
  • allowable range: 0.001 - 1e+05 mm3

cotyledon_hairs

  • label: Cotyledon hairiness
  • description: A plant embryo cotyledon morphology trait [TO:0000775] which is the pilosity [PATO:0000066] of the cotyledons [PO:0020030].;Binary variable distinguishing between glabrous versus hairy cotyledons.
  • type: categorical
  • allowable values:
    • glabrous: Cotyledons lack hairs.
    • hairy: Cotyledons have hairs.

embryo_colour

  • label: Embryo colour
  • description: A plant embryo cotyledon morphology trait [TO:0000775] which is the colour [PATO:0000014] of the plant embryo cotyledon [PO:0025470].;Binary variable distinguishing between embryos that are green versus colourless.
  • type: categorical
  • allowable values:
    • colourless: Colourless embryo.
    • green: Green embryo.

seedling_first_node_leaf_count

  • label: Seedling first node leaf count
  • description: A plant embryo true leaf formation stage [PO:0001095] morphology trait which is the leaf count [NCIT:C25463] at the first stem node [PO:0020141] above the epicotyl [PO:0020035].;Binary variable distinguishing between seedlings where the leaves at the first node are single versus paired.
  • type: categorical
  • allowable values:
    • paired: There is a pair of leaves at the first stem node above the epicotyl.
    • single: There is a single leaf at the first stem node above the epicotyl.

seedling_first_node_leaf_type

  • label: Seedling first true leaf type
  • description: A plant embryo true leaf formation stage [PO:0001095] morphology trait which is whether the leaf [PO:0025034] at the first stem node [PO:0020141] above the epicotyl [PO:0020035] in a true leaf or a scale leaf.;Binary variable distinguishing between seedlings where the first leaf is scale-like (cataphyll) versus leaf-like.
  • type: categorical
  • allowable values:
    • leaf-like: The leaf at the first stem node above the epicotyl is leaf-like (broad).
    • scale-like: The leaf at the first stem node above the epicotyl is scale-like, also referred to as a cataphyll.

seedling_hypocotyl_hairs

  • label: Hypocotyl hairiness
  • description: A seedling hypocotyl morphology trait [TO:0000769] which is the pilosity [PATO:0000066] of the hypocotyl [PO:0020100].;Binary variable distinguishing between glabrous versus hairy hypocotyls, where the hypocotyl is the embryonic axis to which the cotyledons are attached.
  • type: categorical
  • allowable values:
    • glabrous: Hypocotyl lacks hairs.
    • hairy: Hypocotyl has hairs.

bark_water_content_per_saturated_mass

  • label: Bark water content per unit saturated bark mass
  • description: A bark mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] to the fresh mass [APD:glossary/40044] of bark that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].;The ratio of the mass of water in bark (bark water content) to total bark mass, measured in saturated bark. Saturated bark is bark that has been hydrated to its maximal water holding capacity and is at full turgor.
  • type: numeric
  • units: g/g
  • allowable range: 0.1 - 10 g/g

leaf_carotenoid_per_area

  • label: Leaf carotenoid content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of carotenoids in a leaf to leaf area.
  • type: numeric
  • units: umol/m2
  • allowable range: 10 - 1000 umol/m2

leaf_chlorophyll_A_per_area

  • label: Leaf chlorophyll A content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of chlorophyll a in a leaf to leaf area.
  • type: numeric
  • units: umol/m2
  • allowable range: 50 - 2000 umol/m2

leaf_chlorophyll_B_per_area

  • label: Leaf chlorophyll B content per unit leaf area
  • description: The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf area [PATO:0001323].;The ratio of the mass of chlorophyll b in a leaf to leaf area.
  • type: numeric
  • units: umol/m2
  • allowable range: 10 - 1000 umol/m2

leaf_absorption

  • label: Leaf visible light absorption
  • description: A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is absorbed [SWEET:procWave/Absorption] by a leaf.;Proportion of incoming visible light (between 400-700 nm) that is absorbed by a leaf.
  • type: numeric
  • units: umol/umol
  • allowable range: 0.01 - 1 umol/umol

leaf_transmission

  • label: Leaf visible light transmission
  • description: A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is transmitted [SWEET:procWave/Transmission] through a leaf.;Proportion of incoming visible light (between 400-700 nm) that is transmitted through a leaf.
  • type: numeric
  • units: umol/umol
  • allowable range: 0.001 - 1 umol/umol

bark_delta13C

  • label: Bark stable carbon isotope composition (delta13C)
  • description: Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of bark [PO:0004518].;Carbon stable isotope ratio, d13C, of bark.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -50 - 0 {delta}[ppth]

bark_delta15N

  • label: Bark stable nitrogen isotope composition (delta15N)
  • description: Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of bark [PO:0004518].;Nitrogen stable isotope ratio, d15N, of bark.
  • type: numeric
  • units: {delta}[ppth]
  • allowable range: -25 - 75 {delta}[ppth]

life_history_ephemeral_class

  • label: Ephemeral life history class
  • description: A whole plant [PO:0000003] life history [ECOCORE:00000018] which is, for taxa [NCIT_C40098] that display an ephemeral life history, the environmental conditions [ENVO_01000203] associated with seed germination [GO:0009845] and seedling establishment [EnvThes:20410].;For taxa that display an ephemeral life history, the environmental conditions associated with seed germination and seedling establishment.
  • type: categorical
  • allowable values:
    • disturbance_ephemeral: Ephemeral plant the germinates and establishes following disturbance, but the type of disturbance is not specified.
    • fire_ephemeral: Ephemeral plant that germinates and establishes following fire, where it is unknown if the plant is an obligate or facultative fire ephemeral.
    • fire_ephemeral_facultative: Ephemeral plant that increases its germination and establishment following fire, but can also germinate and establish without fire.
    • fire_ephemeral_obligate: Ephemeral plant that only germinates and establishes following fire.
    • rain_ephemeral: Ephemeral plant that germinates and establishes following rain or flood events.

wood_tracheid_fraction

  • label: Wood tracheid fraction
  • description: A vascular tissue morphology trait [TO:0000470] which is the fraction of a cross-section [NCIT:C63795] of sapwood [PO:0004513] comprised of tracheids [PO:0000301], calculated as the ratio [PATO:0001470] of total tracheid cross-sectional area [PATO:0001323] to sapwood cross-sectional area.;The fraction of a cross-section of sapwood comprised of tracheids. It is calculated as the ratio of total tracheid cross-sectional area to sapwood cross-sectional area.
  • type: numeric
  • units: mm2/mm2
  • allowable range: 0 - 1 mm2/mm2