THE KNOWLEDGE OF THE GENUS EOPOMPILUS GUSSAKOVSKIJ , 1932 ( HYMENOPTERA , POMPILIDAE )

Eopompilus himalayensis Wahis, Lelej et Loktionov, nom. n. is proposed for Pompilus ichneumoniformis Cameron, 1897, nom. praeocc., nec Smith, 1864, nec Patton, 1879 and hitherto unknown male of this species is described and illustrated. A new combination is proposed for Platydialepis ichneumoniformis (Smith, 1864), comb. n. The genus Eopompilus Gussakovskij is newly recorded from India (Sikkim), Laos and Indonesia. An updated key to the species of Eopompilus is given for females and males.


INTRODUCTION
The study of the Pompilidae collection deposited in the Hope Entomological Collections of the Oxford University Museum of Natural History (Oxford, United Kingdom) and the Natural History Museum (London, United Kingdom) by R. Wahis discovered the holotype of Pompilus ichneumoniformis Cameron, 1897, female and the series of specimens of this species.He clarified the taxonomic position of the species which belongs to the genus Eopompilus Gussakovskij.At the same time the Pompilus ichneumoniformis Cameron, 1897 is a junior homonym (nec Smith, 1864, nec Patton, 1879) and new name E. himalayensis Wahis, Lelej et Loktionov, nom.n. is proposed here.Additional material of this species and E. pseudominor Loktionov, Lelej et Xu, 2017 from Laos and Indonesia were studied by V. Loktionov in the Biologiezentrum des Oberösterreichischen Landesmuseums (Linz, Austria).
The genus Eopompilus Gussakovskij, 1932 from the tribe Priocnemini of the subfamily Pepsinae currently includes six species, including E. himalayensis nom.n., distributed in the Eastern Palaearctic and Oriental Region.The review of the genus Eopompilus has been done recently (Loktionov et al., 2017).
Herein we propose the new replacement name, describe hitherto unknown male of Eopompilus himalayensis Wahis, Lelej et Loktionov, nom.n., give an updated key to the species for females and males, enlarge the distribution of the genus within the Oriental Region to include India (Sikkim), Laos and Indonesia, and give an updated diagnosis of E. pseudominor.

MATERIAL AND METHODS
The type materials as well as other materials used in this study are deposited in the following collections: BMNH -The Natural History Museum, London, United Kingdom (formerly British Museum (Natural History)); GxABT -Gembloux Agro-Bio-Tech, Department of Entomology, Belgium; HEC -The Hope Entomological Collections of the Oxford University Museum of Natural History, Oxford, United Kingdom; IBSS -The Federal Scientific Center of East Asia Terrestrial Biodiversity (formerly Institute of Biology and Soil Science), Vladivostok, Russia; OLL -Biologiezentrum des Oberösterreichischen Landesmuseums, Linz, Austria.
The terminology for morphology is based on the glossary provided by the Hymenoptera Anatomy Consortium (2013).Terminology is generally standard; F, S and T are used for flagellomeres, metasomal sterna and terga respectively; PODpostocellar (interocellar) distance between posterior ocelli which is measured from above; OOD -ocellocular distance between posterior ocellus and compound eye which is measured from above; UID -upper interocular distance; MID -middle interocular distance; LID -lower interocular distance.Photographs are made by stereomicroscope SteREO Discovery.V12 and digital camera AxioCam MRc, and stacked by using CombineZM (Hadley, 2008).The final illustrations were postprocessed for contrast and brightness using Adobe® Photoshop® software.DIAGNOSIS.FEMALE.Body predominantly yellowish-brown (Fig. 17-19).Wings light yellow; forewing with distinct apical fascia (Fig. 19).Lateral margin of clypeus normal-shape, without concavity.Claw with subbasal additional tooth.MALE.Proclaws bifid; outer mesoclaw with additional tiny sometimes indistinct tooth medially, inner mesoclaw without additional tooth; metaclaws without additional tooth.S6 with transverse and interrupted medially row of short and stout bristles located little before posterior margin (Fig. 10).Hypopygium (Figs 15,16) in ventral view capitate; dorsal surface with suberect lobe; ventral surface subapically and apically with dense and long bristles.Propodeum with dense small coarse punctures forming granulose texture (Fig. 7).Body from brown to dark brown with abundant yellow on: head, mesosoma, metasoma and legs (Figs 1-9).Wings yellowish without any dark bands (Figs 11,12).
Legs.Protarsomere 1 with three rows of spines ventrally; protarsomeres 2 and 3 ventrally with median row of shorter spines than on protarsomere 1. Protarsomeres 4 and 5 with three rows of spines ventrally.Meso-and metatarsomeres 4 and 5 without spines ventrally.Femora and protibia (except apical part) without spines.Meso-and metatibia with scattered short spines.Metatibia inner side with longitudinal furrow along and above brush.Longest spur of metatibia 0.70-0.75times as long as metatarsomere 1. Orbicula with 6-8 long radiating bristles; longest bristle touching claw top.Proclaws bifid; outer mesoclaw with additional tiny sometimes indistinct tooth medially, inner mesoclaw without additional tooth; metaclaws without additional tooth.
Pubescence.Vertex, gena, propleuron and propodeum laterally with dense and long erect setae.Procoxa with scattered erect setae.Disc of pronotum and S1-S5 with shorter than on vertex erect setae.S6 with group of setae lateroapically (Fig. 10).T5 and T6 with very dense and short setae.Head, meso-, metasoma and legs with micropubescence.
COMPARISON.The hitherto unknown male of Eopompilus himalayensis nom.n. is similar to that of E. ungulivarius Ji et Ma from China in having capitate shape of hypopygium and somewhat similar shape of genitalia and resembles to that of Eopompilus luteus Lelej from Russia and China in having abundant yellow colouration of body, yellowish wings and large body size.It can be easily distinguished from both of them as well as from those of other congeners by the following: proclaws bifid, outer mesoclaw with tiny sometimes indistinct additional tooth medially, inner mesoclaw without additional tooth, metaclaws without additional tooth vs claws without additional tooth in E. internalis (Matsumura), E. luteus Lelej and E. minor Gussakovskij or if proclaws bifid then mesoclaw almost bifid and metaclaw with small additional tooth in E. ungulivarius; S6 with transverse and interrupted medially row of short and stout bristles located little before posterior margin (Fig. 10) vs transverse row of spines located right on posterior margin in other congeners; propodeum with dense small coarse punctures giving granulose texture (Fig. 7) vs propodeum with delicate and rarer punctures in other congeners.
ETYMOLOGY.The specific name refers to the north-west distribution of the species in India, along the Himalayan border.
REMARKS.So far, the species was known for holotype only.We give an updated diagnosis due to the examination of the new material from Laos and India.