Species Testis mass (mg) Paragonial mass (mg) Total body mass (mg) Acilius mediatus 3.8177 1.4805 42.6012 Acilius mediatus 2.8078 1.3167 39.7358 Acilius mediatus 3.5215 1.5904 42.2146 Acilius mediatus 3.2602 1.34 42.9689 Acilius mediatus 3.2299 1.7887 42.4841 Acilius mediatus 2.9743 1.3499 34.8496 Acilius mediatus 3.4713 1.4834 38.096 Acilius mediatus 2.9255 1.4181 36.4656 Acilius mediatus 2.408 1.5292 34.0876 Acilius mediatus 3.68 1.3155 45.3255 Acilius mediatus 2.2069 1.5534 31.1883 Acilius mediatus 3.1184 1.5507 33.8591 Acilius semisulcatus 1.3129 1.1422 52.4159 Acilius semisulcatus 3.2063 2.0871 67.7619 Acilius semisulcatus 4.7894 2.4451 70.9741 Acilius semisulcatus 3.4307 2.4991 69.747 Acilius semisulcatus 3.1043 2.12 60.695 Acilius semisulcatus 4.8524 2.3408 77.6178 Acilius semisulcatus 3.0658 1.8688 61.6384 Acilius semisulcatus 2.1714 1.7672 62.0515 Acilius semisulcatus 3.55 2.0498 69.3772 Acilius semisulcatus 2.5141 1.9184 59.7276 Acilius semisulcatus 4.1955 2.3355 66.0581 Acilius semisulcatus 2.6159 2.5342 62.1819 Agabus ambiguus 0.7489 0.5919 16.9609 Agabus ambiguus 0.3514 0.1197 11.2081 Agabus ambiguus 0.7263 0.507 16.6467 Agabus ambiguus 0.6833 0.6007 15.767 Agabus ambiguus 0.9087 0.4701 15.65 Agabus ambiguus 0.689 0.4021 14.3132 Agabus ambiguus 0.7384 0.6044 15.7041 Agabus ambiguus 0.5679 0.3396 12.1454 Agabus ambiguus 0.7598 0.702 18.3095 Agabus ambiguus 0.7743 0.6425 13.844 Celina hubbelli 0.0379 0.0308 1.3919 Celina hubbelli 0.081 0.0395 1.6493 Celina hubbelli 0.0456 0.025 1.3261 Celina hubbelli 0.0595 0.0301 1.3667 Celina hubbelli 0.059 0.0264 1.6639 Celina hubbelli 0.0426 0.0261 1.4444 Celina hubbelli 0.0631 0.0355 1.6304 Celina hubbelli 0.0439 0.0258 1.438 Celina hubbelli 0.0474 0.0294 1.4588 Celina hubbelli 0.0575 0.0371 1.4934 Celina hubbelli 0.0448 0.0247 1.5155 Celina hubbelli 0.0471 0.0225 1.2878 Coptotomus longulus 0.6244 0.4514 13.46 Coptotomus longulus 0.5643 0.1779 12.6583 Coptotomus longulus 1.1384 0.2941 11.0784 Coptotomus longulus 0.663 0.3572 12.5028 Coptotomus longulus 0.5751 0.3645 12.0289 Coptotomus longulus 0.5573 0.3279 13.4821 Coptotomus longulus 0.5125 0.2416 12.1511 Coptotomus longulus 0.4043 0.277 10.7153 Coptotomus longulus 0.3118 0.1908 9.8692 Coptotomus longulus 0.4383 0.2457 12.1719 Coptotomus longulus 0.4158 0.3053 13.318 Coptotomus longulus 0.4983 0.3972 12.113 Desmopachria convexa 0.0016 0.0029 0.2263 Desmopachria convexa 0.0026 0.0042 0.2378 Desmopachria convexa 0.0014 0.003 0.2784 Desmopachria convexa 0.0092 0.0041 0.2614 Desmopachria convexa 0.0086 0.0062 0.2531 Desmopachria convexa 0.0083 0.0075 0.3205 Desmopachria convexa 0.0057 0.0047 0.2632 Desmopachria convexa 0.0028 0.0005 0.1761 Desmopachria convexa 0.0062 0.0051 0.2651 Desmopachria convexa 0.0012 0.0026 0.233 Desmopachria convexa 0.0073 0.0053 0.2954 Desmopachria convexa 0.0073 0.004 0.2968 Graphoderus liberus 2.3667 0.198 32.0334 Graphoderus liberus 3.5116 0.2709 34.9708 Graphoderus liberus 1.5961 0.169 25.8441 Graphoderus liberus 0.9356 0.1339 22.7377 Graphoderus liberus 2.3318 0.1807 29.3654 Graphoderus liberus 2.3263 0.2168 33.6617 Graphoderus liberus 2.3315 0.146 38.3914 Graphoderus liberus 2.8253 0.4282 37.1482 Graphoderus liberus 3.9857 0.3327 38.5495 Graphoderus liberus 2.1203 0.1827 30.6901 Graphoderus liberus 2.8698 0.2746 31.5379 Graphoderus liberus 1.4036 0.2193 26.2861 Heterosternuta pulcher 0.0114 0.0722 1.165 Heterosternuta pulcher 0.0102 0.0896 1.2157 Heterosternuta pulcher 0.0098 0.0868 1.1591 Heterosternuta pulcher 0.0075 0.0284 0.8908 Heterosternuta pulcher 0.0097 0.0778 1.3816 Heterosternuta pulcher 0.0118 0.0682 1.1705 Heterosternuta pulcher 0.0094 0.0622 0.8858 Heterosternuta pulcher 0.0077 0.0832 1.1438 Heterosternuta pulcher 0.0095 0.0718 1.1787 Heterosternuta pulcher 0.0105 0.0783 1.0894 Heterosternuta pulcher 0.0086 0.0777 1.1638 Heterosternuta pulcher 0.0156 0.0816 1.3357 Hydrovatus pustulatus 0.022 0.0101 0.74 Hydrovatus pustulatus 0.0189 0.0057 0.9524 Hydrovatus pustulatus 0.0103 0.0037 0.5883 Hydrovatus pustulatus 0.016 0.0073 0.7578 Hydrovatus pustulatus 0.0221 0.009 0.7679 Hydrovatus pustulatus 0.014 0.0057 0.6427 Hydrovatus pustulatus 0.0146 0.0027 0.5278 Hydrovatus pustulatus 0.0139 0.0058 0.6779 Hydrovatus pustulatus 0.0112 0.0042 0.6137 Hydrovatus pustulatus 0.0129 0.0051 0.6774 Hydrovatus pustulatus 0.0235 0.0084 0.6449 Hydrovatus pustulatus 0.0202 0.009 0.8313 Hygrotus sayi 0.0332 0.0645 1.2853 Hygrotus sayi 0.0219 0.0762 1.5572 Hygrotus sayi 0.0333 0.0777 1.4324 Hygrotus sayi 0.031 0.0684 1.1455 Hygrotus sayi 0.0376 0.0717 1.5357 Hygrotus sayi 0.0301 0.0846 1.3874 Hygrotus sayi 0.0316 0.0738 1.3988 Hygrotus sayi 0.0281 0.0846 1.4133 Hygrotus sayi 0.0263 0.0682 1.3222 Hygrotus sayi 0.0268 0.077 1.5574 Hygrotus sayi 0.0411 0.0744 1.4292 Hygrotus sayi 0.0359 0.0819 1.4985 Hydroporus sp. 0.0401 0.7358 2.9738 Hydroporus sp. 0.0669 0.4145 3.1326 Hydroporus sp. 0.0431 0.6436 3.0563 Hydroporus sp. 0.0426 0.625 3.2536 Hydroporus sp. 0.0393 0.729 3.2729 Hydroporus sp. 0.0233 0.3787 2.8992 Hydroporus sp. 0.0281 0.5014 2.6489 Hydroporus sp. 0.0303 0.5528 3.1508 Hydroporus sp. 0.0295 0.5716 2.83 Hydroporus sp. 0.0389 0.7114 3.2981 Hydroporus sp. 0.032 0.4658 3.0614 Hydroporus sp. 0.0321 0.4478 3.2224 Ilybius oblitus 1.2443 2.0582 27.5435 Ilybius oblitus 2.2946 2.2305 33.767 Ilybius oblitus 1.5387 2.2881 31.6605 Ilybius oblitus 1.556 2.2112 29.5167 Ilybius oblitus 1.5749 2.0786 27.4389 Ilybius oblitus 1.7515 2.3283 30.3283 Ilybius oblitus 1.25 2.0745 25.8936 Ilybius oblitus 1.1959 2.4996 28.0707 Ilybius oblitus 1.562 1.8007 24.2744 Ilybius oblitus 1.2655 2.0702 30.1233 Ilybius oblitus 1.6274 2.4093 32.2882 Ilybius oblitus 1.5098 2.394 32.1959 Laccophilus maculosus 0.0461 0.1232 4.3917 Laccophilus maculosus 0.0486 0.1272 3.5959 Laccophilus maculosus 0.0442 0.1263 3.8869 Laccophilus maculosus 0.0285 0.0668 3.1535 Laccophilus maculosus 0.0436 0.0625 3.7712 Laccophilus maculosus 0.0278 0.0797 3.4926 Laccophilus maculosus 0.0615 0.0765 4.7658 Laccophilus maculosus 0.1013 0.0458 4.6746 Laccophilus maculosus 0.0336 0.0415 3.0008 Laccophilus maculosus 0.0541 0.0379 4.4246 Laccophilus maculosus 0.0487 0.0108 4.5445 Laccophilus maculosus 0.0816 0.0462 4.4508 Neoporus dimidiatus 0.1852 0.156 2.3437 Neoporus dimidiatus 0.1405 0.1183 2.0473 Neoporus dimidiatus 0.136 0.0957 2.1226 Neoporus dimidiatus 0.1634 0.0305 1.5951 Neoporus dimidiatus 0.1287 0.0263 1.5056 Neoporus dimidiatus 0.1522 0.0633 1.8015 Neoporus dimidiatus 0.1501 0.1122 2.3539 Neoporus dimidiatus 0.2001 0.0471 1.8027 Neoporus dimidiatus 0.1669 0.0846 1.8918 Neoporus dimidiatus 0.1399 0.0488 2.1909 Neoporus dimidiatus 0.1901 0.0839 1.9026 Neoporus dimidiatus 0.087 0.1178 2.1266 Nebrioporus rotundatus 0.0699 0.224 3.7136 Nebrioporus rotundatus 0.1189 0.2468 4.0326 Nebrioporus rotundatus 0.0865 0.2503 4.1346 Nebrioporus rotundatus 0.0933 0.2328 3.9306 Nebrioporus rotundatus 0.0946 0.2513 4.3949 Nebrioporus rotundatus 0.0822 0.2303 4.0116 Nebrioporus rotundatus 0.0857 0.2776 4.1908 Nebrioporus rotundatus 0.0942 0.2415 3.9389 Nebrioporus rotundatus 0.0882 0.2485 3.725 Nebrioporus rotundatus 0.0871 0.254 4.1929 Nebrioporus rotundatus 0.1006 0.2354 3.5895 Nebrioporus rotundatus 0.0871 0.2394 3.8495 Neoporus undulatus 0.0836 0.3209 2.8179 Neoporus undulatus 0.0893 0.2369 2.1849 Neoporus undulatus 0.058 0.3937 2.5579 Neoporus undulatus 0.11999 0.3499 2.71869 Neoporus undulatus 0.0845 0.3222 2.6934 Neoporus undulatus 0.0721 0.3326 2.2893 Neoporus undulatus 0.0684 0.3724 2.8585 Neoporus undulatus 0.07 0.3525 2.5097 Neoporus undulatus 0.0807 0.3802 2.7739 Neoporus undulatus 0.0602 0.2941 2.1809 Neoporus undulatus 0.0717 0.3708 2.472 Neoporus undulatus 0.0866 0.396 2.7367 Rhantus binotatus 1.2806 1.5204 28.804 Rhantus binotatus 1.3102 1.804 27.6697 Rhantus binotatus 1.7093 2.2772 29.7775 Rhantus binotatus 1.4833 1.3885 27.8262 Rhantus binotatus 1.4227 2.0616 27.6415 Rhantus binotatus 1.4729 1.5347 26.3595 Rhantus binotatus 1.3207 2.0193 30.9375 Rhantus binotatus 1.4791 2.1526 30.0795 Rhantus binotatus 1.4014 1.5587 26.7136 Rhantus binotatus 1.4025 1.3641 29.992 Rhantus binotatus 1.8721 2.4199 33.4418 Rhantus binotatus 1.4224 1.8541 29.2173 Dineutus ciliatus 2.0903 1.8171 46.6927 Dineutus ciliatus 3.1077 1.8791 61.7999 Dineutus ciliatus 2.7662 2.2176 63.0836 Dineutus ciliatus 2.5298 2.2154 59.2308 Dineutus ciliatus 2.67842 2.16852 57.12284 Dineutus ciliatus 2.8242 2.2173 56.0163 Dineutus ciliatus 2.5308 1.8726 52.1519 Dineutus ciliatus 2.0701 1.4775 46.1988 Dineutus ciliatus 2.4958 1.4765 55.2876 Dineutus ciliatus 2.536 1.926 58.3499 Dineutus ciliatus 2.3791 1.6735 46.6538 Dineutus ciliatus 2.5413 1.9889 50.0062 Dineutus discolor 1.2659 2.7946 35.4903 Dineutus discolor 0.8981 2.8001 33.6013 Dineutus discolor 0.931 2.6797 30.8846 Dineutus discolor 1.0482 3.075 31.7251 Dineutus discolor 1.1053 2.8919 32.0809 Dineutus discolor 0.9597 2.664 30.6913 Dineutus discolor 0.9976 2.7578 34.6165 Dineutus discolor 1.0721 2.5413 32.756 Dineutus discolor 1.0181 2.8343 37.2233 Dineutus discolor 1.2752 2.5886 36.6164 Dineutus discolor 1.0631 2.9582 33.9296 Dineutus discolor 1.0073 2.607 33.0875 Dineutus emarginatus 1.0521 1.0276 20.5148 Dineutus emarginatus 1.0609 1.1887 19.5249 Dineutus emarginatus 0.9556 1.0809 20.3214 Dineutus emarginatus 1.1302 1.0515 20.6319 Dineutus emarginatus 1.2052 1.2543 20.0739 Dineutus emarginatus 1.0807 0.9735 18.4693 Dineutus emarginatus 1.0697 0.7789 18.0842 Dineutus emarginatus 0.9946 0.9011 18.8408 Dineutus emarginatus 1.2087 1.0929 23.8491 Dineutus emarginatus 1.0297 0.9519 19.0645 Dineutus emarginatus 0.9302 0.657 16.8317 Dineutus emarginatus 1.317 1.0298 21.5969 Dineutus horni 1.00E+00 2.0266 19.6396 Dineutus horni 1.4409 2.0683 21.7627 Dineutus horni 1.0833 1.7542 18.4839 Dineutus horni 1.1689 1.7619 20.5995 Dineutus horni 1.1579 1.9021 21.0669 Dineutus horni 1.0548 1.6396 18.7179 Dineutus horni 1.4117 2.1733 23.4313 Dineutus horni 1.1502 1.7437 19.8993 Dineutus horni 1.1815 1.5242 19.84 Dineutus horni 1.2876 2.3244 23.3388 Dineutus horni 1.1004 1.5219 21.767 Dineutus horni 1.3129 1.7933 23.442 Dineutus nigrior 1.2858 2.3279 30.4877 Dineutus nigrior 1.3806 2.4397 28.3154 Dineutus nigrior 1.3755 2.3493 25.6789 Dineutus nigrior 1.4151 2.1381 24.431 Dineutus nigrior 1.3978 2.1134 25.3877 Dineutus nigrior 1.4804 2.4154 26.6066 Dineutus nigrior 1.3838 2.4208 26.399 Dineutus nigrior 1.2841 3.2445 29.2099 Dineutus nigrior 1.3623 1.985 24.6456 Dineutus nigrior 1.3573 2.194 25.6942 Dineutus nigrior 1.3032 2.1757 24.7451 Dineutus nigrior 1.381 2.0479 24.7078 Gyrinus gibber 0.0826 0.1751 3.9784 Gyrinus gibber 0.1285 0.3639 4.5206 Gyrinus gibber 0.099 0.3211 4.273 Gyrinus gibber 0.1118 0.3209 4.8348 Gyrinus gibber 0.1568 0.2106 4.3036 Gyrinus gibber 0.1198 0.2424 4.4346 Gyrinus gibber 0.0789 0.325 4.9388 Gyrinus gibber 0.1681 0.3283 5.0042 Gyrinus gibber 0.1278 0.3517 5.4206 Gyrinus gibber 0.0853 0.2317 4.8102 Gyrinus gibber 0.1166 0.2423 4.2373 Gyrinus gibber 0.1039 0.3476 4.5018 Gyrinus latilimbus 0.044 0.1138 2.1028 Gyrinus latilimbus 0.0367 0.1422 2.0459 Gyrinus latilimbus 0.0405 0.1413 2.3973 Gyrinus latilimbus 0.0324 0.1546 2.148 Gyrinus latilimbus 0.03 0.1613 2.2472 Gyrinus latilimbus 0.0726 0.1591 2.307 Gyrinus latilimbus 0.0267 0.1743 2.2792 Gyrinus latilimbus 0.0325 0.1618 2.3164 Gyrinus latilimbus 0.0315 0.1576 2.4125 Gyrinus latilimbus 0.0355 0.044 2.6509 Gyrinus latilimbus 0.0489 0.0476 2.7549 Gyrinus latilimbus 0.0392 0.0917 2.0293 Gyrinus marginellus 0.0438 0.0315 2.5379 Gyrinus marginellus 0.0828 0.0149 2.996 Gyrinus marginellus 0.0892 0.102 2.2127 Gyrinus marginellus 0.07538 0.12 2.37928 Gyrinus marginellus 0.1602 0.0148 2.9065 Gyrinus marginellus 0.085 0.022 2.4738 Gyrinus marginellus 0.1537 0.0474 3.2207 Gyrinus marginellus 0.1307 0.022 2.6567 Gyrinus marginellus 0.0984 0.0355 1.988 Gyrinus marginellus 0.1142 0.0116 2.573 Gyrinus marginellus 0.0516 0.0602 2.1756 Gyrinus marginellus 0.0947 0.1115 3.0358 Gyretes iricolor 0.2396 0.0781 3.4116 Gyretes iricolor 0.1472 0.0832 2.9508 Gyretes iricolor 0.2041 0.0794 3.2893 Gyretes iricolor 0.2109 0.0819 3.6279 Gyretes iricolor 0.2332 0.0988 3.9058 Gyretes iricolor 0.1993 0.0778 3.5683 Gyretes iricolor 0.2443 0.0973 3.6611 Gyretes iricolor 0.2527 0.0767 3.1192 Gyretes iricolor 0.1294 0.0834 3.4245 Gyretes iricolor 0.1872 0.0621 3.2134 Gyretes iricolor 0.2605 0.0464 3.9242 Gyretes iricolor 0.2107 0.0853 3.4122 Gyrinus fraternus 0.0139 0.2272 4.0374 Gyrinus fraternus 0.0148 0.0187 2.8499 Gyrinus fraternus 0.0121 0.0403 4.0844 Gyrinus fraternus 0.0282 0.0272 3.4997 Gyrinus fraternus 0.0167 0.0267 3.2292 Gyrinus fraternus 0.0117 0.0359 3.4302 Gyrinus fraternus 0.0224 0.0455 4.1227 Gyrinus fraternus 0.0353 0.0332 3.0905 Gyrinus fraternus 0.0155 0.2283 4.766 Gyrinus fraternus 0.02 0.2111 3.7471 Gyrinus fraternus 0.0147 0.0371 3.2427 Gyrinus fraternus 0.018 0.0368 4.485 Haliplus immacucolis 0.0208 0.0306 0.9987 Haliplus immacucolis 0.0184 0.0336 0.7903 Haliplus immacucolis 0.0224 0.0349 1.1461 Haliplus immacucolis 0.0156 0.0389 0.809 Haliplus immacucolis 0.0234 0.036 0.9878 Haliplus immacucolis 0.0184 0.0439 1.0408 Haliplus immacucolis 0.0216 0.0292 0.9594 Haliplus immacucolis 0.0174 0.0302 1.0197 Haliplus immacucolis 0.0243 0.0317 1.1103 Haliplus immacucolis 0.0064 0.0302 0.8826 Haliplus immacucolis 0.0231 0.032 1.0113 Haliplus immacucolis 0.0246 0.0369 1.0017 Haliplus fasciatus 0.0332 0.0169 3.6618 Haliplus fasciatus 0.0296 0.02 3.3532 Haliplus fasciatus 0.0302 0.0271 3.3409 Haliplus fasciatus 0.0313 0.0255 3.369 Haliplus fasciatus 0.0269 0.0193 3.2416 Haliplus fasciatus 0.0387 0.022 3.3971 Haliplus fasciatus 0.0205 0.0204 3.0201 Haliplus fasciatus 0.0182 0.016 3.3782 Haliplus fasciatus 0.0536 0.0175 3.5425 Haliplus fasciatus 0.0178 0.0197 3.0803 Haliplus fasciatus 0.0224 0.0193 3.3154 Haliplus fasciatus 0.0183 0.0241 2.7451 Hydrocanthus iricolor 0.0228 0.1981 3.0402 Hydrocanthus iricolor 0.0208 0.1978 2.4399 Hydrocanthus iricolor 0.0216 0.1339 3.025 Hydrocanthus iricolor 0.0284 0.2695 2.842 Hydrocanthus iricolor 0.035 0.2861 2.9711 Hydrocanthus iricolor 0.0336 0.2837 2.5211 Hydrocanthus iricolor 0.027 0.3114 2.8562 Hydrocanthus iricolor 0.0301 0.2977 2.4473 Hydrocanthus iricolor 0.0301 0.264 2.6641 Hydrocanthus iricolor 0.0278 0.2038 2.8241 Hydrocanthus iricolor 0.0323 0.2817 2.7626 Hydrocanthus iricolor 0.0368 0.3062 2.6465